Trachilos footprints
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The Trachilos footprints are possibly
tetrapod Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids ( pelycosaurs, extinct t ...
footprints which show hominin-like characteristics from the late
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
on the western
Crete Crete ( el, Κρήτη, translit=, Modern: , Ancient: ) is the largest and most populous of the Greek islands, the 88th largest island in the world and the fifth largest island in the Mediterranean Sea, after Sicily, Sardinia, Cyprus, ...
, close to the village of Trachilos, west of
Kissamos Kissamos ( el, Κίσσαμος) is a town and a municipality in the west of the island of Crete, Greece. It is part of the Chania regional unit and of the former Kissamos Province which covers the northwest corner of the island. The town of Kiss ...
, in the
Chania Prefecture Chania ( el, Περιφερειακή ενότητα Χανίων) is one of the four regional units of Crete; it covers the westernmost quarter of the island. Its capital is the city of Chania. Chania borders only one other regional unit: that ...
. Researchers describe the tracks as representing at least one apparent bipedal
hominin The Hominini form a taxonomic tribe of the subfamily Homininae ("hominines"). Hominini includes the extant genera ''Homo'' (humans) and '' Pan'' (chimpanzees and bonobos) and in standard usage excludes the genus ''Gorilla'' (gorillas). The ...
or an unknown
primate Primates are a diverse order of mammals. They are divided into the strepsirrhines, which include the lemurs, galagos, and lorisids, and the haplorhines, which include the tarsiers and the simians ( monkeys and apes, the latter includin ...
. The stratum in which the
footprint Footprints are the impressions or images left behind by a person walking or running. Hoofprints and pawprints are those left by animals with hooves or paws rather than feet, while "shoeprints" is the specific term for prints made by shoes. The ...
s were found was dated to about 5.7 million years ago, which predates the previously earliest discovered hominin footprints by about two million years. Later studies show that the footprints might be more than 6 million years old. The researchers of the tracks suggest that it may imply the possibility of hominin evolution outside of Africa, contrary to the current
theory A theory is a rational type of abstract thinking about a phenomenon, or the results of such thinking. The process of contemplative and rational thinking is often associated with such processes as observational study or research. Theories may ...
.


Discovery

The tracks were originally discovered by Gerard D. Gierliński, from the Polish Research Institute in
Warsaw Warsaw ( pl, Warszawa, ), officially the Capital City of Warsaw,, abbreviation: ''m.st. Warszawa'' is the capital and largest city of Poland. The metropolis stands on the River Vistula in east-central Poland, and its population is officiall ...
in 2002. During a visit to Trachilos on Crete, Gierliński found the tracks, and as he was not planning on staying in Trachilos, Gierliński recorded the footprints to investigate them in the future. In 2012 Gerard D. Gierliński received permission from the Greek government to research the area, returning to Trachilos with other researchers to explore the tracks in detail. The researchers used methods such as laser scans and 3D imaging of the footprints, and compared them to apes and bears as well as humans.


