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Canidae
Canidae (; from Latin, '' canis'', " dog") is a biological family of dog-like carnivorans, colloquially referred to as dogs, and constitutes a clade. A member of this family is also called a canid (). There are three subfamilies found within the canid family, which are the extinct Borophaginae and Hesperocyoninae, and the extant Caninae. The Caninae are known as canines, and include domestic dogs, wolves, coyotes, foxes, jackals and other extant and extinct species. Canids are found on all continents except Antarctica, having arrived independently or accompanied human beings over extended periods of time. Canids vary in size from the gray wolf to the fennec fox. The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails. They are mostly social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a ...
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Borophaginae
The extinct Borophaginae form one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Hesperocyoninae and extant Caninae. Borophaginae, called "bone-crushing dogs", were endemic to North America during the Oligocene to Pliocene and lived roughly 34—2.5 million years ago, existing for about . Origin The Borophaginae descended from the subfamily Hesperocyoninae. The earliest and most primitive borophagine is the genus ''Archaeocyon'', which is a small fox-sized animal mostly found in the fossil beds in western North America. The borophagines soon diversified into several major groups. They evolved to become considerably larger than their predecessors, and filled a wide range of niches in late Cenozoic North America, from small omnivores to powerful, bear-sized carnivores, such as '' Epicyon''. Species There are 66 identified borophagine species, including 18 new ones that range from the Orellan to Blancan ages. A phylogen ...
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Cuon
The dhole (''Cuon alpinus''; ) is a canid native to Central, South, East and Southeast Asia. Other English names for the species include Asian wild dog, Asiatic wild dog, Indian wild dog, whistling dog, red dog, red wolf, and mountain wolf. It is genetically close to species within the genus ''Canis'', but distinct in several anatomical aspects: its skull is convex rather than concave in profile, it lacks a third lower molar and the upper molars sport only a single cusp as opposed to between two and four. During the Pleistocene, the dhole ranged throughout Asia, Europe and North America but became restricted to its historical range 12,000–18,000 years ago. The dhole is a highly social animal, living in large clans without rigid dominance hierarchies and containing multiple breeding females. Such clans usually consist of about 12 individuals, but groups of over 40 are known. It is a diurnal pack hunter which preferentially targets large and medium-sized ungulates ...
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Caninae
The Caninae, known as canines, are one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Borophaginae and Hesperocyoninae. The Caninae includes all living canids and their most recent fossil relatives. Their fossils were first found in North America and dated to the Oligocene era, then spreading to Asia at the end of the Miocene era, some 7 million to 8 million years ago. Taxonomy and lineage The genus ''Leptocyon'' (Greek: ''leptos'' slender + ''cyon'' dog) includes 11 species and was the first primitive canine. They were small and weighed around 2 kg. They first appeared in Sioux County, Nebraska in the Orellan era 34-32 million years ago, which was the beginning of the Oligocene. This was the same time as the appearance of the Borophaginae with whom they share features, indicating that these were two sister groups. Borophaginae skull and dentition were designed for a powerful killing bite compared ...
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Hesperocyoninae
The extinct Hesperocyoninae are one of three subfamilies found within the canid family. The other two canid subfamilies are the extinct Borophaginae and extant Caninae. Taxonomic history Hesperocyoninae are basal canids that gave rise to the other two canid subfamilies, the Borophaginae and Caninae. This subfamily was endemic to North America, living from the Duchesnean stage of the Late Eocene through to the early Barstovian stage of the Miocene, lasting around 20 million years. It comprises 10 recognized genera and 26 recognized species. Four major lineages can be defined based on shared characteristics: *''Mesocyon''-''Enhydrocyon'' clade (includes '' Cynodesmus, Sunkahetanka, Philotrox'') *''Osbornodon'' clade *''Paraenhydrocyon'' *''Ectopocynus clade'' The genus '' Caedocyon'', which is only known from a single partial cranium that shows some ''Paraenhydrocyon'' affinities, probably represents another independent lineage. '' Hesperocyon'', which lacks the shared de ...
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Canis
''Canis'' is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and dentition, long legs, and comparatively short ears and tails.Heptner, V. G.; Naumov, N. P. (1998). ''Mammals of the Soviet Union'' Vol.II Part 1a, SIRENIA AND CARNIVORA (Sea Cows, Wolves and Bears). Science Publishers, Inc. USA. pp. 124–129. . Taxonomy The genus ''Canis'' ( Carl Linnaeus, 1758) was published in the 10th edition of Systema Naturae and included the dog-like carnivores: the domestic dog, wolves, coyotes and jackals. All species within ''Canis'' are phylogenetically closely related with 78 chromosomes and can potentially interbreed. In 1926, the International Commission on Zoological Nomenclature (ICZN) in Opinion 91 included Genus ''Canis'' on its ''Official Lists and Indexes of Names in Zoology''. In 1955, the ICZN's Dire ...
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Canis
''Canis'' is a genus of the Caninae which includes multiple extant species, such as wolves, dogs, coyotes, and golden jackals. Species of this genus are distinguished by their moderate to large size, their massive, well-developed skulls and dentition, long legs, and comparatively short ears and tails.Heptner, V. G.; Naumov, N. P. (1998). ''Mammals of the Soviet Union'' Vol.II Part 1a, SIRENIA AND CARNIVORA (Sea Cows, Wolves and Bears). Science Publishers, Inc. USA. pp. 124–129. . Taxonomy The genus ''Canis'' ( Carl Linnaeus, 1758) was published in the 10th edition of Systema Naturae and included the dog-like carnivores: the domestic dog, wolves, coyotes and jackals. All species within ''Canis'' are phylogenetically closely related with 78 chromosomes and can potentially interbreed. In 1926, the International Commission on Zoological Nomenclature (ICZN) in Opinion 91 included Genus ''Canis'' on its ''Official Lists and Indexes of Names in Zoology''. In 1955, the ICZN's Dire ...
