Arachnomorpha is a proposed subdivision or
clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
of
Arthropod
Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
a, comprising the group formed by the
trilobites
Trilobites (; meaning "three-lobed entities") are extinct marine arthropods that form the class Trilobita. One of the earliest groups of arthropods to appear in the fossil record, trilobites were among the most successful of all early animals, ...
and their close relatives (
Artiopoda
Artiopoda is a clade of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomor ...
),
Megacheira (which may be
paraphyletic
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In co ...
) and
chelicerates
The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, tic ...
. Under this proposed classification scheme, Arachnomorpha is considered the sister group to
Mandibulata
The clade Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes, among others), and all true insects. The name "Mandibul ...
(including
insects
Insects (from Latin ') are hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed ...
,
crustaceans
Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of Arthropod, arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquat ...
and
myriapods
Myriapods () are the members of subphylum Myriapoda, containing arthropods such as millipedes and centipedes. The group contains about 13,000 species, all of them terrestrial.
Although molecular evidence and similar fossils suggests a diversifi ...
).
The arachnomorph concept has been challenged by suggestions that the artiopods are more closely related to
mandibulates, forming the clade
Antennulata instead.
There is no consensus as to assigning Arachnomorpha a formal
Linnean rank.
Classification
Arachnomorpha
Lameere, 1890 Paulus, 1979, = Palaeopoda Packard, 1903"> Arachnata Paulus, 1979, = Palaeopoda Packard, 1903*†''
Nettapezoura''
*†''
Dicranocaris''
*†
Trilobita Walch, 1771
*†
Megacheira Hou & Bergstrom, 1997
*†
Strabopida Hou & Bergström, 1997
*†
Aglaspida Walcott, 1911
*†
Cheloniellida Broili, 1932
*
Chelicerata
The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, tic ...
Heymons 1901
**Family †
Sanctacarididae Legg & Pates, 2017
**Class
Pycnogonida
Sea spiders are marine arthropods of the class Pycnogonida, hence they are also called pycnogonids (; named after ''Pycnogonum'', the type genus; with the suffix '). The class includes the only now-living order Pantopoda ( ‘all feet’), alongs ...
Latreille, 1810
**Clade
Euchelicerata Weygoldt & Paulus, 1979
***Family †
Offacolidae Sutton ''et al''., 2002
***Clade
Prosomapoda
Prosomapoda is a clade of Chelicerata, euchelicerates including the groups Xiphosura (horseshoe crabs) and Planaterga (a group comprising Bunodidae, bunodids, Pseudoniscidae, pseudoniscids, Chasmataspidida, chasmataspidids, Eurypterida, eurypteri ...
Lamsdell, 2013
Phylogeny
Using fossil data, Bergström & Hou (2003) gave an outline of arthropod relationships emphasizing trilobitomorphs (a group that includes trilobites and trilobite-like animals).
References
Arthropod taxonomy
Taxa named by Auguste Lameere
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