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Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
of
monocotyledon Monocotyledons (), commonly referred to as monocots, ( Lilianae '' sensu'' Chase & Reveal) are flowering plants whose seeds contain only one embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but with various ranks ...
ous
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed with ...
s commonly known as grasses. It includes the
cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
grasses,
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
s, the grasses of natural
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
and species cultivated in
lawn A lawn () is an area of soil-covered land planted with Poaceae, grasses and other durable plants such as clover lawn, clover which are maintained at a short height with a lawn mower (or sometimes grazing animals) and used for aesthetic an ...
s and
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Types of pasture Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, c ...
. The latter are commonly referred to collectively as grass. With around 780
genera Genus (; : genera ) is a taxonomic rank above species and below family as used in the biological classification of living and fossil organisms as well as viruses. In binomial nomenclature, the genus name forms the first part of the binomial s ...
and around 12,000 species, the Poaceae is the fifth-largest plant family, following the
Asteraceae Asteraceae () is a large family (biology), family of flowering plants that consists of over 32,000 known species in over 1,900 genera within the Order (biology), order Asterales. The number of species in Asteraceae is rivaled only by the Orchi ...
,
Orchidaceae Orchids are plants that belong to the family (biology), family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant. Orchids are cosmopolitan distribution, cosmopolitan plants that ...
,
Fabaceae Fabaceae () or Leguminosae,International Code of Nomen ...
and
Rubiaceae Rubiaceae () is a family (biology), family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with Petiole ( ...
. The Poaceae are the most economically important plant family, including
staple food A staple food, food staple, or simply staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard diet for an individual or a population group, supplying a large fraction of energy needs an ...
s from domesticated
cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
crops such as
maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
,
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
,
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
,
oats The oat (''Avena sativa''), sometimes called the common oat, is a species of cereal grain grown for its seed, which is known by the same name (usually in the plural). Oats appear to have been domesticated as a secondary crop, as their seed ...
,
barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
, and
millet Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most millets belong to the tribe Paniceae. Millets are important crops in the Semi-arid climate, ...
for people and as feed for meat-producing animals. They provide, through direct human consumption, just over one-half (51%) of all dietary energy; rice provides 20%, wheat supplies 20%, maize (corn) 5.5%, and other
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
s 6%. Some members of the Poaceae are used as building materials (
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
,
thatch Thatching is the craft of building a roof with dry vegetation such as straw, Phragmites, water reed, Cyperaceae, sedge (''Cladium mariscus''), Juncus, rushes, Calluna, heather, or palm branches, layering the vegetation so as to shed water away fr ...
, and
straw Straw is an agricultural byproduct consisting of the dry wikt:stalk, stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the crop yield, yield by weight of cereal crops such as barley, oats, rice, ry ...
); others can provide a source of
biofuel Biofuel is a fuel that is produced over a short time span from Biomass (energy), biomass, rather than by the very slow natural processes involved in the formation of fossil fuels such as oil. Biofuel can be produced from plants or from agricu ...
, primarily via the conversion of maize to
ethanol Ethanol (also called ethyl alcohol, grain alcohol, drinking alcohol, or simply alcohol) is an organic compound with the chemical formula . It is an Alcohol (chemistry), alcohol, with its formula also written as , or EtOH, where Et is the ps ...
. Grasses have stems that are hollow except at the nodes and narrow alternate leaves borne in two ranks. The lower part of each leaf encloses the stem, forming a leaf-sheath. The leaf grows from the base of the blade, an adaptation allowing it to cope with frequent grazing.
Grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s such as
savannah A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
and
prairie Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
where grasses are dominant are estimated to constitute 40.5% of the land area of the
Earth Earth is the third planet from the Sun and the only astronomical object known to Planetary habitability, harbor life. This is enabled by Earth being an ocean world, the only one in the Solar System sustaining liquid surface water. Almost all ...
, excluding
Greenland Greenland is an autonomous territory in the Danish Realm, Kingdom of Denmark. It is by far the largest geographically of three constituent parts of the kingdom; the other two are metropolitan Denmark and the Faroe Islands. Citizens of Greenlan ...
and
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
. Grasses are also an important part of the vegetation in many other habitats, including
wetland A wetland is a distinct semi-aquatic ecosystem whose groundcovers are flooded or saturated in water, either permanently, for years or decades, or only seasonally. Flooding results in oxygen-poor ( anoxic) processes taking place, especially ...
s,
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
s and
tundra In physical geography, a tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. There are three regions and associated types of tundra: #Arctic, Arctic, Alpine tundra, Alpine, and #Antarctic ...
. Though they are commonly called "grasses", groups such as the seagrasses, rushes and sedges fall outside this family. The rushes and sedges are related to the Poaceae, being members of the order
Poales The Poales are a large order (biology), order of flowering plants in the monocotyledons, and includes families of plants such as the Poaceae, grasses, bromeliads, Juncaceae, rushes and Cyperaceae, sedges. 14 plant families are currently recogniz ...
, but the seagrasses are members of the order Alismatales. However, all of them belong to the
monocot Monocotyledons (), commonly referred to as monocots, (Lilianae ''sensu'' Chase & Reveal) are flowering plants whose seeds contain only one Embryo#Plant embryos, embryonic leaf, or cotyledon. A monocot taxon has been in use for several decades, but ...
group of plants.


