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Habelia
''Habelia'' is a genus of extinct arthropod from the Middle Cambrian, thought to be one of the earliest known relatives of chelicerates. Its fossils have been found in the Burgess Shale in British Columbia, Canada. Fifty-four specimens of ''Habelia'' are known from the Greater Phyllopod bed, where they comprise 0.1% of the community. Description The body is divided into two main segments. The cephalon ("head")/prosoma with seven segments/8 somites, and the trunk with 12 segments, making for a total of 19 body segments. The body is around in length excluding the long telson that terminates the body. The head, which is 25% wider than it is long, makes up about 30% of the total body length, and has a bulging shape, with triangular genal spines projecting outward at its edges. It housed the stomach of the animal. The head bore a pair of spherical compound eyes, which were accommodated by notches in the head shield. A hypostome- labrum complex was present, with the mouth openling v ...
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Habeliida
Habeliida is an order of extinct arthropods that existed during the middle Cambrian. It is divided into two families, Habeliidae (monotypic, containing only ''Habelia''), and Sanctacarididae (containing '' Sanctacaris'', '' Utahcaris'' and '' Wisangocaris''). They are thought to have been durophagous, with robust gnathobases (spined basal sections of limbs) used to shred hard-shelled organisms. Remains of trilobites have been found as stomach contents in ''Wisangocaris''. '' Messorocaris'' has been suggested to be part of the order in some studies, but this is uncertain. They are suggested to be stem-group chelicerates, though they lack the chelicerae The chelicerae () are the arthropod mouthparts, mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated ... present in true chelicerates. References Chelicerate orders Prehistoric ch ...
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Sanctacaris
''Sanctacaris'' is a Middle Cambrian arthropod from the Burgess Shale of British Columbia. It was most famously regarded as a stem-group chelicerate, a group which includes horseshoe crabs, spiders and scorpions, although subsequent phylogenetic studies have not always supported this conclusion. Its chelicerate affinities regain support in later observations, alongside the reassignment of ''Habelia optata'' as a sanctacaridid-related basal chelicerate. It has been placed as a member of the extinct family Sanctacarididae alongside '' Wisangocaris'' and '' Utahcaris''. ''Sanctacaris'' specimens range from 46 to 93 mm in length. The head bears five pairs of grasping appendages (corresponding to chelicerate's pedipalps and walking legs) and a sixth pair of posterior appendages (correspond to horseshoe crab's chilaria). The grasping appendages each bear an antenna-like exopods. There are 11 body segments, with the former 10 each bearing a pair of biramous appendages with flap-lik ...
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Arachnomorpha
Arachnomorpha is a proposed subdivision or clade of Arthropoda, comprising the group formed by the trilobites and their close relatives (Artiopoda), Megacheira (which may be paraphyletic) and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered the sister group to Mandibulata (including insects, crustaceans and myriapods). The arachnomorph concept has been challenged by suggestions that the artiopods are more closely related to mandibulates, forming the clade Antennulata instead. There is no consensus as to assigning Arachnomorpha a formal Linnean rank. Classification Arachnomorpha Lameere, 1890 Arachnata Paulus, 1979, = Palaeopoda Packard, 1903*†'' Nettapezoura'' *†'' Dicranocaris'' *† Trilobita Walch, 1771 *† Megacheira Hou & Bergstrom, 1997 *† Strabopida Hou & Bergström, 1997 *† Aglaspida Walcott, 1911 *† Cheloniellida Broili, 1932 *Chelicerata Heymons 1901 **Family † Sanctacarididae Legg & Pates, 2017 **Class Pycnogoni ...
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Hypostome (trilobite)
The hypostome is the hard mouthpart of trilobites found on the ventral side of the cephalon (head). The hypostome can be classified into three types based on whether they are permanently attached to the rostrum or not and whether they are aligned to the anterior dorsal tip of the glabella. Morphology The center of the hypostome is an ovoid, typically convex part called the median body, often divided into an anterior lobe and a posterior lobe. Either side of the median body is a border with various extensions, including anterior and posterior wings, sometimes bearing knob-like processes. The hypostome is hollow, and encloses the mouthparts, the anterior digestive tract, and the bases of the antennae. Trilobite antennae pass through notches between the anterior and posterior wings, then forward. The anterior wings are designed to rest firmly against internal structures (ventral apodemes) on the glabella. Variation in trilobite hypostome morphology is crucial in modern discussi ...
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Chelicerata
The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and '' Mollisonia'' present by this time. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered in a cuti ...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown ...
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Wisangocaris
''Wisangocaris'' is an extinct genus of Cambrian arthropod known from the Emu Bay Shale of Australia. Due to the fact that trilobite fragments have been found in its stomach, it was probably durophagous. It been placed in the family Sanctacarididae alongside ''Sanctacaris'' as stem-group chelicerates The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, tic .... References Prehistoric chelicerate genera Fossil taxa described in 2016 {{paleo-arthropod-stub ...
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Sexual Dimorphism
Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, dioecious species, which consist of most animals and some plants. Differences may include secondary sex characteristics, size, weight, color, markings, or behavioral or cognitive traits. Male-male reproductive competition has evolved a diverse array of sexually dimorphic traits. Aggressive utility traits such as "battle" teeth and blunt heads reinforced as battering rams are used as weapons in aggressive interactions between rivals. Passive displays such as ornamental feathering or song-calling have also evolved mainly through sexual selection. These differences may be subtle or exaggerated and may be subjected to sexual selection and natural selection. The opposite of dimorphism is ''monomorphism'', when both biological sexes are phenotype, ...
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Exopod
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip (anatomy), hip, : ''coxae''), ''trochanter'', ''femur'' (: ''femora''), ''tibia'' (: ''tibiae''), ''tarsus'' (: ''tarsi''), ''ischium'' (: ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (: ''patellae''). Homology (biology), Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of Neontology, extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a Homeobox, ''Hox''-gene, could result in Parallel evolution, parallel gains of leg segments ...
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Seta
In biology, setae (; seta ; ) are any of a number of different bristle- or hair-like structures on living organisms. Animal setae Protostomes Depending partly on their form and function, protostome setae may be called macrotrichia, chaetae, Scale (insect anatomy), scales, or Common name, informally, hairs. The setal membrane is not Cuticle, cuticularized, so movement is possible. Annelid setae are stiff bristles present on the body. They allow earthworms and their relatives to attach to the surface and prevent backsliding during peristaltic motion. These hairs make it difficult to pull a worm straight from the ground. Setae in oligochaetes (the group including earthworms) are largely composed of chitin. They are classified according to the limb to which they are attached; for instance, notosetae are attached to notopodia; neurosetae to neuropodia. The setae on polychaete worms are referred to as chaeta due to their differing morphology. Crustaceans have mechano- and chemos ...
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