Cladoradula Auriculata
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Cladoradula Auriculata
''Cladoradula'' is a genus of liverworts in the family Radulaceae. Distinguished by its thick, brown-pigmented plant stem, stems and distinctive branching pattern, it comprises seven species found primarily in tropical and temperate forest regions worldwide. Originally established as a subgenus of ''Radula'' in 1885, it was elevated to genus rank in 2022 following molecular phylogenetics, molecular studies that revealed it represents one of the oldest lineage (evolution), lineages within Radulaceae, having diverged during the late Permian period about 263 million years ago. The genus is characterised by its specialised plant stem, stem structure, distinctive leaf arrangement, and small protective structures around its reproductive organs. Species in the genus grow on tree bark or shaded rocks from sea level to over in elevation. Taxonomy ''Cladoradula'' was originally established as a subgenus of ''Radula'' by the English bryologist Richard Spruce in 1885, with ''Radula gottsc ...
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Richard Spruce
Richard Spruce (10 September 1817 – 28 December 1893) was an English botanist specializing in bryology. One of the great Victorian botanical explorers, Spruce spent 15 years exploring the Amazon from the Andes to its mouth, and was one of the very first Europeans to observe many of the places where he collected specimens. Spruce discovered and named a number of new plant species, and corresponded with some of the leading botanists of the nineteenth century. Early life and career Richard Spruce was born near Ganthorpe, a small village near Castle Howard in Yorkshire. After training under his father, a local schoolmaster, Spruce began a career as a tutor and then as a mathematics master at St Peter's School in York between 1839 and 1844. Spruce started his botanical collecting in Yorkshire about 1833. In 1834, at age 16, he drew up a neatly written list of all of the plants he had found on trips around Ganthorpe, focusing on bryophytes. Arranged alphabetically and contain ...
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Molecular Phylogenetics
Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to determine the processes by which diversity among species has been achieved. The result of a molecular phylogenetics, phylogenetic analysis is expressed in a phylogenetic tree. Molecular phylogenetics is one aspect of molecular systematics, a broader term that also includes the use of molecular data in Taxonomy (biology), taxonomy and biogeography. Molecular phylogenetics and molecular evolution correlate. Molecular evolution is the process of selective changes (mutations) at a molecular level (genes, proteins, etc.) throughout various branches in the tree of life (evolution). Molecular phylogenetics makes inferences of the evolutionary relationships that arise due to molecular evolution and results in the construction of a phylogenetic tre ...
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Androecia
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament and an anther which contains microsporangia. Most commonly, anthers are two-lobed (each lobe is termed a locule) and are attached to the filament either at the base or in the middle area of the anther. The sterile (i.e. nonreproductive) tissue between the lobes is called the connective, an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte. The size of anthers differs greatly, from a tiny fraction of a millimeter in ''Wolfia'' spp up to five inches (13 centimeters) in '' Canna iridiflora'' and ''Strelitzia nicolai''. The stamens in a flower are collectively called the an ...
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Dioicous
Dioicy () is a sexual system in non-vascular plants where archegonia (female organs) and antheridia (male organs) are produced on separate plants in the gametophyte phase. It is one of the two main sexual systems in bryophytes, the other being monoicy. Both dioicous () and monoicous gametophytes produce gametes in gametangia by mitosis rather than meiosis, so that sperm and eggs are genetically identical with their parent gametophyte. Description Dioicy promotes outcrossing. Sexual dimorphism is commonly found in dioicous species. Dioicy is correlated with reduced sporophyte production, due to spatial separation of male and female colonies, scarcity or absence of males. The term dioecy is inapplicable to bryophytes because it refers to the sexuality of vascular plant sporophytes. Nonetheless dioecy and dioicy are comparable in many respects. Etymology The words dioicous and di(o)ecious are derived from οἶκος or οἰκία and δι- (di-), twice, double. (''(o)e' ...
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Auricle (botany)
In botany, an auricle is a small ear-like projection from the base of a leaf or petal Petals are modified leaves that form an inner whorl surrounding the reproductive parts of flowers. They are often brightly coloured or unusually shaped to attract pollinators. All of the petals of a flower are collectively known as the ''corol .... References External linksDictionary of Botany Leaf morphology {{plant-morphology-stub ...