Description

Grasses may be annual or
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
herbs, generally with the following characteristics (the image gallery can be used for reference): The stems of grasses, called culms, are usually cylindrical (more rarely flattened, but not 3-angled) and are hollow, plugged at the nodes, where the leaves are attached. Grass
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are nearly always alternate and distichous (in one plane), and have parallel veins. Each leaf is differentiated into a lower sheath hugging the stem and a blade with entire (i.e., smooth) margins. The leaf blades of many grasses are hardened with
silica Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , commonly found in nature as quartz. In many parts of the world, silica is the major constituent of sand. Silica is one of the most complex and abundant f ...
phytolith Phytoliths (from Greek language, Greek, "plant stone") are rigid, microscopic mineral deposits found in some plant tissues, often persisting after the decay of the plant. Although some use "phytolith" to refer to all mineral secretions by plants, ...
s, which discourage grazing animals; some, such as sword grass, are sharp enough to cut human skin. A membranous appendage or fringe of hairs called the
ligule A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above g ...
lies at the junction between sheath and blade, preventing water or insects from penetrating into the sheath.
Flower Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
s of Poaceae are characteristically arranged in
spikelet A spikelet, in botany, describes the typical arrangement of the inflorescences of grasses, sedges and some other monocots. Each spikelet has one or more florets. The spikelets are further grouped into panicles or spikes. The part of the sp ...
s, each having one or more florets. The spikelets are further grouped into panicles or spikes. The part of the spikelet that bears the florets is called the rachilla. A spikelet consists of two (or sometimes fewer)
bracts In botany, a bract is a modified or specialized leaf, associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves in size, color, shape or texture. They also loo ...
at the base, called glumes, followed by one or more florets. A floret consists of the flower surrounded by two bracts, one external—the lemma—and one internal—the palea. The flowers are usually hermaphroditic
maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
being an important exception—and mainly anemophilous or wind-pollinated, although insects occasionally play a role. The
perianth The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower. It is a structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla (petals) or tepal ...
is reduced to two scales, called '' lodicules'', that expand and contract to spread the lemma and palea; these are generally interpreted to be modified sepals. The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
of grasses is a caryopsis, in which the seed coat is fused to the fruit wall. A
tiller A tiller or till is a lever used to steer a vehicle. The mechanism is primarily used in watercraft, where it is attached to an outboard motor, rudder post, rudder post or stock to provide leverage in the form of torque for the helmsman to turn ...
is a leafy shoot other than the first shoot produced from the seed.


Growth and development

Grass blades grow at the base of the blade and not from elongated stem tips. This low growth point evolved in response to grazing animals and allows grasses to be grazed or mown regularly without severe damage to the plant. Three general classifications of growth habit present in grasses: bunch-type (also called caespitose),
stolon In biology, a stolon ( from Latin ''wikt:stolo, stolō'', genitive ''stolōnis'' – "branch"), also known as a runner, is a horizontal connection between parts of an organism. It may be part of the organism, or of its skeleton. Typically, animal ...
iferous, and rhizomatous. The success of the grasses lies in part in their morphology and growth processes and in part in their physiological diversity. There are both C3 and C4 grasses, referring to the photosynthetic pathway for carbon fixation. The C4 grasses have a photosynthetic pathway, linked to specialized Kranz leaf anatomy, which allows for increased water use efficiency, rendering them better adapted to hot, arid environments. The C3 grasses are referred to as "cool-season" grasses, while the C4 plants are considered "warm-season" grasses. * Annual cool-season –
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
, rye, annual bluegrass (annual meadowgrass, '' Poa annua''), and oat * Perennial cool-season – orchardgrass (cocksfoot, ''
Dactylis glomerata ''Dactylis glomerata'' is a species of flowering plant in the grass family Poaceae, known as cock's-foot,Interactive Flora of NW Europ''Dactylis glomerata'' (Cock's-foot)/ref> also colloquially as orchard grass, or cat grass (due to its populari ...
''), fescue (''
Festuca ''Festuca'' (fescue) is a genus of flowering plants belonging to the grass family Poaceae (subfamily Pooideae). They are evergreen or herbaceous perennial tufted grasses with a height range of and a cosmopolitan distribution, occurring on ...
'' spp.), Kentucky bluegrass and perennial ryegrass ('' Lolium perenne'') * Annual warm-season –
maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
, sudangrass, and
pearl millet Pearl millet (''Cenchrus americanus'', commonly known as the synonym ''Pennisetum glaucum'') is the most widely grown type of millet. It has been grown in Africa and the Indian subcontinent since prehistoric times. The center of diversity, and ...
* Perennial warm-season –
big bluestem ''Andropogon gerardi'', commonly known as big bluestem, is a species of tall grass native to much of the Great Plains and grassland regions of central and eastern North America. It is also known as tall bluestem, bluejoint, and turkeyfoot. Taxon ...
, Indiangrass, Bermudagrass and
switchgrass ''Panicum virgatum'', commonly known as switchgrass, is a perennial warm season bunchgrass native to North America, where it occurs naturally from 55th parallel north, 55°N latitude in Canada southwards into the United States and Mexico. Switch ...
. Although the C4 species are all in the PACMAD clade (see diagram below), it seems that various forms of C4 have arisen some twenty or more times, in various subfamilies or genera. In the ''
Aristida ''Aristida'' is a very nearly Cosmopolitan distribution, cosmopolitan genus of plants in the Poaceae, grass family. ''Aristida'' is distinguished by having three Awn (Botany), awns (bristles) on each lemma (botany), lemma of each floret. The gen ...
'' genus for example, one species (''A. longifolia'') is C3 but the approximately 300 other species are C4. As another example, the whole tribe of
Andropogoneae The Andropogoneae, sometimes called the sorghum tribe, are a large tribe of grasses (family Poaceae) with roughly 1,200 species in 90 genera, mainly distributed in tropical and subtropical areas. They include such important crops as maize (corn), ...
, which includes
maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
,
sorghum ''Sorghum bicolor'', commonly called sorghum () and also known as great millet, broomcorn, guinea corn, durra, imphee, jowar, or milo, is a species in the Poaceae, grass genus ''Sorghum (genus), Sorghum'' cultivated for its grain. The grain i ...
,
sugar cane Sugarcane or sugar cane is a species of tall, Perennial plant, perennial grass (in the genus ''Saccharum'', tribe Andropogoneae) that is used for sugar Sugar industry, production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fib ...
, " Job's tears", and bluestem grasses, is C4. Around 46 percent of grass species are C4 plants.


Taxonomy

The name Poaceae was given by John Hendley Barnhart in 1895, based on the tribe Poeae described in 1814 by
Robert Brown Robert Brown may refer to: Robert Brown (born 1965), British Director, Animator and author Entertainers and artists * Washboard Sam or Robert Brown (1910–1966), American musician and singer * Robert W. Brown (1917–2009), American printmaker ...
, and the type genus ''
Poa ''Poa'' is a genus of about 570 species of Poaceae, grasses, native to the temperate regions of both hemispheres. Common names include meadow-grass (mainly in Europe and Asia), bluegrass (mainly in North America), tussock (some New Zealand spe ...
'' described in 1753 by
Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...
. The term is derived from the Ancient Greek πόα (póa, "fodder").


