Acercostraca
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Acercostraca
Marrellomorpha are an extinct group of arthropods known from the Cambrian to the Early Devonian. They lacked mineralised hard parts, so are only known from areas of exceptional preservation, limiting their fossil distribution. The best known member is ''Marrella'', with thousands of specimens found in the Cambrian aged Burgess Shale of Canada. Taxonomy The group is divided up into two major orders, Marrellida and Acercostraca. Marrellida is recognised by the possession of head shields with two or three pairs of elongate spine-like projections, and three attached pairs of uniramous (single-branched) appendages, while Acercostraca generally have large ovoid carapaces that cover the entire upper half of the body, and five pairs of uniramous appendages attached to the head section. Both groups have unbranched antennae and a segmented trunk with biramous (divided into two branches) appendages. Recent research has suggested the previously enigmatic Cambrian trilobite-like arthropods ...
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Arachnomorpha
Arachnomorpha is a proposed subdivision or clade of Arthropoda, comprising the group formed by the trilobites and their close relatives (Artiopoda), Megacheira (which may be paraphyletic) and chelicerates. Under this proposed classification scheme, Arachnomorpha is considered the sister group to Mandibulata (including insects, crustaceans and myriapods). The arachnomorph concept has been challenged by suggestions that the artiopods are more closely related to mandibulates, forming the clade Antennulata instead. There is no consensus as to assigning Arachnomorpha a formal Linnean rank. Classification Arachnomorpha Lameere, 1890 Arachnata Paulus, 1979, = Palaeopoda Packard, 1903*†'' Nettapezoura'' *†'' Dicranocaris'' *† Trilobita Walch, 1771 *† Megacheira Hou & Bergstrom, 1997 *† Strabopida Hou & Bergström, 1997 *† Aglaspida Walcott, 1911 *† Cheloniellida Broili, 1932 *Chelicerata Heymons 1901 **Family † Sanctacarididae Legg & Pates, 2017 **Class Pycnogoni ...
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Primicaris
''Primicaris'' is genus of Cambrian arthropod from the Chengjiang biota of China and the Burgess Shale of Canada. It contains a single described species, ''P. larvaformis''. Description Specimens range from in length, and in maximum width. The undivided subovate-shaped dorsal shield covered the entire body, and wrapped around the front edge of the carapace, forming a doublure structure. The midline of the carapace exhibited thickening. There is a pair of slender uniramous segmented antennae, as well as ten pairs of biramous limbs. The exopods of these limbs are flagellate and bear setae, with the exopods decreasing in size posteriorly. The endopods are poorly preserved. Taxonomy Its specimens were once thought to be meraspids of ''Naraoia spinosa''. Although their phylogenetic position has been described as "problematic", recent studies have suggested them to be marrellomorphs belonging to the subgroup Acercostraca. See also * Arthropod * Cambrian explosion * List of Che ...
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Skania
''Skania'' is a Cambrian fossil arthropod. The type species, ''S. fragillis'', is known from the Burgess Shale of British Columbia, Canada. A second possible species ''"S." sundbergi'' is known from the Kaili Formation of China, but its placement within the genus has been questioned. Description Specimens of ''S. fragillis'' range in length from The cordiform dorsal shield/carapace covers the entire body, with its midline having a raised keel. The shield curls around the front to form a doublure, with the posterior edge of the doublure having a pair of lateral spines. A pair of spines is also present on the posterior of the dorsal shield. A hypostome was present on the underside of the head. There are two sets of appendages, the first 5 pairs of appendages are elongate, the first one of which is described as an antenna and has small subquadrate segments/podomeres, while the others have elongate subrectangular segments. These appendages are somewhat curved and extend beyo ...
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Middle Cambrian
Middle or The Middle may refer to: * Centre (geometry), the point equally distant from the outer limits. Places * Middle (sheading), a subdivision of the Isle of Man * Middle Bay (other) * Middle Brook (other) * Middle Creek (other) * Middle Island (other) * Middle Lake (other) * Middle Mountain, California * Middle Peninsula, Chesapeake Bay, Virginia * Middle Range, a former name of the Xueshan Range on Taiwan Island * Middle River (other) * Middle Rocks, two rocks at the eastern opening of the Straits of Singapore * Middle Sound, a bay in North Carolina * Middle Township (other) * Middle East Music *Middle (song), "Middle" (song), 2015 *The Middle (Jimmy Eat World song), "The Middle" (Jimmy Eat World song), 2001 *The Middle (Zedd, Maren Morris and Grey song), "The Middle" (Zedd, Maren Morris and Grey song), 2018 *"Middle", a song by Rocket from the Crypt from their 1995 album ''Scream, Dracula, Scream!'' *"The Middle ...
