Japanese raccoon dog
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The Japanese raccoon dog (''Nyctereutes viverrinus''), also known as the ''tanuki'' ( ja, , , ), is a species of canid
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
. It is one of two species in the genus ''
Nyctereutes ''Nyctereutes'' (Greek: ''nyx, nykt-'' "night" + ''ereutēs'' "wanderer") is a genus of canid which includes only two extant species both known as raccoon dogs; the common raccoon dog (''Nyctereutes procyonoides'') and the Japanese raccoon dog ( ...
'', alongside the
common raccoon dog The common raccoon dog (''Nyctereutes procyonoides''), also called the Chinese or Asian raccoon dog, is a small, heavy-set, fox-like canid native to East Asia. Named for its raccoon-like face markings, it is most closely related to foxes. Common ...
(''N. procyonoides''), of which it was formerly thought to be a subspecies. The Japanese raccoon dog has a relatively smaller stomach and shorter fur of lesser insulation value than mainland raccoon dogs. A rare, white colour type can also be found. Within
Japanese folklore Japanese folklore encompasses the informally learned folk traditions of Japan and the Japanese people as expressed in its oral traditions, customs, and material culture. In Japanese, the term is used to describe folklore. The academic study o ...
, the ''tanuki'' have had a significant role since ancient times. The legendary ''tanuki'' are reputed to be mischievous and jolly, masters of disguise and
shapeshifting In mythology, folklore and speculative fiction, shape-shifting is the ability to physically transform oneself through an inherently superhuman ability, divine intervention, demonic manipulation, sorcery, spells or having inherited the ...
but somewhat gullible and absentminded. The animals have also been common in Japanese art, particularly as subjects for statues.


Japanese etymology

While ''tanuki'' are prominent in Japanese folklore and proverbs, they were not always clearly distinguished from other animals with a similar appearance. In local dialects, ''tanuki'' and '' mujina'' (, '' kyujitai'': ) can refer to raccoon dogs or the
Japanese badger The Japanese badger (''Meles anakuma'') is a species of carnivoran of the family Mustelidae, the weasels and their kin. Endemic to Japan, it is found on Honshu, Kyushu, Shikoku, and Shōdoshima. It shares the genus '' Meles'' with its close re ...
. An animal known as ''tanuki'' in one region may be known as ''mujina'' in another region. In the modern Tokyo standard dialect, ''tanuki'' refers to raccoon dogs and ''anaguma'' refers to badgers. ''Tanuki'' is often mistakenly translated into English as "
badger Badgers are short-legged omnivores in the family Mustelidae (which also includes the otters, wolverines, martens, minks, polecats, weasels, and ferrets). Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by ...
" or "
raccoon The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight of ...
" (as used in the English translation of the film '' Pom Poko'' and outlined in
Tom Robbins Thomas Eugene Robbins (born July 22, 1932) is a best-selling and prolific American novelist. His most notable works are "seriocomedies" (also known as "comedy drama"), such as ''Even Cowgirls Get the Blues''. Tom Robbins has lived in La Conner ...
' book '' Villa Incognito''), two unrelated types of animals with superficially similar appearances. Traditionally, different areas of Japan had different names for raccoon dogs as animals, which would be used to denote different animals in other parts of the country, including badgers and wild cats; however, the official word in the standard Tokyo dialect is now ''tanuki'', a term that also carries the folkloric significance. The North American raccoon (''Procyon lotor'') is translated as in Japanese, while badger is translated as or as .


Behavior

The Japanese raccoon dog is mainly nocturnal, but they are known to be active during daylight. They vocalize by growling or with groans that have pitches resembling those of domesticated cats. Like cats, the Japanese raccoon dog arches its back when it is trying to intimidate other animals; however, they assume a defensive posture similar to that of other canids, lowering their bodies and showing their bellies to submit. Usually social groups are limited to a breeding pair, but individual Japanese raccoon dogs may stay in a group of non-paired individuals until they find a mate. The species is predominantly monogamous. The breeding period for the species is synchronized between females and males and lasts between February and April. A litter (typically with 4–6 pups) is born after a
gestation period In mammals, pregnancy is the period of reproduction during which a female carries one or more live offspring from implantation in the uterus through gestation. It begins when a fertilized zygote implants in the female's uterus, and ends once ...
of 9 weeks. The parents look after their pups at a den for around a month, and then for another month after the pups leave the den. Japanese raccoon dogs live for 7–8 years in the wild, and have reached the age of 13 in captivity. They have been observed to climb trees to forage for fruits and berries, using their curved claws to climb.


