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Alienoptera
Alienopteridae is an extinct family of dictyopterans, known from the Mid-Cretaceous to Eocene. They are noted for their unusual combination of features not found in other dictyopterans. Taxonomy It was originally assigned to its own order Alienoptera by Bai et al., 2016. It was reassigned to the dictyopteran superfamily Umenocoleoidea as sister family to the beetle-like Umenocoleidae by Vršanský ''et al.'' (2018), and a more recent analysis similarly places Alienopteridae and Umenocoleidae as sister taxa within Dictyoptera, but placing both lineages outside of Blattodea. A 2021 study revived the order Alienoptera for the clade containing Alienopteridae and Umenocoleidae, with a cladistic analysis placing Alienoptera as the sister clade to Mantodea (praying mantises). Distribution The majority of the alienopterid genera are known from the mid Cretaceous (latest Albian-earliest Cenomanian ~ 100 million years ago) Burmese amber found in Myanmar; though an additional two gener ...
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Alienopterus
''Alienopterus brachyelytrus'' is an extinct insect described from a 99 million year old fossil found in Burmese amber from the Hukawng Valley of Myanmar. It was the first known member of the order Alienoptera until 2018, when the second and third members of the order, '' Caputoraptor elegans'', and '' Alienopterella stigmatica'' were described. ''A. brachyelytrus'' has characters that are shared with cockroaches and mantids and is thought to represent either the sister taxon, or an ancestor to mantids Mantidae is one of the largest families in the order of praying mantises, based on the type species '' Mantis religiosa''; however, most genera are tropical or subtropical. Historically, this was the only family in the order, and many referen .... ''Alienopterus'' has shortened forewings and functional hindwings capable of flight that are attached to pads as in the Mantophasmatodea. The foreleg has a femoral brush which is a characteristic of Mantodea. The mouth points down ...
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Umenocoleoidea
Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). While all modern Dictyoptera have short ovipositors, the oldest fossils of Dictyoptera have long ovipositors, much like members of the Orthoptera. Classification and phylogeny The use of the term Dictyoptera has changed over the years, and while largely out of use for much of the last century, it is becoming more widely used. It has usually been considered a superorder, with Isoptera, Blattodea and Mantodea being its three orders. In some classifications, however, Dictyoptera is shifted to order status and in others the order Isoptera has been subsumed under Blattodea while retaining Dictyoptera as a superorder. Regardless, in all classifications the constituent groups are the same, just treated at different rank. ...
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Dictyoptera
Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). While all modern Dictyoptera have short ovipositors, the oldest fossils of Dictyoptera have long ovipositors, much like members of the Orthoptera. Classification and phylogeny The use of the term Dictyoptera has changed over the years, and while largely out of use for much of the last century, it is becoming more widely used. It has usually been considered a superorder, with Isoptera, Blattodea and Mantodea being its three orders. In some classifications, however, Dictyoptera is shifted to order status and in others the order Isoptera has been subsumed under Blattodea while retaining Dictyoptera as a superorder. Regardless, in all classifications the constituent groups are the same, just treated at different ran ...
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Dictyoptera
Dictyoptera (from Greek δίκτυον ''diktyon'' "net" and πτερόν ''pteron'' "wing") is an insect superorder that includes two extant orders of polyneopterous insects: the order Blattodea (termites and cockroaches together) and the order Mantodea (mantises). While all modern Dictyoptera have short ovipositors, the oldest fossils of Dictyoptera have long ovipositors, much like members of the Orthoptera. Classification and phylogeny The use of the term Dictyoptera has changed over the years, and while largely out of use for much of the last century, it is becoming more widely used. It has usually been considered a superorder, with Isoptera, Blattodea and Mantodea being its three orders. In some classifications, however, Dictyoptera is shifted to order status and in others the order Isoptera has been subsumed under Blattodea while retaining Dictyoptera as a superorder. Regardless, in all classifications the constituent groups are the same, just treated at different ran ...
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Blattodea
Blattodea is an order of insects that contains cockroaches and termites. Formerly, termites were considered a separate order, Isoptera, but genetic and molecular evidence suggests they evolved from within the cockroach lineage, cladistically making them cockroaches as well. The Blattodea and the mantis (order Mantodea) are now all considered part of the superorder Dictyoptera. Blattodea includes approximately 4,400 species of cockroach in almost 500 genera, and about 3,000 species of termite in around 300 genera. Termites are pale-coloured, soft-bodied eusocial insects that live in colonies, whereas cockroaches are darker-coloured (often brown), sclerotized, segmented insects. Within the colony, termites have a caste system, with a pair of mature reproductives, the king and the queen, and numerous sterile workers and soldiers. Cockroaches are not colonial but do have a tendency to aggregate and may be considered pre-social, as all adults are capable of breeding. Other simil ...