Evolutionary history

Grasses include some of the most versatile
plant life-form Plant life-form schemes constitute a way of classifying plants alternatively to the ordinary species-genus-family scientific classification. In colloquial speech, plants may be classified as trees, shrubs, herbs (forbs and graminoids), etc. The sci ...
s. They became widespread toward the end of the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
period, and fossilized
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
dung (
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name ...
s) belonging to the
sauropod Sauropoda (), whose members are known as sauropods (; from '' sauro-'' + '' -pod'', 'lizard-footed'), is a clade of saurischian ('lizard-hipped') dinosaurs. Sauropods had very long necks, long tails, small heads (relative to the rest of their b ...
titanosaurs (though this provenance has been questioned) have been found containing
phytolith Phytoliths (from Greek language, Greek, "plant stone") are rigid, microscopic mineral deposits found in some plant tissues, often persisting after the decay of the plant. Although some use "phytolith" to refer to all mineral secretions by plants, ...
s of a variety that include grasses that are related to modern
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
and
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
. Grasses have adapted to conditions in lush
rain forest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
s, dry
desert A desert is a landscape where little precipitation occurs and, consequently, living conditions create unique biomes and ecosystems. The lack of vegetation exposes the unprotected surface of the ground to denudation. About one-third of the la ...
s, cold mountains and even intertidal habitats, and are currently the most widespread plant type; grass is a valuable source of food and energy for all sorts of wildlife. A cladogram shows subfamilies and approximate species numbers in brackets: Before 2005, fossil findings indicated that grasses evolved around 55 million years ago. Finds of grass-like
phytolith Phytoliths (from Greek language, Greek, "plant stone") are rigid, microscopic mineral deposits found in some plant tissues, often persisting after the decay of the plant. Although some use "phytolith" to refer to all mineral secretions by plants, ...
s in
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
dinosaur
coprolite A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name ...
s from the latest Cretaceous (
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
) aged Lameta Formation of India have pushed this date back to 66 million years ago. Due to high phosphatic content of 12.2-16.2% in Type A coprolites collected from the Lameta, an omnivore is hypothesized to be the source, contradicting the hypothesis of a sauropod origin. In 2011, fossils from the same deposit were found to belong to the modern rice tribe
Oryzeae Oryzeae is a Tribe (biology), tribe of flowering plants in the true grass family, Poaceae. It contains 11 genera, including both cultivated rice (''Oryza'') and wild rice (''Zizania''). Genera There are 11 genera classified in two subtribes: ...
, suggesting substantial diversification of major lineages by this time. In 2018, a study described grass microfossils extracted from the teeth of the hadrosauroid
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
''
Equijubus ''Equijubus'' (; ''Mǎzōng'' meaning "horse mane" after the area Mǎzōng Mountain 马鬃山 in which it was found), is a genus of herbivorous hadrosauroid dinosaur from the Early Cretaceous (Albian stage) of northwestern China. Discovery and ...
normani'' from northern China, dating to the
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
stage of the
Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...
approximately 113–100 million years ago, which were found to belong to primitive lineages within Poaceae, similar in position to the Anomochlooideae. These are currently the oldest known grass fossils. Fossils of ''Phragmites'' have been found in the Late Cretaceous of North America, particularly in the Maastrichtian aged
Laramie Formation The Laramie Formation is a geologic formation (geology), formation of the Late Cretaceous (Maastrichtian) age, named by Clarence King in 1876 for exposures in northeastern Colorado, in the United States.King, C. 1876. Report of the Geological Exp ...
. However slightly older fossils of ''Phragmites'' have been found in the Eastern coast of the US dating the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
(such as in the Black Creek Formation). The relationships among the three subfamilies Bambusoideae, Oryzoideae and Pooideae in the BOP clade have been resolved: Bambusoideae and Pooideae are more closely related to each other than to Oryzoideae. This separation occurred within the relatively short time span of about 4 million years. According to Lester Charles King, the spread of grasses in the Late Cenozoic would have changed patterns of
hillslope evolution Hillslope evolution is the changes in the erosion rates, erosion styles and form of slopes of hills and mountains over time. Conceptual models During most of the 20th century three models of hillslope evolution were widely diffused: slope decline, ...
favouring slopes that are convex upslope and concave downslope and lacking a free face were common. King argued that this was the result of more slowly acting surface wash caused by carpets of grass which in turn would have resulted in relatively more soil creep.


