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Sidneyia Inexpectans
''Sidneyia'' is an extinct marine arthropod known from fossils found from the Early to the Mid Cambrian of China and the Mid Cambrian Burgess Shale of British Columbia, Canada. Description Sidneyia size comparison.svg, Size comparison of ''Sidneyia'' species Rsos211134f08 d.jpg, Biramous limb of ''Sidneyia inexpectans'' ''Sidneyia inexpectans'' reached lengths of at least . The largest known specimen of ''S. minor'' is around long and wide, while the largest specimen of ''S. malongensis'' is long and wide. The head shield is short, with notches present on the sides to accommodate stalked eyes, with the underside having a hypostome. The head has a pair of segmented antennae, as well as three pairs of post-antenal appendages. This was followed by a thorax, which had eight to ten segments/ tergites, each associated with a pair of biramous (two branched) appendages, this was followed with one to three abdomen segments/tergites, with the body terminating with a telson, which c ...
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Charles Doolittle Walcott
Charles Doolittle Walcott (March 31, 1850February 9, 1927) was an American paleontologist, administrator of the Smithsonian Institution from 1907 to 1927, and director of the United States Geological Survey. He is famous for his discovery in 1909 of well-preserved fossils, including some of the oldest soft-part imprints, in the Burgess Shale of British Columbia, Canada. Early life Charles Doolittle Walcott was born on March 31, 1850, in New York Mills, New York. His grandfather, Benjamin S. Walcott, moved from Rhode Island in 1822. His father, also Charles Doolittle Walcott, died when Charles Jr. was only two. Walcott was the youngest of four children. He was interested in nature from an early age, collecting minerals and bird eggs and, eventually, fossils. He attended various schools in the Utica area but left at the age of eighteen without completing high school, the end of his formal education. His interest in fossils solidified as he became a commercial fossil collector ...
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Brachiopod
Brachiopods (), phylum (biology), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachio ...
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Thulaspis
''Thulaspis'' is an extinct genus of artiopodan arthropod from the Cambrian Stage 3 aged Sirius Passet site in Greenland. It is thought to be a close relative of '' Squamacula'', and is possibly one of the most basal members of Artiopoda. Description ''Thulaspsis'' was relatively large-sized. The largest specimen is about wide and long. The carapace is overall oval-shaped was convex and likely stiff in life, with a raised central axis. The head is broad and bears a pair of antennae, which have at least 20 rectangular segments, and likely two pairs of biramous (two branched) limbs.The body has 15 thoracic segments, which curve forwards in the first few segments, before becoming straight in the fourth and fifth segments, then increasingly posteriorly curved in subsequent segments. At least the first 11 thoracic segments are associated with pairs of biramous limbs. The biramous limbs have robust endopods (lower, leg-like branches), and flap-shaped exopods (upper branches). The b ...
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Artiopoda
Artiopoda is a clade of extinct arthropods that includes trilobites and their close relatives. It was erected by Hou and Bergström in 1997 to encompass a wide diversity of arthropods that would traditionally have been assigned to the Trilobitomorpha. Trilobites, in part due to abundance of findings owing to their mineralized exoskeletons, are by far the best recorded, diverse, and long lived members of the clade. Other members, which lack mineralised exoskeletons, are known mostly from Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ... deposits. Description According to Stein and Selden (2012) artiopods are recognised by the possession of filiform antennulae, limbs with bilobate exopods (upper branches), with the proximal (closest to base of the limb) lobe being elong ...
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Vicissicaudata
Vicissicaudata is an unranked group of artiopodan arthropods, containing Cheloniellida, Aglaspidida and several other genera outside these groups like ''Sidneyia'' and ''Emeraldella.'' Description Vicissicaudatans are similar to most other artiopods in shape, with rounded bodies consisting of many wide tergites, sometimes with a distinguishable axis along the midline. This clade is defined by a differentiated posterior region, often ending in a telson and caudal appendages. The morphology of this posterior region varies between clades. Aglaspidids, alongside '' Eozetetes'' and '' Carimersa'', have long styliform telsons and relatively small, flap-like caudal appendages while cheloniellids and '' Tardisia'' have much longer caudal appendages similar to the caudal furcae of various taxa, alongside much smaller telsons more resembling the postabdominal segments. The paraphyletic "xenopods" are more similar to those of aglaspidids, also with long telsons and flap-like caudal app ...
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Emeraldella
''Emeraldella'' is a genus of arthropod known from the Middle Cambrian of North America. The type species ''E. brocki'' was described in 1912 from the Burgess Shale. 21 specimens of ''Emeraldella'' are known from the Greater Phyllopod bed, where they comprise < 0.1% of the community. A re-study on the species was done in 2012. A second species ''E. brutoni'' is known from the Wheeler Shale, which was described in 2011. An additional specimen of ''E. brutoni'' was described in 2019, which revealed more of the anatomy. It has been placed as a basal member of the clade Vicissicaudata within Artiopoda, a group of arthropods containing trilobites and their relatives.


