In
phylogenetics
In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an
evolutionary tree.
Definition
The expression is most easily illustrated by a
cladogram
A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
:
Taxon
In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
A and taxon B are sister groups to each other. Taxa A and B, together with any other
extant or
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
descendants of their
most recent common ancestor (MRCA), form a
monophyletic
In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria:
# the grouping contains its own most recent co ...
group, the
clade
In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...
AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC.
The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the
leaves
A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
and among larger, more deeply
rooted clades. The tree structure shown connects through its root to the rest of the
universal tree of life.
In
cladistic standards, taxa A, B, and C may represent specimens,
species
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
,
genera, or any other taxonomic units. If A and B are at the same taxonomic level, terminology such as ''sister species'' or ''sister genera'' can be used.
Example
The term ''sister group'' is used in
phylogenetic analysis, however, only groups identified in the analysis are labeled as "sister groups".
An example is
birds, whose commonly cited ''living'' sister group is the
crocodiles, but that is true only when discussing
extant organisms; when other, extinct groups are considered, the relationship between birds and crocodiles appears distant.
Although the bird family tree is rooted in the
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s, there were a number of other, earlier groups, such as the
pterosaurs, that branched off the line leading to the dinosaurs after the last common ancestor of
birds and crocodiles.
The term ''sister group'' must thus be seen as a relative term, with the caveat that the sister group is only the closest relative among the groups/species/specimens that are included in the analysis.
Notes
References
{{Phylogenetics
Evolutionary biology
Phylogenetics
Biological classification
Taxa