Subdivisions

There are about 12,000 grass species in about 771 genera that are classified into 12 subfamilies. See the full
list of Poaceae genera Poaceae, also known as the true grasses, is the fourth largest plant family in the world with around 12,000 species and roughly 800 genera. They contain, among others, the cereal crop species and other plants of economic importance, such as the b ...
. *
Anomochlooideae Anomochlooideae is a subfamily of the true grass family (biology), family Poaceae. It is sister to all the other grasses. It includes perennial herbs that grow on the shaded floor of forests in the Neotropics. There are two genera, ''Anomochloa'' ...
Pilg. ex Potztal, a small lineage of broad-leaved grasses that includes two genera (''Anomochloa'', ''Streptochaeta'') * Pharoideae L.G.Clark & Judz., a small lineage of grasses of three genera, including ''Pharus'' and ''Leptaspis'' * Puelioideae L.G.Clark, M.Kobay., S.Mathews, Spangler & E.A.Kellogg, a small lineage of the African genus ''Puelia'' *
Pooideae The Pooideae are the largest subfamily of the grass family (biology), family Poaceae, with about 4,000 species in 15 tribes and roughly 200 genera. They include some major cereals such as wheat, barley, oat, rye and many lawn and pasture grasses. ...
, including
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
,
barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
, oats, brome-grass ('' Bromus''), reed-grasses (''Calamagrostis'') and many lawn and pasture grasses such as bluegrass (''Poa'') *
Bambusoideae Bamboos are a diverse group of mostly evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in the case of '' Dendrocalamus sinicus'' ...
, including
bamboo Bamboos are a diverse group of mostly evergreen perennial plant, perennial flowering plants making up the subfamily (biology), subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family, in th ...
* Ehrhartoideae, including
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
and wild rice * Aristidoideae, including ''
Aristida ''Aristida'' is a very nearly Cosmopolitan distribution, cosmopolitan genus of plants in the Poaceae, grass family. ''Aristida'' is distinguished by having three Awn (Botany), awns (bristles) on each lemma (botany), lemma of each floret. The gen ...
'' *
Arundinoideae The Arundinoideae are a subfamily of the true grass family Poaceae with around 40 species, including giant reed and common reed. Unlike many other members of the PACMAD clade of grasses, the Arundinoideae all use C3 photosynthesis. Their sist ...
, including giant reed and common reed *
Chloridoideae Chloridoideae is one of the largest subfamily, subfamilies of Poaceae, grasses, with roughly 150 genera and 1,600 species, mainly found in arid tropical or subtropical grasslands. Within the PACMAD clade, their sister group is the Danthonioideae. ...
, including the lovegrasses (''Eragrostis'', about 350 species, including teff), dropseeds (''Sporobolus'', some 160 species),
finger millet Finger millet (''Eleusine coracana'') is an Annual plant, annual herbaceous plant widely grown as a cereal crop in the arid and Semi-arid climate, semiarid areas in Africa and Asia. It is a tetraploid and Self-pollination, self-pollinating speci ...
(''Eleusine coracana'' (L.) Gaertn.), and the muhly grasses (''Muhlenbergia'', about 175 species) *
Panicoideae Panicoideae is the second-largest subfamily of the grasses with over 3,500 species, mainly distributed in warm temperate and tropical regions. It comprises some important agricultural crops, including sugarcane, maize (or corn), sorghum, and s ...
, including panic grass,
maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
,
sorghum ''Sorghum bicolor'', commonly called sorghum () and also known as great millet, broomcorn, guinea corn, durra, imphee, jowar, or milo, is a species in the Poaceae, grass genus ''Sorghum (genus), Sorghum'' cultivated for its grain. The grain i ...
,
sugarcane Sugarcane or sugar cane is a species of tall, Perennial plant, perennial grass (in the genus ''Saccharum'', tribe Andropogoneae) that is used for sugar Sugar industry, production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fib ...
, most
millet Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most millets belong to the tribe Paniceae. Millets are important crops in the Semi-arid climate, ...
s, fonio, " Job's tears", and bluestem grasses * Micrairoideae * Danthonioideae, including pampas grass


Distribution

The grass family is one of the most widely distributed and abundant groups of plants on
Earth Earth is the third planet from the Sun and the only astronomical object known to Planetary habitability, harbor life. This is enabled by Earth being an ocean world, the only one in the Solar System sustaining liquid surface water. Almost all ...
. Grasses are found on every continent, including
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
. The Antarctic hair grass, '' Deschampsia antarctica'' is one of only two flowering plant species native to the western
Antarctic Peninsula The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martin in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctica. ...
.


Ecology

Grasses are the dominant vegetation in many habitats, including
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
, salt-marsh, reedswamp and
steppes In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropical gr ...
. They also occur as a smaller part of the vegetation in almost every other terrestrial habitat. Grass-dominated
biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...
s are called grasslands. If only large, contiguous areas of grasslands are counted, these biomes cover 31% of the planet's land. Grasslands include
pampas The Pampas (; from Quechua 'plain'), also known as the Pampas Plain, are fertile South American low grasslands that cover more than and include the Argentine provinces of Buenos Aires, La Pampa, Santa Fe, Entre Ríos, and Córdoba; all o ...
,
steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...
s, and
prairie Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
s. Grasses provide food to many
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to free range (roam around) and consume wild vegetations in order to feed conversion ratio, convert the otherwise indigestible (by human diges ...
mammals, as well as to many species of
butterflies Butterflies are winged insects from the lepidopteran superfamily Papilionoidea, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The oldest butterfly fossi ...
and
moth Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s. Many types of animals eat grass as their main source of food, and are called '' graminivores'' – these include
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
,
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
,
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s,
rabbit Rabbits are small mammals in the family Leporidae (which also includes the hares), which is in the order Lagomorpha (which also includes pikas). They are familiar throughout the world as a small herbivore, a prey animal, a domesticated ...
s and many
invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...
s, such as
grasshopper Grasshoppers are a group of insects belonging to the suborder Caelifera. They are amongst what are possibly the most ancient living groups of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago. Grassh ...
s and the caterpillars of many brown butterflies. Grasses are also eaten by
omnivorous An omnivore () is an animal that regularly consumes significant quantities of both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize ...
or even occasionally by primarily
carnivorous A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose nutrition and energy requirements are met by consumption of animal tissues (mainly mu ...
animals. Grasses dominate certain
biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...
s, especially temperate grasslands, because many species are adapted to grazing and fire. Grasses are unusual in that the
meristem In cell biology, the meristem is a structure composed of specialized tissue found in plants, consisting of stem cells, known as meristematic cells, which are undifferentiated cells capable of continuous cellular division. These meristematic c ...
is near the bottom of the plant; hence, grasses can quickly recover from cropping at the top. The evolution of large grazing animals in the
Cenozoic The Cenozoic Era ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, insects, birds and angiosperms (flowering plants). It is the latest of three g ...
contributed to the spread of grasses. Without large grazers, fire-cleared areas are quickly colonized by grasses, and with enough rain, tree seedlings. Trees eventually outcompete most grasses. Trampling grazers kill seedling trees but not grasses.


Sexual reproduction and meiosis

Sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...
and
meiosis Meiosis () is a special type of cell division of germ cells in sexually-reproducing organisms that produces the gametes, the sperm or egg cells. It involves two rounds of division that ultimately result in four cells, each with only one c ...
have been studied in
rice Rice is a cereal grain and in its Domestication, domesticated form is the staple food of over half of the world's population, particularly in Asia and Africa. Rice is the seed of the grass species ''Oryza sativa'' (Asian rice)—or, much l ...
,
maize Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout grass that produces cereal grain. It was domesticated by indigenous peoples in southern Mexico about 9,000 years ago from wild teosinte. Native American ...
,
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
and
barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
. Meiosis research in these crop species is linked to crop improvement, since meiotic recombination is an important component of
plant breeding Plant breeding is the science of changing the traits of plants in order to produce desired characteristics. It is used to improve the quality of plant products for use by humans and animals. The goals of plant breeding are to produce crop varie ...
. Unlike in animals, the specification of both male and female plant germlines occurs late in development during flowering. The transition from the
sporophyte A sporophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the biological life cycle, life cycles of plants and algae. It is a diploid multicellular organism which produces asexual Spo ...
phase to the
gametophyte A gametophyte () is one of the two alternating multicellular phases in the life cycles of plants and algae. It is a haploid multicellular organism that develops from a haploid spore that has one set of chromosomes. The gametophyte is the se ...
state is initiated by meiotic entry.