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Chelicerata
The subphylum Chelicerata (from Neo-Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. Chelicerates include the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and '' Mollisonia'' present by this time. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered in a cuti ...
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Stem Group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown ...
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Burgessia
''Burgessia'' is a genus of arthropod known from the mid-Cambrian aged Burgess Shale of British Columbia, Canada. It is relatively abundant, with over 1,700 specimens having been collected. Description The body had an unsegmented approximately circular carapace that was somewhat convex. It was likely thin and had only weak sclerotization. A pair of tapering flexible segmented antennae projected forwards from the head, which were about equal in length to the carapace, these were likely tactile in function. There are three pairs of cephalic appendages excluding the antennae that functioned as walking limbs, as well as seven pairs of biramous (two-branched) walking limbs with gills, otherwise similar to the cephalic limbs running along the trunk, which decreased in size posteriorly. Although not visible on any specimens, the mouth was almost certainly located on the underside of the body. The circular carapace was largely occupied by the guts, which were divided into two sect ...
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Sea Spider
Sea spiders are marine arthropods of the class (biology), class Pycnogonida, hence they are also called pycnogonids (; named after ''Pycnogonum'', the type genus; with the suffix '). The class includes the only now-living order (biology), order Pantopoda ( ‘all feet’), alongside a few fossil species which could trace back to the early or mid Paleozoic. They are cosmopolitan distribution, cosmopolitan, found in oceans around the world. The over 1,300 known species have leg spans ranging from to over . Most are toward the smaller end of this range in relatively shallow depths; however, they can grow to be quite large in Antarctic and deep-sea gigantism, deep waters. Despite their name and brief resemblance, "sea spiders" are not spiders, nor even arachnids. While some literature around the 2000s suggests they may be a sister group to all other living arthropods, their traditional classification as a member of chelicerates alongside Xiphosura, horseshoe crabs and arachnids has r ...
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Cladistical
Cladistics ( ; from Ancient Greek 'branch') is an approach to biological classification in which organisms are categorized in groups ("clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics (synapomorphies) that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a ' grade', which are fruitless to precisely delineate, especially when in ...
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Mandibulata
The clade Mandibulata constitutes one of the major subdivisions of the phylum Arthropoda, alongside Chelicerata. Mandibulates include the crustaceans, myriapods (centipedes and millipedes, among others), and all true insects. The name "Mandibulata" refers to the mandibles, a modified pair of limbs used in food processing, the presence of which are characteristic of most members of the group. The mandibulates are divided between the extant groups Myriapoda (millipedes and centipedes, among others) and Pancrustacea (including crustaceans and hexapods, the latter group containing insects). Molecular phylogenetic studies suggest that the living arthropods are related as shown in the cladogram below. Crustaceans do not form a monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent ...
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Isoxyida
Isoxyids are members of the order Isoxyida and the family Isoxyidae, a group of basal arthropods that existed during the Cambrian period. It contains two genera, ''Isoxys'', with 20 species found worldwide, and ''Surusicaris'' known from a single species found in the Burgess Shale of Canada. They are distinguished by their bivalved carapaces and pair of upward curving grasping frontal appendages. Description Isoxyids have a combination of features seen in both stem-group arthropods, as well as more advanced taxa. Isoxyids have semicircular bivalved Carapace, carapaces, large, spherical Arthropod eye, eyes, a pair of large upward curling frontal appenages covered in spines, and pairs of biramous limbs running along the body. The trunk region appears to be unsegmented and lacks sclerotisation, similar to Radiodonta, radiodonts, with a segmented and sclerotised (arthropodized) trunk being characteristic of most arthropods. Currently only two genera have been accepted to be members ...
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Miaolingian
The Miaolingian is the third Series of the Cambrian Period, and was formally named in 2018. It lasted from about to million years ago and is divided in ascending order into 3 stages: the Wuliuan, Drumian, and Guzhangian. The Miaolingian is preceded by the unnamed Cambrian Series 2 and succeeded by the Furongian series. It is named after the Miaoling Mountains in southeastern Guizhou Province, China. Definition A number of proposals for fossils and type sections were made before it was formally ratified in 2018. The most promising fossil markers were seen to be the respective first appearances of either trilobite species ''Ovatoryctocara granulata'' or ''Oryctocephalus indicus'', which both have an age close to million years ago. After some deliberation, the FAD of ''Oryctocephalus indicus'' was chosen to be the lower boundary marker, and the GSSP was placed in the Kaili Formation, Wuliu-Zengjiayan, Guizhou, China. The Miaolingian-Furongian boundary has the same definition a ...
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