Dates

The tracks were dated by using the underlying rock bed, predominantly
sedimentary Sedimentary rocks are types of rock that are formed by the accumulation or deposition of mineral or organic particles at Earth's surface, followed by cementation. Sedimentation is the collective name for processes that cause these particles ...
, and
foraminifera Foraminifera (; Latin for "hole bearers"; informally called "forams") are single-celled organisms, members of a phylum or class of amoeboid protists characterized by streaming granular ectoplasm for catching food and other uses; and commonly ...
. The study explains, "The coastal rocks at Trachilos lie within the Platanos Basin, and present a succession of shallow marine late Miocene
carbonates A carbonate is a salt of carbonic acid (H2CO3), characterized by the presence of the carbonate ion, a polyatomic ion with the formula . The word ''carbonate'' may also refer to a carbonate ester, an organic compound containing the carbonate g ...
and siliciclastics ..At the top, this marine succession terminates abruptly in the coarse-grained terrigenous sedimentary rocks of the Hellenikon Group The study continues, stating that the sedimentary rocks would have been created around 5.6 million years ago, at the time of the
Messinian salinity crisis The Messinian salinity crisis (MSC), also referred to as the Messinian event, and in its latest stage as the Lago Mare event, was a geological event during which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (d ...
(mya). The researchers also found evidence of foraminifera, which were dated at 8.5 mya and 3.5 mya. Given the date of the sedimentary rocks and the foraminifera samples, the researches created an approximate interval of 8.5 mya to 5.6 mya. As the rock sediment containing the tracks resembled that of Hellenikon minerals, the tracks were estimated to be 5.7 mya within the given interval. In 2021, further research was published in regard to dating by Kirscher et al. Cyclostratigraphic data based on
magnetic susceptibility In electromagnetism, the magnetic susceptibility (Latin: , "receptive"; denoted ) is a measure of how much a material will become magnetized in an applied magnetic field. It is the ratio of magnetization (magnetic moment per unit volume) to the ap ...
indicated that the Trachilos footprints are about 6.05 Ma old, which is 350,000 years older than was found previously. This puts the Trachilos footprints in the same period as the fossils of ''
Orrorin ''Orrorin tugenensis'' is a postulated early species of Homininae, estimated at and discovered in 2000. It is not confirmed how ''Orrorin'' is related to modern humans. Its discovery was used to argue against the hypothesis that australopithecin ...
tugenensis'' from
Kenya ) , national_anthem = "Ee Mungu Nguvu Yetu"() , image_map = , map_caption = , image_map2 = , capital = Nairobi , coordinates = , largest_city = Nairobi , ...
.


Characteristics

The footprints were measured to range at 94-223 mm (3.7-8.8 inches) long and determined to be oriented in a south-west direction. There are clear pressure indexes, resembling that of a modern Homo sapiens
plantigrade 151px, Portion of a human skeleton, showing plantigrade habit In terrestrial animals, plantigrade locomotion means walking with the toes and metatarsals flat on the ground. It is one of three forms of locomotion adopted by terrestrial mammals. T ...
structure. The researchers also determined the presence of five digits in the imprints, classifying the track maker as pentadactyl, and lacking claws. As there was no visible evidence of forelimbs from the tracks, the track maker was identified as bipedal. Through 3D printing and laser scanning, there are impressions found which indicate a ball region, a pulling up motion of the foot, a
hallux Toes are the digits (fingers) of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being '' digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being '' pl ...
, and small possible gaps between the first and other digit impressions. Poorly preserved prints lack these gaps, however. The lateral digit impressions become progressively smaller so that the digital region as a whole is strongly asymmetrical. The impression of the
hallux Toes are the digits (fingers) of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being '' digitigrade''. Humans, and other animals that walk on the soles of their feet, are described as being '' pl ...
has a narrow neck and bulbous asymmetrical distal pad, indicating that the tracks were entaxonic.
Morphometric Morphometrics (from Greek μορϕή ''morphe'', "shape, form", and -μετρία ''metria'', "measurement") or morphometry refers to the quantitative analysis of ''form'', a concept that encompasses size and shape. Morphometric analyses are co ...
analysis showed the footprints to have outlines that are distinct from modern non-hominin primates and resemble those of hominins. While younger than fossil records of hominins such as ''
Sahelanthropus ''Sahelanthropus tchadensis'' is an extinct species of the Homininae (African apes) dated to about , during the Miocene epoch. The species, and its genus ''Sahelanthropus'', was announced in 2002, based mainly on a partial cranium, nicknamed '' ...
'', found in
Chad Chad (; ar, تشاد , ; french: Tchad, ), officially the Republic of Chad, '; ) is a landlocked country at the crossroads of North and Central Africa. It is bordered by Libya to the north, Sudan to the east, the Central African Republic ...
and dated around seven million years ago, the discovery potentially challenges the generally accepted theory that all early hominins were only present in Africa. The print morphology suggests that the trackmaker could be a basal member of the clade Hominini, but as Crete is some distance outside the known geographical range of pre-
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
hominins, researchers say that there is also a possibility that they represent a hitherto unknown late
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
primate that convergently evolved human-like foot anatomy.