Uses

Grasses are, in human terms, perhaps the most economically important plant family. Their economic importance stems from several areas, including food production, industry, and
lawn A lawn () is an area of soil-covered land planted with Poaceae, grasses and other durable plants such as clover lawn, clover which are maintained at a short height with a lawn mower (or sometimes grazing animals) and used for aesthetic an ...
s. They have been grown as food for
domesticated animals This page gives a list of domesticated animals, also including a list of domestication of animals, animals which are or may be currently undergoing the process of domestication and animals that have an extensive relationship with humans beyond simp ...
for up to 6,000 years and the grains of grasses such as
wheat Wheat is a group of wild and crop domestication, domesticated Poaceae, grasses of the genus ''Triticum'' (). They are Agriculture, cultivated for their cereal grains, which are staple foods around the world. Well-known Taxonomy of wheat, whe ...
, rice, maize (corn) and
barley Barley (), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains; it was domesticated in the Fertile Crescent around 9000 BC, giving it nonshattering spikele ...
have been the most important human food crops. Grasses are also used in the manufacture of
thatch Thatching is the craft of building a roof with dry vegetation such as straw, Phragmites, water reed, Cyperaceae, sedge (''Cladium mariscus''), Juncus, rushes, Calluna, heather, or palm branches, layering the vegetation so as to shed water away fr ...
,
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, Textile, rags, poaceae, grasses, Feces#Other uses, herbivore dung, or other vegetable sources in water. Once the water is dra ...
,
fuel A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work (physics), work. The concept was originally applied solely to those materials capable of releasing chem ...
,
clothing Clothing (also known as clothes, garments, dress, apparel, or attire) is any item worn on a human human body, body. Typically, clothing is made of fabrics or textiles, but over time it has included garments made from animal skin and other thin s ...
, insulation, timber for
fencing Fencing is a combat sport that features sword fighting. It consists of three primary disciplines: Foil (fencing), foil, épée, and Sabre (fencing), sabre (also spelled ''saber''), each with its own blade and set of rules. Most competitive fe ...
,
furniture Furniture refers to objects intended to support various human activities such as seating (e.g., Stool (seat), stools, chairs, and sofas), eating (table (furniture), tables), storing items, working, and sleeping (e.g., beds and hammocks). Furnitur ...
, scaffolding and
construction Construction are processes involved in delivering buildings, infrastructure, industrial facilities, and associated activities through to the end of their life. It typically starts with planning, financing, and design that continues until the a ...
materials, floor matting, sports turf and
baskets A basket is a container that is traditionally constructed from stiff fibers, and can be made from a range of materials, including wood splints, runners, and cane. While most baskets are made from plant materials, other materials such as horse ...
.


Food production

Of all crops grown, 70% are grasses. Agricultural grasses grown for their edible seeds are called ''
cereal A cereal is a grass cultivated for its edible grain. Cereals are the world's largest crops, and are therefore staple foods. They include rice, wheat, rye, oats, barley, millet, and maize ( Corn). Edible grains from other plant families, ...
s'' or ''
grain A grain is a small, hard, dry fruit (caryopsis) – with or without an attached husk, hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and ...
s'' (although the latter term, when used agriculturally, refers to both cereals and similar seeds of other plant species, such as
buckwheat Buckwheat (''Fagopyrum esculentum'') or common buckwheat is a flowering plant in the knotweed family Polygonaceae cultivated for its grain-like seeds and as a cover crop. Buckwheat originated around the 6th millennium BCE in the region of what ...
and
legumes Legumes are plants in the pea family Fabaceae (or Leguminosae), or the fruit or seeds of such plants. When used as a dry grain for human consumption, the seeds are also called pulses. Legumes are grown agriculturally, primarily for human consu ...
). Three cereals—rice, wheat, and maize (corn)—provide more than half of all calories consumed by humans. Cereals constitute the major source of
carbohydrate A carbohydrate () is a biomolecule composed of carbon (C), hydrogen (H), and oxygen (O) atoms. The typical hydrogen-to-oxygen atomic ratio is 2:1, analogous to that of water, and is represented by the empirical formula (where ''m'' and ''n'' ...
s for humans and perhaps the major source of protein; these include rice (in southern and
eastern Asia East Asia is a geocultural region of Asia. It includes China, Japan, Mongolia, North Korea, South Korea, and Taiwan, plus two special administrative regions of China, Hong Kong and Macau. The economies of China, Japan, South Korea, and Taiwan ...
), maize (in Central and
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
), and wheat and barley (in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
, northern Asia and the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
).
Sugarcane Sugarcane or sugar cane is a species of tall, Perennial plant, perennial grass (in the genus ''Saccharum'', tribe Andropogoneae) that is used for sugar Sugar industry, production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fib ...
is the major source of
sugar Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose Glucose is a sugar with the Chemical formula#Molecular formula, molecul ...
production. Additional food uses of sugarcane include sprouted grain, shoots, and
rhizome In botany and dendrology, a rhizome ( ) is a modified subterranean plant stem that sends out roots and Shoot (botany), shoots from its Node (botany), nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from ...
s, and in drink they include
sugarcane juice Sugarcane juice is the liquid extracted from pressed sugarcane. It is consumed as a beverage in many places, especially where sugarcane is commercially grown, such as Southeast Asia, the Indian subcontinent, North Africa, mainly Egypt, and also ...
and plant milk, as well as
rum Rum is a liquor made by fermenting and then distilling sugarcane molasses or sugarcane juice. The distillate, a clear liquid, is often aged in barrels of oak. Rum originated in the Caribbean in the 17th century, but today it is produced i ...
,
beer Beer is an alcoholic beverage produced by the brewing and fermentation of starches from cereal grain—most commonly malted barley, although wheat, maize (corn), rice, and oats are also used. The grain is mashed to convert starch in the ...
,
whisky Whisky or whiskey is a type of liquor made from Fermentation in food processing, fermented grain mashing, mash. Various grains (which may be Malting, malted) are used for different varieties, including barley, Maize, corn, rye, and wheat. Whisky ...
, and
vodka Vodka ( ; is a clear distilled beverage, distilled alcoholic beverage. Its varieties originated in Poland and Russia. Vodka is composed mainly of water and ethanol but sometimes with traces of impurities and flavourings. Traditionally, it is ...
. Bamboo shoots are used in numerous Asian dishes and broths, and are available in supermarkets in various sliced forms, in both fresh, fermented and canned versions. Lemongrass is a grass used as a culinary herb for its citrus-like flavor and scent. Many species of grass are grown as
pasture Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Types of pasture Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, c ...
for foraging or as
fodder Fodder (), also called provender (), is any agriculture, agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, domestic rabbit, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food ...
for prescribed
livestock Livestock are the Domestication, domesticated animals that are raised in an Agriculture, agricultural setting to provide labour and produce diversified products for consumption such as meat, Egg as food, eggs, milk, fur, leather, and wool. The t ...
feeds, particularly in the case of
cattle Cattle (''Bos taurus'') are large, domesticated, bovid ungulates widely kept as livestock. They are prominent modern members of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Mature female cattle are calle ...
,
horse The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 mi ...
s, and
sheep Sheep (: sheep) or domestic sheep (''Ovis aries'') are a domesticated, ruminant mammal typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to d ...
. Such grasses may be cut and stored for later feeding, especially for the winter, in the form of bales of hay or
straw Straw is an agricultural byproduct consisting of the dry wikt:stalk, stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the crop yield, yield by weight of cereal crops such as barley, oats, rice, ry ...
, or in silos as
silage Silage is fodder made from green foliage crops which have been preserved by fermentation (food), fermentation to the point of souring. It is fed to cattle, sheep and other ruminants. The fermentation and storage process is called ''ensilage'', ' ...
. Straw (and sometimes hay) may also be used as bedding for animals. An example of a sod-forming perennial grass used in agriculture is '' Thinopyrum intermedium''.


