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Lycosoid
Lycosoidea is a clade or superfamily of araneomorph spiders. The traditional circumscription was based on a feature of the eyes. The tapetum is a reflective layer at the back of the eye, thought to increase sensitivity in low light levels. Lycosoids were then defined by having a "grate-shaped" tapetum. Research from the late 1990s onwards suggests that this feature has evolved more than once, possibly as many as five times, so that the original Lycosoidea is paraphyletic. Studies published in 2014 and 2015 suggest that a smaller group of families does form a clade. The reflective attributes of the tapeta vary significantly betwixt lycosoid spiders. Phylogeny The phylogeny of the Lycosoidea and related groups has been studied using both morphological and molecular data. Comparing these studies is complicated by the redrawing of family boundaries. Thus potential lycosoid genera placed in the family Miturgidae in 1993, such as '' Uliodon'', have since been placed in the Zoropsidae ...
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Udubidae
Udubidae is a family of araneomorph spiders, most of whose members were formerly placed in the family Zorocratidae, which is no longer accepted. Phylogeny A study investigating the phylogenetic relationships of lycosoid spiders concluded that the genera formerly placed in the family Zorocratidae fell into two groups. The largest group formed the sister clade to the "grate-shaped tapetum clade" (see the cladogram below). The type genus, '' Zorocrates'', grouped with '' Zoropsis'' inside the grate-shaped tapetum clade. Some earlier studies had also cast doubt on the monophyly of the Zorocratidae. A 2003 study found that ''Raecius'', ''Uduba'', and ''Zorodictyna'' formed a clade somewhat separated from ''Zorocrates''. A 2014 study including ''Zorocrates'' and ''Raecius'' did not find that they grouped together., p. 289 Moving the type genus to a different family meant that a new family name was needed for the remaining members. Griswold and Polotow proposed "Udubidae", with t ...
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Zoropsidae
Zoropsidae, also known as false wolf spiders for their physical similarity to wolf spiders, is a family of cribellate araneomorph spiders first described by Philipp Bertkau in 1882. They can be distinguished from wolf spiders by their two rows of eyes that are more equal in size than those of Lycosidae. The families Tengellidae and Zorocratidae are now included in Zoropsidae. Genera , the World Spider Catalog accepts the following genera: *''Akamasia'' Bosselaers, 2002 – Cyprus *'' Anachemmis'' Chamberlin, 1919 – United States, Mexico *'' Austrotengella'' Raven, 2012 – Australia *'' Birrana'' Raven & Stumkat, 2005 – Australia *''Cauquenia'' Piacentini, Ramírez & Silva, 2013 *''Chinja'' Polotow & Griswold, 2018 – Tanzania *'' Ciniflella'' Mello-Leitão, 1921 – Brazil *'' Devendra'' Lehtinen, 1967 – Sri Lanka *'' Griswoldia'' Dippenaar-Schoeman & Jocqué, 1997 – South Africa *''Hoedillus'' Simon, 1898 – Guatemala, Nicaragua *''Huntia'' Gray & Thompson, 2001 � ...
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Psechridae
Psechridae is a family of araneomorph spiders with about 70 species in two genera. These are among the biggest cribellate spiders with body lengths up to and funnel webs more than in diameter. The family belongs to the RTA clade of spiders because they all have a Retrolateral Tibial Apophysis on the male pedipalp. A recent phylogenetic analysis places Psechridae as close relatives of the lynx spiders, wolf spiders, and nursery web spiders. They feature several characteristics normally found in ecribellate spiders, for example brood care behavior, and a colulus with no apparent function. They have greatly elongated legs, with the last element being very flexible. Female '' Psechrus'' carry their egg-sac in the chelicerae, similar to their relatives, the ecribellate Pisauridae. Members of ''Psechrus'' construct horizontal webs lace webs, while ''Fecenia'' construct pseudo-orbs, similar to orb webs of Orbiculariae spiders in an example of evolutionary convergence. Distribu ...