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Cambrian Stage 3
Cambrian Stage 3 is the still unnamed third stage of the Cambrian. It succeeds Cambrian Stage 2 and precedes Cambrian Stage 4, although neither its base nor top have been formally defined. The plan is for its lower boundary to correspond approximately to the first appearance of trilobites, about million years ago, though the globally asynchronous appearance of trilobites warrants the use of a separate, globally synchronous marker to define the base. The upper boundary and beginning of Cambrian Stage 4 is informally defined as the first appearance of the trilobite genera ''Olenellus'' or ''Redlichia'' around million years ago. Naming The International Commission on Stratigraphy has not officially named the 3rd stage of the Cambrian. The stage approximately corresponds to the "Atdabanian", which is used by geologists working in Siberia. Biostratigraphy The oldest trilobite known is ''Lemdadella'' which appears at the beginning of the ''Fallotaspis'' zone. The Cambrian radiation of ...
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Mantou Formation
''Mantou'' ( zh, t=饅頭, s=馒头, first=t), often referred to as a Chinese steamed bun, is a white and soft type of steamed bread or bun popular in northern China. Folk etymology connects the name ''mantou'' to a tale about Zhuge Liang. Description ''Mantou'' are typically eaten as a staple food in northern parts of China where wheat, rather than rice, is grown. They are made with milled wheat flour, water and leavening agents. In size and texture, they range from , soft and fluffy in the most elegant restaurants, to over , firm and dense for the working man's lunch. As white flour, being more heavily processed, was once more expensive, white ''mantou'' were something of a luxury in preindustrial China. Traditionally, ''mantou'', ''bing'', and wheat noodles were the staple carbohydrates of the northern Chinese diet, analogous to rice, which forms the mainstay of the southern Chinese diet. They are also known in the south but are often served as street food or in rest ...
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Wuliuan
The Wuliuan stage is the fifth stage of the Cambrian, and the first stage of the Miaolingian Series of the Cambrian. It was formally defined by the International Commission on Stratigraphy in 2018. Its base is defined by the first appearance of the trilobite species '' Oryctocephalus indicus''; it ends with the beginning of the Drumian Stage, marked by the first appearance of the trilobite '' Ptychagnostus atavus'' around million years ago. The 'golden spike' that formally defines the base of the age is driven into the Wuliu-Zengjiayan (乌溜-曾家崖) section of the Kaili formation, near Balang Village in the Miaoling Mountains, Guizhou, China. GSSP Three sections were discussed as GSSP candidates: the Wuliu-Zengjiayan section near Balang in Guizhou province (China), a section on Split Mountain in Nevada (USA) and the "Molodo river section" along the Molodo river (Sakha Republic, Russia). The Wuliu-Zengjiayan section is an outcrop of the Kaili Formation in the Wuliu quarr ...
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Sirius Passet
Sirius Passet is a Cambrian Lagerstätte in Peary Land, Greenland. The Sirius Passet Lagerstätte was named after the Slædepatruljen Sirius, Sirius sledge patrol that operates in North Greenland. It comprises six places in Nansen Land, on the east shore of J.P. Koch Fjord in the far north of Greenland. It was discovered in 1984 by A. Higgins of the Geological Survey of Denmark and Greenland, Geological Survey of Greenland. A preliminary account was published by Simon Conway Morris and others in 1987 and expeditions led by J. S. Peel and Conway Morris have returned to the site several times between 1989 and the present. A field collection of perhaps 10,000 fossil specimens has been amassed. It is a part of the Buen Formation. Age The fauna is inevitably compared to that of the Burgess Shale, although it is probably ten to fifteen million years older – vs. ) – and more closely contemporaneous with the fauna of the Maotianshan shales from Chengjiang, which are dated to . Pr ...
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Spence Shale
The Spence Shale is the middle member of the Langston Formation in southeastern Idaho and northeastern Utah. It is exposed in the Bear River Range, the Wasatch Range and the Wellsville Mountains. It is known for its abundant Cambrian trilobites and the preservation of Burgess Shale-type fossils. The type locality is Spence Gulch in southeastern Idaho, near the town of Liberty. It was first described by Charles Doolittle Walcott in 1908. Stratigraphy The Spence Shale spans the '' Albertella'' and '' Glossopleura'' biozones. Fauna Generic list of the fauna of the Spence Shale:Hammersburg, S.R., Hasiotis, S.T., Robison, R.R. 2018. Ichnotaxonomy of the Cambrian Spence Shale Member of the Langston Formation, Wellsville Mountains, Northern Utah, USA. Paleontological Contributions, 20, 1–66. Arthropoda Soft-bodied *''Anomalocaris'' *'' Canadaspis'' *''Caryosyntrips'' *'' Dioxycaris'' *'' Hurdia'' *''Isoxys'' *''Leanchoilia'' *'' Meristosoma'' *'' Mollisonia'' *''Sidneyia' ...
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Chengjiang Biota
The Maotianshan Shales () are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation or Heilinpu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized organisms or traces. The Maotianshan Shales form one of some forty Cambrian fossil locations worldwide exhibiting exquisite preservation of rarely preserved, non-mineralized soft tissue, comparable to the fossils of the Burgess Shale of British Columbia, Canada. They take their name from Maotianshan Hill () in Chengjiang County, Yunnan Province, China. The most famous assemblage of organisms are referred to as the Chengjiang biota for the multiple scattered fossil sites in Chengjiang. The age of the Chengjiang Lagerstätte is locally termed Qiongzhusian, a stage correlated to the late Atdabanian Stage in Siberian sequences of the middle of the Early Cambrian. The shales date to ≤. Along with the Burgess Shale, the Maotianshan Shales ar ...
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