Industry

Grasses are used as raw material for a multitude of purposes, including construction and in the composition of building materials such as cob, for insulation, in the manufacture of paper and board such as oriented structural straw board. Grass
fiber Fiber (spelled fibre in British English; from ) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often inco ...
can be used for making
paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, Textile, rags, poaceae, grasses, Feces#Other uses, herbivore dung, or other vegetable sources in water. Once the water is dra ...
,
biofuel Biofuel is a fuel that is produced over a short time span from Biomass (energy), biomass, rather than by the very slow natural processes involved in the formation of fossil fuels such as oil. Biofuel can be produced from plants or from agricu ...
production, nonwoven fabrics, and as replacement for glass fibers used in reinforced plastics. Bamboo scaffolding is able to withstand typhoon-force winds that would break steel scaffolding. Larger bamboos and '' Arundo donax'' have stout culms that can be used in a manner similar to timber, ''Arundo'' is used to make reeds for
woodwind instrument Woodwind instruments are a family of musical instruments within the greater category of wind instruments. Common examples include flute, clarinet, oboe, bassoon, and saxophone. There are two main types of woodwind instruments: flutes and ...
s, and bamboo is used for innumerable implements. '' ''Phragmites australis'''' (common reed) is important for thatching and wall construction of homes in Africa. Grasses are used in
water treatment Water treatment is any process that improves the quality of water to make it appropriate for a specific end-use. The end use may be drinking, industrial water supply, irrigation, river flow maintenance, water recreation or many other uses, ...
systems, in
wetland conservation Wetland conservation is aimed at protecting and preserving areas of land including marshes, swamps, bogs, anfensthat are covered by water seasonally or permanently due to a variety of threats from both natural and Anthropogenic hazard, anthropoge ...
and
land reclamation Land reclamation, often known as reclamation, and also known as land fill (not to be confused with a waste landfill), is the process of creating new Terrestrial ecoregion, land from oceans, list of seas, seas, Stream bed, riverbeds or lake ...
, and used to lessen the erosional impact of urban storm water runoff.


Palaeoecological reconstructions

Pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...
morphology, particularly in the ''Poaceae'' family, is key to figuring out their evolutionary relationships and how environments have changed over time. Grass pollen grains, however, often look the same, making it hard to use them for detailed climate or environmental reconstructions. Grass pollen has a single pore and can vary a lot in size, from about 20 to over 100 micrometers, and this size difference has been looked into for clues about past habitats, to tell apart
domesticated Domestication is a multi-generational mutualistic relationship in which an animal species, such as humans or leafcutter ants, takes over control and care of another species, such as sheep or fungi, to obtain from them a steady supply of reso ...
grasses from wild ones, and to indicate various biological features like how they perform
photosynthesis Photosynthesis ( ) is a system of biological processes by which photosynthetic organisms, such as most plants, algae, and cyanobacteria, convert light energy, typically from sunlight, into the chemical energy necessary to fuel their metabo ...
, their breeding systems, and genetic complexity. Yet, there's ongoing debate about how effective pollen size is for piecing together historical landscapes and weather patterns, considering other factors such as genetic material amount might also affect pollen size. Despite these challenges, new techniques in Fourier-Transform Infrared Spectroscopy and improved statistical methods are now helping to better identify these similar-looking pollen types.