Taxonomy

The Japanese raccoon dog is sometimes classified as its own distinct species due to unique
chromosomal A chromosome is a long DNA molecule with part or all of the genetic material of an organism. In most chromosomes the very long thin DNA fibers are coated with packaging proteins; in eukaryotic cells the most important of these proteins ar ...
,
behavior Behavior (American English) or behaviour (British English) is the range of actions and mannerisms made by individuals, organisms, systems or artificial entities in some environment. These systems can include other systems or organisms as we ...
al, and morphological characteristics absent in mainland raccoon dogs. Researchers have suggested that they be considered a separate species, ''N. viverrinus'', or that raccoon dogs of Japan could be further divisible into separate subspecies as ''N. p. procyonoides'' and ''N. p. albus'', but both views were controversial. However, following morphological and genetic analysis across multiple studies, all of which indicated that ''N. viverrinus'' was a distinct species, it was later classified as such by the
American Society of Mammalogists The American Society of Mammalogists (ASM) was founded in 1919. Its primary purpose is to encourage the study of mammals, and professions studying them. There are over 4,500 members of this society, and they are primarily professional scientists w ...
. Genetic analysis has confirmed unique sequences of
mtDNA Mitochondrial DNA (mtDNA or mDNA) is the DNA located in mitochondria, cellular organelles within eukaryotic cells that convert chemical energy from food into a form that cells can use, such as adenosine triphosphate (ATP). Mitochondrial DNA ...
, classifying the Japanese raccoon dog as a distinct isolation species, based on evidence of eight
Robertsonian translocation Robertsonian translocation (ROB) is a chromosomal abnormality wherein a certain type of a chromosome becomes attached to another. It is the most common form of chromosomal translocation in humans, affecting 1 out of every 1,000 babies born. It doe ...
s. The
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
Canid Group's Canid Biology and Conservation Conference in September 2001 rejected the classification of the Japanese raccoon dog as a separate species, but its status is still disputed, based on its elastic genome. The
karyotype A karyotype is the general appearance of the complete set of metaphase chromosomes in the cells of a species or in an individual organism, mainly including their sizes, numbers, and shapes. Karyotyping is the process by which a karyotype is disce ...
of Japanese raccoon dogs is different from that of the mainland raccoon dogs. Though it is unknown whether mainland raccoon dogs and Japanese raccoon dogs can produce fertile offspring, it is assumed that the chromosomal differences between them would have deleterious effects on the fertility of the potential offspring and this would be indicative of
speciation Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
. Aggregators on mammal taxonomy are inconsistent: Like the IUCN, '' Mammal Species of the World'' (2005) considers the Japanese raccoon dog to be a subspecies, whereas the American Association of Mammologists include ''N. viverrinus'' as a valid species in their Mammal Diversity Database. The raccoon dogs from
Hokkaido is Japan, Japan's Japanese archipelago, second largest island and comprises the largest and northernmost Prefectures of Japan, prefecture, making up its own List of regions of Japan, region. The Tsugaru Strait separates Hokkaidō from Honshu; th ...
are sometimes recognized as a different subspecies from the mainland tanuki as ''Nyctereutes procyonoides albus'' (Hornaday, 1904) (or ''N. viverrinus albus'' if recognized as a distinct species). This taxon is synonymized with ''N. p. viverrinus'' in ''Mammal Species of the World'', but comparative morphometric analysis supports recognizing the Hokkaido population as a distinct subspecific unit.


Conservation

The IUCN places the raccoon dog at "least concern" status due to the animal's wide distribution in Japan and abundant population, including as an introduced species throughout northeastern Europe. In many European countries, it is legal to hunt raccoon dogs, as they are considered a harmful and invasive species. In Japan the species is hunted mainly to prevent them from damaging crops; however, their fur is desired for use in calligraphy brushes and was exported chiefly to the United States before the outbreak of World War II. The animal is a common victim of vehicle accidents, with conservative estimates of up to 370,000 Japanese raccoon dogs being killed by vehicles each year in Japan.