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Micrometre
The micrometre (English in the Commonwealth of Nations, Commonwealth English as used by the International Bureau of Weights and Measures; SI symbol: μm) or micrometer (American English), also commonly known by the non-SI term micron, is a unit of length in the International System of Units (SI) equalling (SI standard prefix "micro-" = ); that is, one millionth of a metre (or one thousandth of a millimetre, , or about ). The nearest smaller common SI Unit, SI unit is the nanometre, equivalent to one thousandth of a micrometre, one millionth of a millimetre or one billionth of a metre (). The micrometre is a common unit of measurement for wavelengths of infrared radiation as well as sizes of biological cell (biology), cells and bacteria, and for grading wool by the diameter of the fibres. The width of a single human hair ranges from approximately 20 to . Examples Between 1 μm and 10 μm: * 1–10 μm – length of a typical bacterium * 3–8 μm – width of str ...
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Perianth
The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepals when called a perigone. The term ''perianth'' is derived from Greek περί (, "around") and άνθος (, "flower"), while ''perigonium'' is derived from περί () and γόνος (, "seed, sex organs"). In the mosses and liverworts (Marchantiophyta), the perianth is the sterile (neither male nor female) tubelike tissue that surrounds the female reproductive structure or developing sporophyte. Flowering plants In flowering plants, the perianth may be described as being either dichlamydeous/heterochlamydeous in which the calyx and corolla are clearly separate, or homochlamydeous, in which they are indistinguishable (and the sepals and petals are collectively referred to as tepals). When the perianth is in two whorls, it is desc ...
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Cortex (botany)
In botany, a cortex is an outer layer of a stem or root in a vascular plant, lying below the epidermis but outside of the vascular bundles. The cortex is composed mostly of large thin-walled parenchyma cells of the ground tissue system and shows little to no structural differentiation. The outer cortical cells often acquire irregularly thickened cell walls, and are called collenchyma cells. Plants Stems and branches In the three dimensional structure of herbaceous stems, the epidermis, cortex and vascular cambium form concentric cylinders around the inner cylindrical core of pith. Some of the outer cortical cells may contain chloroplasts, giving them a green color. They can therefore produce simple carbohydrates through photosynthesis. In woody plants, the cortex is located between the periderm (bark) and the vascular tissue ( phloem, in particular). It is responsible for the transportation of materials into the central cylinder of the root through diffusion and ma ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxono ...
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Chloroplast
A chloroplast () is a type of membrane-bound organelle, organelle known as a plastid that conducts photosynthesis mostly in plant cell, plant and algae, algal cells. Chloroplasts have a high concentration of chlorophyll pigments which capture the Radiant energy, energy from sunlight and convert it to chemical energy and release oxygen. The chemical energy created is then used to make sugar and other organic molecules from carbon dioxide in a process called the Calvin cycle. Chloroplasts carry out a number of other functions, including fatty acid synthesis, amino acid synthesis, and the immune response in plants. The number of chloroplasts per cell varies from one, in some unicellular algae, up to 100 in plants like ''Arabidopsis'' and wheat. Chloroplasts are highly dynamic—they circulate and are moved around within cells. Their behavior is strongly influenced by environmental factors like light color and intensity. Chloroplasts cannot be made anew by the plant cell and must ...
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Porella
''Porella'' is a large, common, and widespread genus of liverworts in order Porellales. It is a member of the family Porellaceae within that order. There were 84 species recognized in 2016, most of them from East Asia. World Flora Online accepts 125 species, and GBIF accepts 150 species (as of 4 July 2023). The genus has a wide cosmopolitan distribution In biogeography, a cosmopolitan distribution is the range of a taxon that extends across most or all of the surface of the Earth, in appropriate habitats; most cosmopolitan species are known to be highly adaptable to a range of climatic and en ... in temperate areas, where it is commonly found growing attached to the bark of trees. Members of the genus have lobed leaves with a large upper lobe and small lower lobe. Species ''Porella arboris-vitae'', ''Porella bolanderi'', ''Porella cordeana'', ''Porella navicularis'', ''Porella pinnata'', ''Porella platyphylla'' and ''Porella roellii'', were all found in Montana, USA. ...
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Eustace Wilkinson Jones
Eustace Wilkinson Jones (6 June 1909 – 21 August 1992) was a forester and bryologist whose expertise on African hepatics is widely recognized for its accuracy and attention to detail. His contributions to bryology were made in his leisure hours since he spent most of his career teaching and conducting research on forestry. It was only after he retired that he could devote himself entirely to bryology. Life and career Born in Walsall in Staffordshire, Eustace showed an interest in field botany at a young age. Educated in Walsall, he entered Jesus College, Cambridge in 1928. There he earned a first-class honours degree in Natural Sciences Tripos, and then went on to pursue a Ph.D. on the ionic relationships of ''Cladophora'', graduating with this degree in 1937. He joined the Forestry School at Oxford in 1934 as a teacher, where he remained until his retirement in 1972. His work on the structure and reproduction of the virgin forest of the North Temperate Zone and Nigerian fo ...
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