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Amborellaceae
''Amborella'' is a monotypic genus of understory shrubs or small trees endemic to the main island, Grande Terre, of New Caledonia in the southwest Pacific Ocean. The genus is the only member of the family Amborellaceae and the order Amborellales and contains a single species, ''Amborella trichopoda''. ''Amborella'' is of great interest to plant systematists because molecular phylogenetic analyses consistently place it as the sister group to all other flowering plants, meaning it was the earliest group to evolve separately from all other flowering plants. Description ''Amborella'' is a sprawling shrub or small tree up to high. It bears alternate, simple evergreen leaves without stipules. The leaves are two-ranked, with distinctly serrated or rippled margins, and about long. ''Amborella'' has xylem tissue that differs from that of most other flowering plants. The xylem of ''Amborella'' contains only tracheids; vessel elements are absent. Xylem of this form has long been regard ...
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Monotypic
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described. Theoretical implications Monotypic taxa present several important theoretical challenges in biological classification. One key issue is known as "Gregg's Paradox": if a single species is the only member of multiple hierarchical levels (for example, being the only species in its genus, which is the only genus in its family), then each level needs a distinct definition to maintain logical structure. Otherwise, the different taxonomic ranks become effectively identical, which creates problems for organizing biological diversity in a hierarchical syste ...
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Vessel Element
A vessel element or vessel member (also called a xylem vessel) is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are found in most angiosperms (flowering plants) and in some gymnosperms such as cycads and ''Ephedra (plant), Ephedra'', but absent in conifers. Vessel elements are the main feature distinguishing the "hardwood" of angiosperms from the "softwood" of conifers. Anatomy Xylem is the tissue in vascular plants that conducts water (and substances dissolved in it) upwards from the roots to the shoots. Two kinds of cell are involved in xylem transport: tracheids and vessel elements. Vessel elements are the building blocks of vessels, the conducting pathways that constitute the major part of the water transporting system in flowering plants. Vessels form an efficient system for transporting water (including necessary minerals) from the root to the leaves and other parts of the plant. In secondary xylem – the xylem that is prod ...
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Xylem
Xylem is one of the two types of transport tissue (biology), tissue in vascular plants, the other being phloem; both of these are part of the vascular bundle. The basic function of the xylem is to transport water upward from the roots to parts of the plants such as stems and leaves, but it also transports plant nutrition, nutrients. The word ''xylem'' is derived from the Ancient Greek word (), meaning "wood"; the best-known wood organism is plants, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cell (biology), cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Wood also contains two other type of cells: Ground tissue#Parenchyma, parenchyma and ground tissue#Fibres, fibers. Xylem can be found: * in vascular bundles, present in ...
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Aleksander Pavlovich Melikyan
Alexander () is a male given name, name of Greek origin. The most prominent bearer of the name is Alexander the Great, the king of the Ancient Greek kingdom of Macedonia (ancient kingdom), Macedonia who created one of the largest empires in ancient history. Variants listed here are Aleksandar, Aleksander, Oleksandr, Oleksander, Aleksandr, and Alekzandr. Related names and diminutives include Iskandar (name), Iskandar, Alec, Alek, Alex, Alexsander, Alexandre (given name), Alexandre, Aleks (given name), Aleks, Aleksa (given name), Aleksa, Aleksandre, Alejandro, Alessandro, Alasdair, Sasha (name), Sasha, Sandy (given name), Sandy, Sandro, Sikandar (other), Sikandar, Skander, Sander (name), Sander and Xander; feminine forms include Alexandra, Alexandria (given name), Alexandria, and Sasha (name), Sasha. Etymology The name ''Alexander'' originates from the (; 'defending men' or 'protector of men'). It is a compound of the verb (; 'to ward off, avert, defend') and the no ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta. Angiosperms are by far the most diverse group of Embryophyte, land plants with 64 Order (biology), orders, 416 Family (biology), families, approximately 13,000 known Genus, genera and 300,000 known species. They include all forbs (flowering plants without a woody Plant stem, stem), grasses and grass-like plants, a vast majority of broad-leaved trees, shrubs and vines, and most aquatic plants. Angiosperms are distinguished from the other major seed plant clade, the gymnosperms, by having flowers, xylem consisting of vessel elements instead of tracheids, endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from the commo ...
