Leptoceratopsinae
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Leptoceratopsinae
Leptoceratopsidae is an extinct family of neoceratopsian dinosaurs from Asia, North America and possibly Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the families Protoceratopsidae and Ceratopsidae, but they were more primitive and generally smaller. Phylogeny Leptoceratopsidae was originally named by Franz Nopcsa von Felső-Szilvás in 1923 as a subfamily Leptoceratopsinae, and its type species is ''Leptoceratops gracilis''. Mackovicky, in 2001, defined it as a stem-based taxon and a family consisting of ''Leptoceratops gracilis'' and all species closer to ''Leptoceratops'' than to ''Triceratops horridus''.Makovicky, P.J. 2001. A ''Montanoceratops cerorhynchus'' (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta, In: Tanke, D.H. & Carpenter, K. (Eds.). ''Mesozoic Vertebrate Life''. Bloomington: Indiana University Press. Pp. 243-262. All previously published neoceratopsian phylogenetic analyses we ...
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Leptoceratops
''Leptoceratops'' (meaning 'small horn face') is a genus of ceratopsian dinosaur from the Late Cretaceous of North America. First found in Alberta in 1910, the type species ''Leptoceratops gracilis'' was named in 1914 by Barnum Brown for a partial skull and skeleton of two individuals found in the Scollard Formation of Alberta. Additional specimens found in the Scollard include one complete and two mostly complete skeletons together, uncovered in 1947 by Charles M. Sternberg. Specimens from Montana that were among the earliest referred to ''Leptoceratops'' have since been moved to their own genera ''Montanoceratops'' and ''Cerasinops'', while new specimens of ''L. gracilis'' include bonebed remains from the Hell Creek Formation of Montana and a partial skeleton from the Lance Formation of Wyoming. Together with related taxa, ''Leptoceratops'' is the eponymous genus of the family Leptoceratopsidae. ''Leptoceratops'' is known from more than ten individuals, all from Maastrichtian de ...
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Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the Latin word for the white limestone known as chalk. The chalk of northern France and the white cliffs of south-eastern England date from the Cretaceous Period. Climate During the Late Cretaceous, the climate was warmer than present, although throughout the period a cooling trend is evident. The tropics became restricted to equatorial regions and northern latitudes experienced markedly more seasonal climatic conditions. Geography Due to plate tectonics, the Americas were gradually moving westward, causing the Atlantic Ocean to expand. The Western Interior Seaway divided North America into eastern and western halves; Appalachia and Laramidia. India maintained a northward course towards Asia. In the Southern Hemisphere, Aus ...
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Neoceratopsian
Ceratopsia or Ceratopia ( or ; Greek: "horned faces") is a group of herbivorous, beaked dinosaurs that thrived in what are now North America, Asia and Europe, during the Cretaceous Period, although ancestral forms lived earlier, in the Late Jurassic of Asia. The earliest known ceratopsian, '' Yinlong downsi'', lived between 161.2 and 155.7 million years ago.Holtz, Thomas R. Jr. (2011) ''Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,'Winter 2010 Appendix./ref> The last ceratopsian species, '' Triceratops prorsus'', became extinct during the Cretaceous–Paleogene extinction event, . ''Triceratops'' is by far the best-known ceratopsian to the general public. It is traditional for ceratopsian genus names to end in "''-ceratops''", although this is not always the case. One of the first named genera was '' Ceratops'' itself, which lent its name to the group, although it is considered a ''nomen dubium'' today as its fossil remains have no disti ...
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Platyceratops
''Bagaceratops'' (meaning "small-horned face") is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 72 to 71 million years ago. ''Bagaceratops'' remains have been reported from the Barun Goyot Formation and Bayan Mandahu Formation. One specimen may argue the possible presence of ''Bagaceratops'' in the Djadochta Formation. ''Bagaceratops'' was among the smallest ceratopsians, growing up to in length, with a weight about . Although emerging late in the reign of the dinosaurs, ''Bagaceratops'' had a fairly primitive anatomy—when compared to the much derived ceratopsids—and kept the small body size that characterized early ceratopsians. Unlike its close relative, ''Protoceratops'', ''Bagaceratops'' lacked premaxillary teeth (cylindrical, blunt teeth near the tip of the upper jaw). History of discovery During the large field work of the Polish-Mongolian Palaeontological Expeditions in the 1970s, abundant protoceratopsid specimen ...
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Archaeoceratops Yujingziensis
''Archaeoceratops'', meaning "ancient horned face", is a genus of basal neoceratopsian dinosaur from the Early Cretaceous (Aptian stage) of north central China. It appears to have been bipedal and quite small, reaching in length and in body mass. It had a comparatively large head but no horns, possessing only a small bony frill projecting from the back of its head. Discovery and species Two specimens were found in the Middle Gray Unit of the Xinminbao Group, Gongpoquan Basin of the Mazong Shan area of Gansu Province, north central China. The type species, ''A. oshimai'', was named by Dong Zhiming and Azuma in 1997. It is the first basal neoceratopsian discovered in this area. The type specimen, IVPP V11114, consists of a partially complete skeleton including skull, caudal vertebrae, pelvis, and most of a hind foot. The second specimen (paratype), IVPP V11115, consists of an incomplete skeleton with a relatively well preserved caudal series, a partial hind limb, and a co ...