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Otocyon
The bat-eared fox (''Otocyon megalotis'') is a species of fox found on the African savanna. It is the only extant species of the genus ''Otocyon'' and considered a basal canid species. Fossil records indicate this canid first appeared during the middle Pleistocene. It is named for its large ears, which have a role in thermoregulation. The bat referred to in its colloquial name is possibly the Egyptian slit-faced bat (''Nycteris thebaica''), which is abundant in the region and has very large ears. Although not commonly used, other vernacular names include big-eared fox, black-eared fox, long-eared fox, Delalande's fox, cape fox, and motlosi. Systematics and evolution The bat-eared fox is the only living species of the genus ''Otocyon''. Its scientific name, given by Anselme Gaëtan Desmarest, was initially ''Canis megalotis'' (due to its close resemblance to jackals), and later changed by Salomon Müller which placed it in its own genus, ''Otocyon''; its huge ears and differen ...
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Wolf
The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly understood, comprise wild subspecies. The wolf is the largest extant member of the family Canidae. It is also distinguished from other '' Canis'' species by its less pointed ears and muzzle, as well as a shorter torso and a longer tail. The wolf is nonetheless related closely enough to smaller ''Canis'' species, such as the coyote and the golden jackal, to produce fertile hybrids with them. The banded fur of a wolf is usually mottled white, brown, gray, and black, although subspecies in the arctic region may be nearly all white. Of all members of the genus ''Canis'', the wolf is most specialized for cooperative game hunting as demonstrated by its physical adaptations to tackling large prey, its more social nature, and its highly adv ...
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Coyote
The coyote (''Canis latrans'') is a species of canine native to North America. It is smaller than its close relative, the wolf, and slightly smaller than the closely related eastern wolf and red wolf. It fills much of the same ecological niche as the golden jackal does in Eurasia. The coyote is larger and more predatory and was once referred to as the American jackal by a behavioral ecologist. Other historical names for the species include the prairie wolf and the brush wolf. The coyote is listed as least concern by the International Union for Conservation of Nature, due to its wide distribution and abundance throughout North America. The species is versatile, able to adapt to and expand into environments modified by humans. It is enlarging its range by moving into urban areas in the eastern U.S. and Canada. The coyote was sighted in eastern Panama (across the Panama Canal from their home range) for the first time in 2013. The coyote has 19 recognized subspecies. The average ...
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Jackals
Jackals are medium-sized canids native to Africa and Eurasia. While the word "jackal" has historically been used for many canines of the subtribe canina, in modern use it most commonly refers to three species: the closely related black-backed jackal (''Lupulella mesomelas'') and side-striped jackal (''Lupulella adusta'') of sub-Saharan-Africa, and the golden jackal (''Canis aureus'') of south-central Europe and Asia. The African golden wolf (''Canis lupaster'') was also formerly considered as a jackal. While they do not form a monophyletic clade, all jackals are opportunistic omnivores, predators of small to medium-sized animals and proficient scavengers. Their long legs and curved canine teeth are adapted for hunting small mammals, birds, and reptiles, and their large feet and fused leg bones give them a physique well-suited for long-distance running, capable of maintaining speeds of for extended periods of time. Jackals are crepuscular, most active at dawn and dusk. ...
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Vulpes
'' Vulpes '' is a genus of the sub-family Caninae. The members of this genus are colloquially referred to as true foxes, meaning they form a proper clade. The word " fox" occurs in the common names of all species of the genus, but also appears in the common names of other canid species. True foxes are distinguished from members of the genus '' Canis'', such as domesticated dogs, wolves, jackals and coyotes, by their smaller size (5–11 kg), longer, bushier tail, and flatter skull. They have black, triangular markings between their eyes and nose, and the tip of their tail is often a different color from the rest of their pelt. The typical lifespan for this genus is between two and four years, but can reach up to a decade. Extant species Within ''Vulpes'', 12 separate extant species and four fossil species are described: Early history The oldest known fossil species within ''Vulpes'' is ''V. riffautae'', dating back to the late Miocene of Chad, which is within the ...
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Dusicyon
''Dusicyon'' is an extinct genus of South American canids. The type species is ''Dusicyon australis'', the Falkland Islands wolf. In 1914, Oldfield Thomas established this genus, in which he included the culpeo and other South American foxes. These other canids were removed to '' Lycalopex'' by Langguth in 1975. '' Dusicyon avus'', widely distributed in the late Pleistocene from Uruguay through Buenos Aires Province to southernmost Chile, is the closest known relative of the Falkland Islands wolf; the two lineages split only about 16,000 years ago. It died out in the late Holocene, earlier estimates suggested about 2,980 years ago on the island of Tierra del Fuego and almost 1,700 years ago in the continent. More recent research confirms much later extinction dates, with the latest confirmed records in the Pampean Region being 700 BP (1232–1397 AD) and southernmost Patagonia at 400 years BP (1454–1626 AD). There is still much debate about the classification of ''"Dusicyo ...
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