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Lycosidae
Wolf spiders are members of the family Lycosidae (). They are robust and agile hunters with excellent eyesight. They live mostly in solitude, hunt alone, and do not spin webs. Some are opportunistic hunters, pouncing upon prey as they find it or chasing it over short distances; others wait for passing prey in or near the mouth of a burrow. Wolf spiders resemble nursery web spiders (family Pisauridae), but wolf spiders carry their egg sacs by attaching them to their spinnerets, while the Pisauridae carry their egg sacs with their chelicerae and pedipalps. Two of the wolf spider's eight eyes are large and prominent; this distinguishes them from nursery web spiders, whose eyes are all of roughly equal size. This can also help distinguish them from the similar-looking grass spiders. Description The many genera of wolf spiders range in body size (legs not included) from less than . They have eight eyes arranged in three rows. The bottom row consists of four small eyes, the middle r ...
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Dionycha
The Dionycha are a clade of spiders (Araneomorphae:Entelegynae), characterized by the possession of two tarsal claws with tufts of hairs (setae) beside them, which produce strong adhesion, enabling some species to climb glass. The circumscription of the group has varied widely; a 2021 analysis resulted in about 20 families, including Salticidae (jumping spiders), Gnaphosidae (ground spiders), and Clubionidae. The Dionycha are considered to be a subgroup of the larger RTA clade. Most species hunt their prey instead of building webs. There are no cribellate members in the Dionycha.Griswold ''et al.'' 1999 Today it is thought that the reduction of the third claw present in ancestral spiders evolved several times independently, so this alone is not a criterion that defines the clade. Families In 2021, a group of several spider taxonomists published a major study of the phylogeny of Dionycha, using genetic and phenotypic data. It included the families listed below: ;Dionycha *Pr ...
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RTA Clade
The RTA clade is a clade of araneomorph spiders, united by the possession of a retrolateral tibial apophysis – a backward-facing projection on the tibia of the male pedipalp. The clade contains over 21,000 species, almost half the current total of about 46,000 known species of spider. Most of the members of the clade are wanderers and do not build webs. Despite making up approximately half of all modern spider diversity, there are no unambiguous records of the group from the Mesozoic and molecular clock evidence suggests that the group began to diversify during the Late Cretaceous. Families In 2005, Coddington included 39 families in a cladogram showing the RTA clade: *Agelenidae *Amaurobiidae *Ammoxenidae *Amphinectidae (paraphyletic; merged into Desidae) *Anyphaenidae *Cithaeronidae *Clubionidae *Corinnidae * Cryptothelidae *Ctenidae *Desidae *Dictynidae * Gallieniellidae *Gnaphosidae *Lamponidae * Liocranidae *Lycosidae *Miturgidae (paraphyletic) * Oxyopidae *Philodromidae ...
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Trechaleidae
Trechaleidae (''tre-kah-LEE-ih-dee'') is a family of araneomorph spiders first described by Eugène Simon in 1890, and includes about 140 described species in 16 genera. They all live in Central and South America except for '' Shinobius orientalis'', which is endemic to Japan. Other names for the family are longlegged water spiders and fishing spiders. Genera , the World Spider Catalog accepts the following genera: *'' Amapalea'' Silva & Lise, 2006 – Brazil *''Barrisca'' Chamberlin & Ivie, 1936 – South America, Panama *''Caricelea'' Silva & Lise, 2007 – Peru *''Cupiennius'' Simon, 1891 – Mexico to northwestern South America *''Dossenus'' Simon, 1898 – Trinidad, South America *''Dyrines'' Simon, 1903 – South America, Panama *'' Enna'' O. Pickard-Cambridge, 1897 – South America, Central America, Mexico *''Heidrunea'' Brescovit & Höfer, 1994 – Brazil *''Hesydrus'' Simon, 1898 – South America, Central America *''Neoctenus'' Simon, 1897 – Brazil, ...
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Thomisidae
The Thomisidae are a family of spiders, including about 170 genera and over 2,100 species. The common name crab spider is often linked to species in this family, but is also applied loosely to many other families of spiders. Many members of this family are also known as flower spiders or flower crab spiders. Description Members of this family of spiders do not spin webs, and are ambush predators. The two front legs are usually longer and more robust than the rest of the legs. The back two legs are smaller, and are usually covered in a series of strong spines. They have dull colorations such as brown, grey, or very bright green, pink, white or yellow. They gain their name from the shape of their body, and they usually move sideways or backwards. These spiders are quite easy to identify and can very rarely be confused with Sparassidae family, though the crab spiders are usually smaller. Etymology Spiders in this family are called "crab spiders" due to their resemblance to crabs ...