In folklore and tradition

The ''tanuki'' has a long history in Japanese legend and folklore. '' Bake-danuki'' () are a kind of ''tanuki yōkai'' (supernatural beings) found in the classics and in the folklore and legends of various places in Japan. Although the ''tanuki'' is a real, extant animal, the ''bake-danuki'' that appears in literature has always been depicted as a strange, even supernatural animal. The earliest appearance of the ''bake-danuki'' in literature, in the chapter about
Empress Suiko (554 – 15 April 628) was the 33rd monarch of Japan,Imperial Household Agency (''Kunaichō'') 推古天皇 (33)/ref> according to the traditional order of succession. Suiko reigned from 593 until her death in 628. In the history of Japa ...
in the '' Nihon Shoki'' written during the
Nara period The of the history of Japan covers the years from CE 710 to 794. Empress Genmei established the capital of Heijō-kyō (present-day Nara). Except for a five-year period (740–745), when the capital was briefly moved again, it remained the c ...
, there are such passages as "in two months of spring, there are tanuki in the country of Mutsu (), they turn into humans and sing songs ()". Bake-danuki subsequently appear in such classics as the Nihon Ryōiki and the Uji Shūi Monogatari. In some regions of Japan, ''bake-danuki'' are reputed to have abilities similar to those attributed to ''kitsune'' (foxes): they can shapeshift into other things or people, and can possess human beings. Many legends of ''tanuki'' exist in the Sado Islands of
Niigata Prefecture is a prefecture in the Chūbu region of Honshu of Japan. Niigata Prefecture has a population of 2,227,496 (1 July 2019) and is the fifth-largest prefecture of Japan by geographic area at . Niigata Prefecture borders Toyama Prefecture and ...
and in
Shikoku is the smallest of the four main islands of Japan. It is long and between wide. It has a population of 3.8 million (, 3.1%). It is south of Honshu and northeast of Kyushu. Shikoku's ancient names include ''Iyo-no-futana-shima'' (), '' ...
, and among them, like the Danzaburou-danuki of Sado, the Kinchō-tanuki and Rokuemon-tanuki of Awa Province (
Tokushima Prefecture is a prefecture of Japan located on the island of Shikoku. Tokushima Prefecture has a population of 728,633 (1 October 2019) and has a geographic area of 4,146 km2 (1,601 sq mi). Tokushima Prefecture borders Kagawa Prefecture to the nort ...
), and the Yashima no Hage-tanuki of
Kagawa Prefecture is a prefecture of Japan located on the island of Shikoku. Kagawa Prefecture has a population of 949,358 (as of 2020) and is the smallest prefecture by geographic area at . Kagawa Prefecture borders Ehime Prefecture to the southwest and Tok ...
, the ''tanuki'' that possessed special abilities were given names, and even became the subject of rituals. Apart from these places, ''tanuki'' are treated with special regard in a few cases.