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Pollen
Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male Conifer cone, cone to the female cone of gymnosperms. If pollen lands on a compatible pistil or female cone, it Germination, germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require magnification to see detail. The study of pollen is called palynology and is highly useful in paleoecology, paleontology, archaeology, and Forensic science, forensics. Pollen in plants is used for transferring Ploidy#Haploid and monoploid, haploid male genetic ma ...
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Staminode
In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co. London, 4th ed 1928 Staminodes are frequently inconspicuous and stamen-like, usually occurring at the inner whorl of the flower, but are also sometimes long enough to protrude from the corolla. Sometimes, the staminodes are modified to produce nectar, as in the witch-hazel ''(Hamamelis)''.jin lu mei shu. Hamamelis Linnaeus, Sp. Pl. 1: 124. 1753. Flora of China 9: 32. 2003 Staminodes can be a critical characteristic for differentiating between species, for instance in the orchid genus ''Paphiopedilum'', and among the penstemons. In the case of cannas, the petals are inconsequential and the staminodes are refined into eye-catching petal-like replacements. A spectacular example of staminode is given by ''Couroupita guianensis ''C ...
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Carpel
Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils'' and is typically surrounded by the pollen-producing reproductive organs, the stamens, collectively called the androecium. The gynoecium is often referred to as the "female" portion of the flower, although rather than directly producing female gametes (i.e. egg cells), the gynoecium produces megaspores, each of which develops into a female gametophyte which then produces egg cells. The term gynoecium is also used by botanists to refer to a cluster of archegonia and any associated modified leaves or stems present on a gametophyte shoot in mosses, liverworts, and hornworts. The corresponding terms for the male parts of those plants are clusters of antheridia within the androecium. Flowers that bear a gynoecium but no stame ...
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Stamen
The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament and an anther which contains sporangium, microsporangia. Most commonly, anthers are two-lobed (each lobe is termed a locule) and are attached to the filament either at the base or in the middle area of the anther. The sterile (i.e. nonreproductive) tissue between the lobes is called the Connective (botany), connective, an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte. The size of anthers differs greatly, from a tiny fraction of a millimeter in ''Wolfia'' spp up to five inches (13 centimeters) in ''Canna iridiflora'' and ''Strelitzia nicolai''. The stamens in a flower ...
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Dioecious
Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is biparental reproduction. Dioecy has costs, since only the female part of the population directly produces offspring. It is one method for excluding self-fertilization and promoting allogamy (outcrossing), and thus tends to reduce the expression of recessive deleterious mutations present in a population. Plants have several other methods of preventing self-fertilization including, for example, dichogamy, herkogamy, and self-incompatibility. In zoology In zoology, dioecy means that an animal is either male or female, in which case the synonym gonochory is more often used. Most animal species are gonochoric, almost all vertebrate species are gonochoric, and all bird and mammal species are gonochoric. Dioecy may also describe colonies ...
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Primitive (phylogenetics)
In phylogenetics, a primitive (or ancestral) character, trait, or feature of a lineage or taxon is one that is inherited from the common ancestor of a clade (or clade group) and has undergone little change since. Conversely, a trait that appears ''within'' the clade group (that is, is present in any subgroup within the clade but not all) is called advanced or derived. A clade is a group of organisms that consists of a common ancestor and all its lineal descendants. A primitive trait is the original condition of that trait in the common ancestor; advanced indicates a notable change from the original condition. These terms in biology contain no judgement about the sophistication, superiority, value or adaptiveness of the named trait. "Primitive" in biology means only that the character appeared first in the common ancestor of a clade group and has been passed on largely intact to more recent members of the clade. "Advanced" means the character has evolved within a later subgroup ...
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Tracheid
A tracheid is a long and tapered Lignin, lignified cell in the xylem of Tracheophyta, vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms also use another type of conductive cell, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to Transpiration, transport water and inorganic salts, and to provide structural support for trees. There are often Pit (botany), pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast. The wood (softwood) of gymnosperms such as pines and other conifers is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of ferns. The tracheid was first named by the German botanist Carl Gustav Sanio in 1863, from the German ''Tracheide''. Evolution Tracheids were the main conductive cells found in early vascular plants. In the first 140� ...
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