Lawn and ornamental use

Grasses are the primary plants used in lawns, which themselves derive from grazed
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s in Europe. They also provide an important means of erosion control (e.g. along roadsides), especially on sloping land. Grass lawns are an important covering of playing surfaces in many sports, including
football (soccer) Association football, more commonly known as football or soccer, is a team sport played between two teams of 11 Football player, players who almost exclusively use their feet to propel a Ball (association football), ball around a rectangular f ...
,
American football American football, referred to simply as football in the United States and Canada and also known as gridiron football, is a team sport played by two teams of eleven players on a rectangular American football field, field with goalposts at e ...
,
tennis Tennis is a List of racket sports, racket sport that is played either individually against a single opponent (singles (tennis), singles) or between two teams of two players each (doubles (tennis), doubles). Each player uses a tennis racket st ...
,
golf Golf is a club-and-ball sport in which players use various Golf club, clubs to hit a Golf ball, ball into a series of holes on a golf course, course in as few strokes as possible. Golf, unlike most ball games, cannot and does not use a standa ...
,
cricket Cricket is a Bat-and-ball games, bat-and-ball game played between two Sports team, teams of eleven players on a cricket field, field, at the centre of which is a cricket pitch, pitch with a wicket at each end, each comprising two Bail (cr ...
,
softball Softball is a Variations of baseball, variation of baseball, the difference being that it is played with a larger ball, on a smaller field, and with only underhand pitches (where the ball is released while the hand is primarily below the ball) ...
and
baseball Baseball is a bat-and-ball games, bat-and-ball sport played between two team sport, teams of nine players each, taking turns batting (baseball), batting and Fielding (baseball), fielding. The game occurs over the course of several Pitch ...
. Ornamental grasses, such as
perennial In horticulture, the term perennial ('' per-'' + '' -ennial'', "through the year") is used to differentiate a plant from shorter-lived annuals and biennials. It has thus been defined as a plant that lives more than 2 years. The term is also ...
bunch grasses, are used in many styles of
garden design Garden design is the art and process of designing and creating plans for layout and planting of garden, gardens and landscapes. Garden design may be done by the garden owner themselves, or by professionals of varying levels of experience and expe ...
for their foliage, inflorescences and seed heads. They are often used in
natural landscaping Natural landscaping, also called native gardening, is the use of native plants including trees, shrubs, groundcover, and grasses which are local to the geographic area of the garden. Benefits Maintenance Natural landscaping is adapted to t ...
,
xeriscaping Xeriscaping is the process of Garden design, landscaping, or gardening, that reduces or eliminates the need for irrigation. It is promoted in regions that do not have accessible, plentiful, or reliable supplies of fresh water and has gained acce ...
and slope and beach stabilization in contemporary landscaping,
wildlife garden A wildlife garden (or habitat garden or backyard restoration) is an Biophysical environment, environment created with the purpose to serve as a sustainable haven for surrounding wildlife. Wildlife gardens contain a variety of habitats that cater t ...
ing, and native plant gardening. They are used as screens and hedges.


Sports turf

Grass playing fields, courses and pitches are the traditional playing surfaces for many
sports Sport is a physical activity or game, often competitive and organized, that maintains or improves physical ability and skills. Sport may provide enjoyment to participants and entertainment to spectators. The number of participants in ...
, including
American football American football, referred to simply as football in the United States and Canada and also known as gridiron football, is a team sport played by two teams of eleven players on a rectangular American football field, field with goalposts at e ...
,
association football Association football, more commonly known as football or soccer, is a team sport played between two teams of 11 Football player, players who almost exclusively use their feet to propel a Ball (association football), ball around a rectangular f ...
,
baseball Baseball is a bat-and-ball games, bat-and-ball sport played between two team sport, teams of nine players each, taking turns batting (baseball), batting and Fielding (baseball), fielding. The game occurs over the course of several Pitch ...
,
cricket Cricket is a Bat-and-ball games, bat-and-ball game played between two Sports team, teams of eleven players on a cricket field, field, at the centre of which is a cricket pitch, pitch with a wicket at each end, each comprising two Bail (cr ...
,
golf Golf is a club-and-ball sport in which players use various Golf club, clubs to hit a Golf ball, ball into a series of holes on a golf course, course in as few strokes as possible. Golf, unlike most ball games, cannot and does not use a standa ...
, and rugby. Grass surfaces are also sometimes used for
horse racing Horse racing is an equestrian performance activity, typically involving two or more horses ridden by jockeys (or sometimes driven without riders) over a set distance for competition. It is one of the most ancient of all sports, as its bas ...
and
tennis Tennis is a List of racket sports, racket sport that is played either individually against a single opponent (singles (tennis), singles) or between two teams of two players each (doubles (tennis), doubles). Each player uses a tennis racket st ...
. Type of maintenance and species of grass used may be important factors for some sports, less critical for others. In some sports facilities, including indoor domes and other places where maintenance of a grass field would be difficult, grass may be replaced with
artificial turf Artificial turf is a surface of synthetic fibers made to look like natural grass, used in sports arenas, residential lawns and commercial applications that traditionally use grass. It is much more durable than grass and easily maintained wi ...
, a synthetic grass-like substitute.


Cricket

In cricket, the pitch is the strip of carefully mowed and rolled grass where the bowler bowls. In the days leading up to the match it is repeatedly mowed and rolled to produce a very hard, flat surface for the ball to bounce off.


Golf

Grass on golf courses is kept in three distinct conditions: that of the ''rough'', the ''fairway'', and the ''putting green''. Grass on the fairway is mown short and even, allowing the player to strike the ball cleanly. Playing from the rough is a disadvantage because the long grass may affect the flight of the ball. Grass on the putting green is the shortest and most even, ideally allowing the ball to roll smoothly over the surface. An entire industry revolves around the development and marketing of turf grass varieties.


Tennis

In tennis, grass is grown on very hard-packed soil, and the bounce of a
tennis ball A tennis ball is a small, hollow ball used in games of tennis and real tennis. Tennis balls are fluorescent yellow in Professional sports, professional competitions, but in Amateur sports, recreational play other colors are also used. Tennis bal ...
may vary depending on the grass's health, how recently it has been mowed, and the wear and tear of recent play. The surface is softer than hard courts and
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolinite, ). Most pure clay minerals are white or light-coloured, but natural clays show a variety of colours from impuriti ...
(other tennis surfaces), so the ball bounces lower, and players must reach the ball faster resulting in a different style of play which may suit some players more than others. Among the world's most prestigious court for grass tennis is Centre Court at
Wimbledon, London Wimbledon () is a suburb of southwest London, England, southwest of Charing Cross; it is the main commercial centre of the London Borough of Merton. Wimbledon had a population of 68,189 in 2011 which includes the electoral wards of Abbey, Wimb ...
, which hosts the final of the annual
Wimbledon Championships The Wimbledon Championships, commonly called Wimbledon, is a tennis tournament organised by the All England Lawn Tennis and Croquet Club in collaboration with the Lawn Tennis Association annually in Wimbledon, London. It is chronologically the ...
in England, one of the four Grand Slam tournaments.