News and controversy

When Gierliński and his team tried to publish the study, they received harsh criticism due to the findings going against the widely-accepted theory of early hominins evolving in
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
alone. According to the study, the Trachilos footprints may represent an early hominin or primate species that may have evolved hominin-like feet independently, outside of Africa. It also suggests the possibility of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
, wherein unrelated species adapt similar traits and characteristics to each other. Although convergent evolution is not an implausible explanation of the hominin-like trackways,, strong evidence is needed to support this interpretation. There were doubts if the tracks were footprints at all. This resulted in rejections from many scientific journals when offered to publish the study's findings. In an interview at the
CBC News CBC News is a division of the Canadian Broadcasting Corporation responsible for the news gathering and production of news programs on the corporation's English-language operations, namely CBC Television, CBC Radio, CBC News Network, and CBC.ca. ...
, researchers claimed that while they were trying to publish their work about the footprints at high-profile publications they got "ferociously aggressive responses", criticism and rejection from reviewers and editors. According to the researchers, “Basically, it wasn't a true
peer review Peer review is the evaluation of work by one or more people with similar competencies as the producers of the work (peers). It functions as a form of self-regulation by qualified members of a profession within the relevant field. Peer review ...
process at all,” “They were just trying to shut us down.” After multiple rejections from other publications, the study was eventually published in the journal, "Proceedings of the
Geologists' Association The Geologists' Association, founded in 1858, is a British organisation with charitable status for those concerned with the study of geology. It publishes the ''Proceedings of the Geologists' Association'' and jointly with the Geological Society ...
." Shortly after the research about the footprints was published, eight prints were chiseled out of the rock and stolen. According to the newspaper ''
Proto Thema Proto or PROTO may refer to: Language * Proto-, an English prefix meaning "first" Media * ''Proto'' (magazine), an American science magazine *Radio Proto in Cyprus Music * ''Proto'' (Holly Herndon album), 2019 * ''Proto'' (Leo O'Kelly ...
'', the culprit was a high school teacher, who was later arrested by Crete authorities at Kasteli, Chania.Chung, Emily (February 23, 2018)
"One Hell of an Impression"
''CBC News''.
The prints were later found in his house and on a farm. David Begun, a paleoanthropologist at the
University of Toronto The University of Toronto (UToronto or U of T) is a public research university in Toronto, Ontario, Canada, located on the grounds that surround Queen's Park. It was founded by royal charter in 1827 as King's College, the first institution ...
, Canada, argues that the footprints might not necessarily belong to a human ancestor, despite its appearance. Similarly, Robin Crompton, a
biological anthropologist Biological anthropology, also known as physical anthropology, is a scientific discipline concerned with the biological and behavioral aspects of human beings, their extinct hominin ancestors, and related non-human primates, particularly from an e ...
at the
University of Liverpool , mottoeng = These days of peace foster learning , established = 1881 – University College Liverpool1884 – affiliated to the federal Victoria Universityhttp://www.legislation.gov.uk/ukla/2004/4 University of Manchester Act 200 ...
, England, believes that although the footprints belong to a bipedal organism, they might not be human ancestors but “made by a member of the great ape clade”. Multiple fossilized footprints must be found and analyzed in order to accurately categorize the tracks by species. Meldrum and Sarmiento (2018) who studied the Trachilos prints (ichnites) did not agree with Gierlinski et al. (2017) that these ichnites were made by a primate or even a vertebrate. Principally, Gierlinski et al. did not provide objective criteria for identifying prints as primate prints. The prints show no consistent and repetitive details that could identify them as being made by a primate, or even an animal with bilateral symmetry. Fatally, Gierlinski et al. did not explore alternative agents that could have produced the prints and provided no explanation as to how their analyses of print outlines deals with missing data and comparisons of non-homologous print outlines. Meldrum and Sarmiento concluded that there was insufficient evidence to argue the prints were made by an animal with bilateral symmetry. The Trachilos ichnites could have been made by non-vertebrate lifeforms or could even be the result of non-organic agents.


See also

*
Convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
*
Early expansions of hominins out of Africa Several expansions of populations of archaic humans (genus ''Homo'') out of Africa and throughout Eurasia took place in the course of the Lower Paleolithic, and into the beginning Middle Paleolithic, between about 2.1 million and 0.2 million ...
*
Messinian salinity crisis The Messinian salinity crisis (MSC), also referred to as the Messinian event, and in its latest stage as the Lago Mare event, was a geological event during which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (d ...


References

{{reflist Fossil trackways Archaeology of Greece Prehistoric Greece Crete Human evolution Hominin fossils