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Calyptapis
''Calyptapis'' is an extinct bombini genus related to bumblebees with one described species ''Calyptapis florissantensis''. It is known only from the Late Eocene Chadronian age shales of the Florissant Formation in Colorado. the genus and species were described by Theodore Dru Alison Cockerell in 1906. References † A dagger, obelisk, or obelus is a typographical mark that usually indicates a footnote if an asterisk has already been used. The symbol is also used to indicate death (of people) or extinction (of species). It is one of the modern descendan ... Eocene insects Fossil bee taxa Fossil taxa described in 1906 Insects described in 1906 Prehistoric insects of North America Fossil bee genera {{paleo-insect-stub ...
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Bumblebee
A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related genera (e.g., '' Calyptapis'') are known from fossils. They are found primarily in higher altitudes or latitudes in the Northern Hemisphere, although they are also found in South America, where a few lowland tropical species have been identified. European bumblebees have also been introduced to New Zealand and Tasmania. Female bumblebees can sting repeatedly, but generally ignore humans and other animals. Most bumblebees are social insects that form colonies with a single queen. The colonies are smaller than those of honey bees, growing to as few as 50 individuals in a nest. Cuckoo bumblebees are brood parasitic and do not make nests or form colonies; their queens aggressively invade the nests of other bumblebee species, kill the resident q ...
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Bumblebees
A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only extant group in the tribe Bombini, though a few extinct related genera (e.g., ''Calyptapis'') are known from fossils. They are found primarily in higher altitudes or latitudes in the Northern Hemisphere, although they are also found in South America, where a few lowland tropical species have been identified. European bumblebees have also been introduced to New Zealand and Tasmania. Female bumblebees can sting repeatedly, but generally ignore humans and other animals. Most bumblebees are social insects that form colonies with a single queen. The colonies are smaller than those of honey bees, growing to as few as 50 individuals in a nest. Cuckoo bumblebees are brood parasitic and do not make nests or form colonies; their queens aggressively invade the nests of other bumblebee species, kill the resident que ...
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Bombini
The Bombini are a tribe of large bristly apid bees which feed on pollen or nectar. Many species are social, forming nests of up to a few hundred individuals; other species, formerly classified as '' Psithyrus'' cuckoo bees, are brood parasites of nest-making species. The tribe contains a single living genus, '' Bombus'', the bumblebees, and some extinct genera such as '' Calyptapis'' and '' Oligobombus''. The tribe was described by Pierre André Latreille in 1802. Fossils '' Bombus cerdanyensis'' was described from Late Miocene lacustrine beds of La Cerdanya, Spain in 2014. '' Calyptapis florissantensis'' was described by Theodore Dru Alison Cockerell in 1906 from the Chadronian ( Eocene) lacustrine – large shale of Florissant in the US. '' Oligobombus cuspidatus'' was described by Antropov ''et al'' (2014) from the Late Eocene Insect Bed of the Bembridge Marls on the Isle of Wight, England. The holotype fossil was described by re-examining a specimen in the Smith ...
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Chadronian
The Chadronian age within the North American Land Mammal Ages chronology is the North American faunal stage typically set from 38,000,000 to 33,900,000 years BP, a period of . It is usually considered to fall within the Eocene epoch. The Chadronian is preceded by the Duchesnean and followed by the Orellan NALMA stages. The Chadronian can be further divided into the substages of: *Late/Upper Chadronian (shares upper boundary). Lower boundary source of the base of the Priabonian (approximate) *Middle Chadronian. Lower boundary source and base of the Priabonian (approximate). Upper boundary source of the base of the Orellan (approximate). *Early/Lower Chadronian (shares lower boundary). Upper boundary source: base of Orellan (approximate). Geological time The Chadronian maintains a period of time within the Priabonian through Rupelian of the Late Eocene through Early Oligocene in the geologic time scale The geologic time scale, or geological time scale, (GTS) is a representati ...
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Theodore Dru Alison Cockerell
Theodore Dru Alison Cockerell (1866–1948) was an American zoologist, born at Norwood, England, and brother of Sydney Cockerell. He was educated at the Middlesex Hospital Medical School, and then studied botany in the field in Colorado in 1887–90. Subsequently, he became a taxonomist and published numerous papers on the Hymenoptera, Hemiptera, Mollusca and plants, as well as publications on paleontology and evolution. Personal life Cockerell was born in Norwood, Greater London and died in San Diego, California. He married Annie Sarah Fenn in 1891 (she died in 1893) and Wilmatte Porter in 1900. In 1901, he named the ultramarine blue chromodorid ''Mexichromis porterae'' (now '' Felimare porterae'') in her honor. After their marriage in 1900, they frequently went on collecting expeditions together and assembled a large private library of natural history films, which they showed to schoolchildren and public audiences to promote the cause of environmental conservation. After h ...
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Late Eocene
The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope 13C in the atmosphere was exceptionally low in comparison with the more common isotope 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Siberia and in what is now Chesapeake Bay. As with other geologic periods, the strata that define the start and end ...
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Shale
Shale is a fine-grained, clastic sedimentary rock formed from mud that is a mix of flakes of clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4) and tiny fragments (silt-sized particles) of other minerals, especially quartz and calcite.Blatt, Harvey and Robert J. Tracy (1996) ''Petrology: Igneous, Sedimentary and Metamorphic'', 2nd ed., Freeman, pp. 281–292 Shale is characterized by its tendency to split into thin layers ( laminae) less than one centimeter in thickness. This property is called '' fissility''. Shale is the most common sedimentary rock. The term ''shale'' is sometimes applied more broadly, as essentially a synonym for mudrock, rather than in the more narrow sense of clay-rich fissile mudrock. Texture Shale typically exhibits varying degrees of fissility. Because of the parallel orientation of clay mineral flakes in shale, it breaks into thin layers, often splintery and usually parallel to the otherwise indistinguishable bed ...
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Florissant Formation
The Florissant Formation is a sedimentary geologic formation outcropping around Florissant, Teller County, Colorado. The formation is noted for the abundant and exceptionally preserved insect and plant fossils that are found in the mudstones and shales. Based on argon radiometric dating, the formation is Eocene (approximately 34 million years old ) in age and has been interpreted as a lake environment. The fossils have been preserved because of the interaction of the volcanic ash from the nearby Thirtynine Mile volcanic field with diatoms in the lake, causing a diatom bloom. As the diatoms fell to the bottom of the lake, any plants or animals that had recently died were preserved by the diatom falls. Fine layers of clays and muds interspersed with layers of ash form "paper shales" holding beautifully-preserved fossils. The Florissant Fossil Beds National Monument is a national monument established to preserve and study the geology and history of the area. History The name ''Fl ...
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Eocene Insects
The Eocene ( ) Epoch is a geological epoch that lasted from about 56 to 33.9 million years ago (mya). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''ēṓs'', "dawn") and (''kainós'', "new") and refers to the "dawn" of modern ('new') fauna that appeared during the epoch. The Eocene spans the time from the end of the Paleocene Epoch to the beginning of the Oligocene Epoch. The start of the Eocene is marked by a brief period in which the concentration of the carbon isotope 13C in the atmosphere was exceptionally low in comparison with the more common isotope 12C. The end is set at a major extinction event called the ''Grande Coupure'' (the "Great Break" in continuity) or the Eocene–Oligocene extinction event, which may be related to the impact of one or more large bolides in Siberia and in what is now Chesapeake Bay. As with other geologic periods, the strata that define the start and end of th ...
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Fossil Bee Taxa
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolute ...
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Insects Described In 1906
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch from eggs. Insect ...
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