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Stiphidiidae
Stiphidiidae, also called sheetweb spiders, is a family of araneomorph spiders first described in 1917. Most species are medium size (''Stiphidion facetum'' is about long) and speckled brown with long legs. All members of this family occur in New Zealand and Australia except for '' Asmea''. They build a horizontal sheet-like web under rocks, hence the name "sheetweb spiders". The largest of New Zealand's species is ''Cambridgea foliata'', with a body length up to and a span of up to . Hikers and trampers often find their sheet-like webs that can be up to across, but the spider itself is nocturnal, spending the day time inside its web tunnel. It can also be found in gardens and males may enter human homes. Their large size, including mouth parts up to long, may be intimidating, but it is considered harmless to humans and bites are extremely rare. Genera , the World Spider Catalog accepts the following genera: *'' Aorangia'' Forster & Wilton, 1973 — New Zealand *'' Asm ...
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Senoculidae
''Senoculus'' is a genus of araneomorph spiders in the family Senoculidae, and was first described by Władysław Taczanowski in 1872. It is the only genus in the family Senoculidae. Species it contains thirty-one species, found in South America, Guatemala, Panama, Mexico, and on Trinidad: *'' Senoculus albidus'' ( F. O. Pickard-Cambridge, 1897) – Brazil *'' Senoculus barroanus'' Chickering, 1941 – Panama *'' Senoculus bucolicus'' Chickering, 1941 – Panama *'' Senoculus cambridgei'' Mello-Leitão, 1927 – Brazil *'' Senoculus canaliculatus'' F. O. Pickard-Cambridge, 1902 – Mexico to Panama *'' Senoculus carminatus'' Mello-Leitão, 1927 – Brazil *'' Senoculus darwini'' (Holmberg, 1883) – Argentina *'' Senoculus fimbriatus'' Mello-Leitão, 1927 – Brazil *'' Senoculus gracilis'' ( Keyserling, 1879) – Guyana to Argentina *'' Senoculus guianensis'' Caporiacco, 1947 – Guyana *''Senoculus iricolor'' (Simon, 1880) – Brazil *'' Senoculus maronicus'' Taczanowski, 187 ...
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Pisauridae
Nursery web spiders (Pisauridae) is a family of araneomorph spiders first described by Eugène Simon in 1890. They resemble wolf spiders (Lycosidae) except for several key differences. Wolf spiders have two very prominent eyes in addition to the other six, while a nursery web spider's eyes are all about the same size. Additionally, female nursery web spiders carry their egg sacs with their jaws and pedipalps instead of attaching them to their spinnerets as wolf spiders do. When the eggs are about to hatch, a female spider builds a nursery "tent", places her egg sac inside, and stands guard outside, hence the family's common name. Like the wolf spiders, however, the nursery web spiders are roaming hunters that don't use webs for catching prey. Species occur throughout the world except for extremely dry or cold environments, and are common just about everywhere. Many can walk on the surface of still bodies of water and may even dive beneath the surface temporarily to escape enem ...
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Oxyopidae
Lynx spider (Oxyopidae) is a family of araneomorph spiders first described by Tamerlan Thorell in 1870. Most species make little use of webs, instead spending their lives as hunting spiders on plants. Many species frequent flowers in particular, ambushing pollinators, much as crab spiders do. They tend to tolerate members of their own species more than most spiders do, and at least one species has been identified as exhibiting social behaviour. Description There are several genera of Oxyopidae and they differ in their habits and adaptations. Most of them have large spiny bristles on their legs and in many species the bristles form almost a basket-like structure that may assist in confining the prey that they grasp, and protect the spider from its struggles. Most '' Oxyopes'' and '' Hamataliwa'' species are small to medium in size. Lynx spiders, in spite of being largely ambush hunters, are very speedy runners and leapers, alert and with good vision. Oxyopidae in general rely on k ...
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