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Caputoraptor
''Caputoraptor'' is an extinct genus of insect from the Cenomanian aged Burmese amber containing two species: the type species ''Caputoraptor elegans'', as well as ''Caputoraptor vidit''. It is part of the extinct order Alienoptera. ''C. elegans'' is notable for the presence of a scissor like mechanism consisting of a straight edge on the back of the head and corresponding serrated edges on the first thoracic segment, these were initially suggested to used by the female to grasp the male during mating, but the structure is not Sexual dimorphism, sexually dimorphic, so a use to hold prey was subsequently suggested. Its morphology suggests a predatory habit inhabiting shrubs and trees. In 2020 a ''Caputoraptor elegans'' nymph was described that was in the process of being predated upon by a ''Ceratomyrmex'' Haidomyrmecinae, hell ant. References

Prehistoric insect genera Burmese amber Fossil taxa described in 2018 {{cretaceous-insect-stub ...
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Green River Formation
The Green River Formation is an Eocene geologic formation that records the sedimentation in a group of intermountain lakes in three basins along the present-day Green River in Colorado, Wyoming, and Utah. The sediments are deposited in very fine layers, a dark layer during the growing season and a light-hue inorganic layer in the dry season. Each pair of layers is called a varve and represents one year. The sediments of the Green River Formation present a continuous record of six million years. The mean thickness of a varve here is 0.18 mm, with a minimum thickness of 0.014 mm and maximum of 9.8 mm.Bradley, W. H. The varves and climate of the Green River epoch: U.S. Geol. Survey Prof. Paper 158, pp 87–110, 1929. The sedimentary layers were formed in a large area named for the Green River, a tributary of the Colorado River. The three separate basins lie around the Uinta Mountains (north, east, and south) of northeastern Utah: * an area in northwestern Colorado eas ...
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Brazil
Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area and the List of countries and dependencies by population, seventh most populous. Its capital is Brasília, and List of cities in Brazil by population, its most populous city is São Paulo. The federation is composed of the union of the 26 States of Brazil, states and the Federal District (Brazil), Federal District. It is the largest country to have Portuguese language, Portuguese as an List of territorial entities where Portuguese is an official language, official language and the only one in the Americas; one of the most Multiculturalism, multicultural and ethnically diverse nations, due to over a century of mass Immigration to Brazil, immigration from around the world; and the most populous Catholic Church by country, Roman Catholic-major ...
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Colorado
Colorado (, other variants) is a state in the Mountain states, Mountain West subregion of the Western United States. It encompasses most of the Southern Rocky Mountains, as well as the northeastern portion of the Colorado Plateau and the western edge of the Great Plains. Colorado is the List of U.S. states and territories by area, eighth most extensive and List of U.S. states and territories by population, 21st most populous U.S. state. The 2020 United States Census, 2020 United States census enumerated the population of Colorado at 5,773,714, an increase of 14.80% since the 2010 United States Census, 2010 United States census. The region has been inhabited by Indigenous peoples of the Americas, Native Americans and their Paleo-Indians, ancestors for at least 13,500 years and possibly much longer. The eastern edge of the Rocky Mountains was a major migration route for early peoples who spread throughout the Americas. "''Colorado''" is the Spanish adjective meaning "ruddy", th ...
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Myanmar
Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explains, the English spellings of both Myanmar and Burma assume a non-rhotic variety of English, in which the letter r before a consonant or finally serves merely to indicate a long vowel: �mjænmɑː, ˈbɜːmə So the pronunciation of the last syllable of Myanmar as ɑːror of Burma as ɜːrməby some speakers in the UK and most speakers in North America is in fact a spelling pronunciation based on a misunderstanding of non-rhotic spelling conventions. The final ''r'' in ''Myanmar'' was not intended for pronunciation and is there to ensure that the final a is pronounced with the broad ''ah'' () in "father". If the Burmese name my, မြန်မာ, label=none were spelled "Myanma" in English, this would be pronounced at the end by all ...
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Burmese Amber
Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. The amber is of significant palaeontological interest due to the diversity of flora and fauna contained as inclusions, particularly arthropods including insects and arachnids but also birds, lizards, snakes, frogs and fragmentary dinosaur remains. The amber has been known and commercially exploited since the first century AD, and has been known to science since the mid-nineteenth century. Research on the deposit has attracted controversy due to its alleged role in funding internal conflict in Myanmar and hazardous working conditions in the mines where it is collected. Geological context, depositional environment and age The amber is found within the Hukawng Basin, a large Cretaceous-Cenozoic sedimentary basin within northern Myanmar. The s ...
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Cenomanian
The Cenomanian is, in the ICS' geological timescale, the oldest or earliest age of the Late Cretaceous Epoch or the lowest stage of the Upper Cretaceous Series. An age is a unit of geochronology; it is a unit of time; the stage is a unit in the stratigraphic column deposited during the corresponding age. Both age and stage bear the same name. As a unit of geologic time measure, the Cenomanian Age spans the time between 100.5 and 93.9 million years ago (Mya). In the geologic timescale, it is preceded by the Albian and is followed by the Turonian. The Upper Cenomanian starts around at 95 Mya. The Cenomanian is coeval with the Woodbinian of the regional timescale of the Gulf of Mexico and the early part of the Eaglefordian of the regional timescale of the East Coast of the United States. At the end of the Cenomanian, an anoxic event took place, called the Cenomanian-Turonian boundary event or the "Bonarelli event", that is associated with a minor extinction event for m ...
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