In popular culture

Tanuki (or their folklore version) are a recurring theme in Japanese popular culture. The first exposure of non-Japanese to tanuki usually comes through exported Japanese media. However, they are often described as "raccoons" in translation or assumed as such if no species is given. Notable appearances of tanuki in popular culture include: * In the 1992
Super NES The Super Nintendo Entertainment System (SNES), commonly shortened to Super NES or Super Nintendo, is a 16-bit home video game console developed by Nintendo that was released in 1990 in Japan and South Korea, 1991 in North America, 1992 in ...
video game Pocky & Rocky, one of the two playable characters is a tanuki named Rocky who defeats his enemies by shooting leaves at them. Rocky also makes appearances in the 1994, 2001, and 2022 releases in the hit series. * In
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's video games ''
Super Mario Bros. 3 ''Super Mario Bros. 3'' is a platform game developed and published by Nintendo for the Nintendo Entertainment System (NES). It was released for home consoles in Japan on October 23, 1988, in North America on February 12, 1990 and in Europe on ...
'', '' Super Mario 3D Land'', ''
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'', and '' Super Mario 3D World'',
Mario is a character (arts), character created by Japanese video game designer Shigeru Miyamoto. He is the title character of the ''Mario (franchise), Mario'' franchise and the mascot of Japanese video game company Nintendo. Mario has appeared in ...
can wear a "Tanooki Suit". By doing so, he takes on the appearance of a tanuki and gains the ability to fly, spin his tail to attack enemies, and shapeshift into a statue, much like a ''bake-danuki''. The same games also feature the "Super Leaf", which gives Mario tanuki ears and a tail and allows him to fly and use his tail to attack, although this form is known as Raccoon Mario; in ''Super Mario 3D Land'' and ''Super Mario 3D World'', Mario can only transform into his Tanooki form after obtaining a Super Leaf. This power-up is based on the mythology of tanuki using leaves to help themselves transform. * The 1994 Studio Ghibli film '' Pom Poko'' features a group of tanuki who use their shapeshifting powers to defend their habitat against human developers. * In 2015 GitLab.com adopted a new logo of an abstracted raccoon dog and the term Tanuki. * Tom Nook, a recurring character in the '' Animal Crossing'' video game series, is a tanuki, as well as his two employees, Timmy and Tommy. In the English versions of the games he is localized as a raccoon, although his name still alludes to tanuki. The furniture that these characters sell transforms into leaves for easy transport. * A tanuki appears as a newscaster in the Japanese version of the American Disney computer-animated film ''
Zootopia ''Zootopia'' (titled ''Zootropolis'' in various regions) is a 2016 American computer-animated buddy cop action comedy film produced by Walt Disney Animation Studios and released by Walt Disney Pictures. The 55th Disney animated feature fi ...
''; the standard release of the film and releases in other countries use other animals.Loughrey, Clarisse.
Zootropolis' new anchors change animal depending on what country you're in
" ''
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''. March 7, 2016. Retrieved on March 7, 2016.
* In the 2019 ''
Sonic Sonic or Sonics may refer to: Companies *Sonic Drive-In, an American drive-in fast-food restaurant chain * Sonic (ISP), an Internet provider and CLEC, serving more than 100 California communities * Sonic Foundry, a computer software company wh ...
'' game ''
Team Sonic Racing ''Team Sonic Racing'' is a 2019 kart racing game and a spin-off from Sega's ''Sonic the Hedgehog'' series. Controlling one of 15 characters from the series' cast, players compete in races using sports cars. Gameplay is from a third-person pe ...
'', one of the characters is a tanuki named Dodon Pa. * The 2020 Studio Trigger TV anime '' BNA: Brand New Animal'' features main protagonist Michiru Kagemori, a humanoid shape-shifting tanuki who is often mistaken as a raccoon. * '' Genshin Impact'' by miHoYo features the region based on Japan called Inazuma where the player can encounter tanuki who plays with them and offer rewards when solving their puzzles. The tanuki even play a role in the multi-part story quest, Sacred Sakura Cleansing Ritual. In Japanese slang, ''tanuki gao'' ("raccoon dog face") can refer to a face that looks like that of the animal, or a person's facial expression of feigned ignorance. By contrast, ''
kitsune In Japanese folklore, , are foxes that possess paranormal abilities that increase as they get older and wiser. According to '' yōkai'' folklore, all foxes have the ability to shapeshift into human form. While some folktales speak of employing ...
gao'' ("fox face") refers to people with narrow faces, close-set eyes, thin eyebrows, and high cheekbones. A dish called ' ("''tanuki'' soup") ceased to contain actual tanuki meat, but some rural stews do use tanuki.Nicol, C.W.,
Talking tanuki — or whatever you call them
, ''
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'', 4 January 2015, p. 21
Of Japanese noodles, the words "tanuki" and "kitsune" designate two varieties of the
udon Udon ( or ) is a thick noodle made from wheat flour, used in Japanese cuisine. It is a comfort food for many Japanese people. There are a variety of ways it is prepared and served. Its simplest form is in a hot soup as with a mild broth called ...
or
soba Soba ( or , "buckwheat") is a thin Japanese noodle made from buckwheat. The noodles are served either chilled with a dipping sauce, or hot in a noodle soup. The variety ''Nagano soba'' includes wheat flour. In Japan, soba noodles can be found ...
dishes. Neither contain any of those meats. Tanuki udon/soba contains flakes of fried tempura batter ("
tenkasu are crunchy bits of deep fried flour- batter used in Japanese cuisine, specifically in dishes such as '' soba'', ''udon'', ''takoyaki'' and ''okonomiyaki''. Hot plain ''soba'' and ''udon'' with added ''tenkasu'' are called ''tanuki-soba'' and ...
"), while kitsune udon/soba contains fried tofu (" abura-age").


References


Further reading

* * * *


External links


"Report: ''Nyctereutes procyonoides viverrinus''"
in the
Integrated Taxonomic Information System The Integrated Taxonomic Information System (ITIS) is an American partnership of federal agencies designed to provide consistent and reliable information on the taxonomy of biological species. ITIS was originally formed in 1996 as an interagen ...
{{Taxonbar, from=Q6111761 Nyctereutes Mammals of Japan Endemic fauna of Japan Mammals described in 1838 Taxa named by Coenraad Jacob Temminck Shapeshifting Japanese folklore Articles containing video clips Taxobox binomials not recognized by IUCN