Economically important grasses

A number of grasses are invasive species that damage natural ecosystems, including forms of ''Phragmites australis'' which are native to Eurasia but has spread around the world.


Role in society

Grasses have long had significance in human society. They have been cultivated as feed for people and
domesticated animals This page gives a list of domesticated animals, also including a list of domestication of animals, animals which are or may be currently undergoing the process of domestication and animals that have an extensive relationship with humans beyond simp ...
for thousands of years. The primary ingredient of
beer Beer is an alcoholic beverage produced by the brewing and fermentation of starches from cereal grain—most commonly malted barley, although wheat, maize (corn), rice, and oats are also used. The grain is mashed to convert starch in the ...
is usually barley or wheat, each of which has been used for this purpose for over 4,000 years. In some places, particularly in
suburb A suburb (more broadly suburban area) is an area within a metropolitan area. They are oftentimes where most of a metropolitan areas jobs are located with some being predominantly residential. They can either be denser or less densely populated ...
an areas, the maintenance of a grass lawn is a sign of a homeowner's responsibility to the overall appearance of their neighborhood. One work credits lawn maintenance to: In communities with
drought A drought is a period of drier-than-normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D.  Jiang, A.  Khan, W.  Pokam Mba, D.  Rosenfeld, J. Tierney, ...
problems, watering of lawns may be restricted to certain times of day or days of the week. Many US municipalities and homeowners' associations have rules that require lawns to be maintained to certain specifications, sanctioning those who allow the grass to grow too long. The smell of freshly cut grass is produced mainly by cis-3-Hexenal. Some common
aphorism An aphorism (from Greek ἀφορισμός: ''aphorismos'', denoting 'delimitation', 'distinction', and 'definition') is a concise, terse, laconic, or memorable expression of a general truth or principle. Aphorisms are often handed down by tra ...
s involve grass. For example: * "The grass is always greener on the other side" suggests an alternate state of affairs will always seem preferable to one's own. * "Don't let the grass grow under your feet" tells someone to get moving. * "A
snake Snakes are elongated limbless reptiles of the suborder Serpentes (). Cladistically squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales much like other members of the group. Many species of snakes have s ...
in the grass" means dangers that are hidden. * "When
elephant Elephants are the largest living land animals. Three living species are currently recognised: the African bush elephant ('' Loxodonta africana''), the African forest elephant (''L. cyclotis''), and the Asian elephant ('' Elephas maximus ...
s fight, it is the grass that suffers" tells of bystanders caught in the crossfire. A folk myth about grass is that it refuses to grow where any violent death has occurred.Olmert, Michael (1996). ''Milton's Teeth and Ovid's Umbrella: Curiouser & Curiouser Adventures in History'', p. 208. Simon & Schuster, New York. .


Image gallery

File:Gräsfrön - Grass seeds - 2025.jpg, Grass seeds, a mixture of 90% Festuca rubra and 10% Festuca ovina. File:Ruwbeemdgras Poa trivialis ligula.jpg, Leaves of '' Poa trivialis'' showing the
ligule A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf A leaf (: leaves) is a principal appendage of the plant stem, stem of a vascular plant, usually borne laterally above g ...
s File:Bamboo DSCN2465.jpg, Bamboo stem and leaves, nodes are evident File:Chasmanthium latifolium-spikelet.jpg, A '' Chasmanthium latifolium'' spikelet File:En Spica spiculae.png, Wheat spike and spikelet File:En Aperta.png, Spikelet opened to show caryopsis File:Harestail grass.jpg, Harestail grass File:Grass.jpg, Grass File:Saccharum-officinarum2.JPG, Sugarcane ('' Saccharum officinarum'') File:Bromus hordeaceus unten.jpeg, Roots of '' Bromus hordeaceus'' File:Ohra.jpg, Barley mature spikes ('' Hordeum vulgare'') File:Koeh-283.jpg, Illustration depicting both staminate and pistillate flowers of maize (''
Zea mays Maize (; ''Zea mays''), also known as corn in North American English, is a tall stout Poaceae, grass that produces cereal grain. It was domesticated by indigenous peoples of Mexico, indigenous peoples in southern Mexico about 9,000 years ago ...
'') File:Flowering Grass.JPG, A grass flower head (meadow foxtail) showing the plain-coloured flowers with large anthers File:Grass Anthers.JPG, Anthers detached from a meadow foxtail flower File:Setaria verticillata W IMG 1084.jpg, '' Setaria verticillata'', bristly foxtail File:Setaria verticillata W IMG 1083.jpg, '' Setaria verticillata'', bristly foxtail File:Oryza sativa in Kadavoor.jpg, ''
Oryza sativa ''Oryza sativa'', having the common name Asian cultivated rice, is the much more common of the two rice species cultivated as a cereal, the other species being ''Oryza glaberrima, O. glaberrima'', African rice. It was History of rice cultivation ...
'',
Kerala Kerala ( , ) is a States and union territories of India, state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile ...
,
India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...


See also

* * * * * *


References


External links


Need a Definition of Grass?

Vegetative Key to Grasses

Poaceae
a
''The Plant List''

Learn about grasses
a
''The Story of the Poaceae''
*

a
''The Families of Flowering Plants (DELTA)''


at th
''Angiosperm Phylogeny Website''

''Poaceae Classification''
from the onlin
''Catalogue of New World Grasses''

Poaceae
at the onlin
''Guide to the Flora of Mongolia''

Poaceae
at the onlin
''Flora of Taiwan''

Poaceae
at the onlin
''Flora of Pakistan''

Poaceae
at the onlin
''Flora of Zimbabwe''

Poaceae
at the onlin
''Flora of Western Australia''
* Grasses of Australia (AusGrass2) �
AusGrass2 , Grasses of Australia

Gramineae
at the onlin
''Flora of New Zealand''

NZ Grass Key
An Interactive Key to New Zealand Grasses a
''Landcare Research''

The Grass Genera of the World
a
''DELTA intkey''



''GrassWorld''
{{Authority control Extant Albian first appearances Grasslands Plant life-forms Plants by habit Poales families