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2019 In Paleontology
Flora Plants Fungi Paleomycological research * Fossil sporocarps indistinguishable from sporocarps of members of the extant genus ''Stemonitis'' are described from the Cretaceous amber from Myanmar by Rikkinen, Grimaldi & Schmidt (2019). * A study on the impact of major historical events such as the Cretaceous–Paleogene extinction event on the evolution of two major subclasses of lichen-forming fungi ( Lecanoromycetidae and Ostropomycetidae) is published by Huang ''et al.'' (2019). * Description of crustose lichens from European Paleogene amber is published by Kaasalainen ''et al.'' (2019). * Fungi belonging to the genera '' Periconia'', ''Penicillium'' and '' Scopulariopsis'', representing the first and the oldest known fossil record of these taxa, are described from the Eocene Baltic amber by Tischer ''et al.'' (2019). Sponges Research * Sponge spicules and spicule-like structures that probably represent sponge fossils are described from four sections of the Ediacara ...
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Triceratops Horridus
''Triceratops'' ( ; ) is a genus of Chasmosaurinae, chasmosaurine Ceratopsia, ceratopsian dinosaur that lived during the late Maastrichtian age of the Late Cretaceous Period (geology), period, about 68 to 66 million years ago on the island continent of Laramidia, now forming western North America. It was one of the last-known non-avian dinosaurs and lived until the Cretaceous-Paleogene extinction event, Cretaceous–Paleogene extinction event 66 million years ago. The name ''Triceratops'', which means 'three-horned face', is derived from the Ancient Greek, Greek words () meaning 'three', () meaning 'horn', and () meaning 'face'. Bearing a large bony neck frill, frill, three horns on the skull, and a large, four-legged body, exhibiting convergent evolution with bovines and rhinoceroses, ''Triceratops'' is one of the most recognizable of all dinosaurs and the best-known ceratopsian. It was also one of the largest, measuring around long and weighing up to . It shared the la ...
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Stem-based Taxon
Phylogenetic nomenclature is a method of nomenclature for taxon, taxa in biology that uses phylogenetics, phylogenetic definitions for taxon names as explained below. This contrasts with Biological classification, the traditional method, by which taxon names are defined by a ''Type (biology), type'', which can be a specimen or a taxon of lower Taxonomic rank, rank, and a description in words. Phylogenetic nomenclature is regulated currently by the ''PhyloCode, International Code of Phylogenetic Nomenclature'' (''PhyloCode''). Definitions Phylogenetic nomenclature associates names with clades, groups consisting of an ancestor and all its descendants. Such groups are said to be Monophyly, monophyletic. There are slightly different methods of specifying the ancestor, which are discussed below. Once the ancestor is specified, the meaning of the name is fixed: the ancestor and all organisms which are its descendants are included in the taxon named. Listing all these organisms (i.e. prov ...
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Ceratopsidae
Ceratopsidae (sometimes spelled Ceratopidae) is a family of ceratopsian dinosaurs including ''Triceratops'', ''Centrosaurus'', and ''Styracosaurus''. All known species were quadrupedal herbivores from the Upper Cretaceous. All but one species are known from western North America, which formed the island continent of Laramidia during most of the Late Cretaceous. Ceratopsids are characterized by beaks, rows of shearing teeth in the back of the jaw, elaborate nasal horns, and a thin parietal-squamosal shelf that extends back and up into a frill. The group is divided into two subfamilies—Chasmosaurinae and Centrosaurinae. The chasmosaurines are generally characterized by long, triangular frills and well-developed brow horns. The centrosaurines had well-developed nasal horns or nasal bosses, shorter and more rectangular frills, and elaborate spines on the back of the frill. These horns and frills show remarkable variation and are the principal means by which the various species have be ...
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Protoceratopsidae
Protoceratopsidae is a family of basal (primitive) ceratopsians from the Late Cretaceous period. Although ceratopsians have been found all over the world, protoceratopsids are only definitively known from Cretaceous strata in Asia, with most specimens found in China and Mongolia. As ceratopsians, protoceratopsids were herbivorous, with constantly replacing tooth batteries made for slicing through plants and a hooked beak for grabbing them. Protoceratopsids were small ceratopsians around in length. Their bony frill and horns were much smaller than more derived members of Ceratopsia, such as ceratopsids. Description Protoceratopsids were relatively small ceratopsians, averaging around 1-2.5 m in length from head to tail. Protoceratopsids have a frill and rostral bone characteristic of all ceratopsians. Their snout is particularly wedge-shaped with tall and narrow nostrils situated high on it. The antorbital fenestra is unusually small, and the antorbital fossa sits high on the sku ...
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