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Protorothyrididae
Protorothyrididae is an extinct family of small, lizard-like reptiles belonging to Eureptilia. Their skulls did not have fenestrae, like the more derived diapsids. Protorothyridids lived from the Late Carboniferous to Early Permian periods, in what is now North America. Many genera of primitive reptiles were thought to be protorothyridids. '' Brouffia'', '' Coelostegus'', '' Paleothyris'' and '' Hylonomus'', for example, were found to be more basal eureptiles in Muller and Reisz (2006), making the family as historically defined paraphyletic, though three genera, ''Protorothyris, Anthracodromeus,'' and '' Cephalerpeton'', were recovered as a monophyletic group. ''Anthracodromeus'', ''Paleothyris'', and ''Protorothyris w''e recovered as a monophyletic group in Ford and Benson (2020) (who did not sample ''Cephalerpeton''), who recovered them as more derived than captorhinids Captorhinidae (also known as cotylosaurs) is an extinct family of tetrapods, traditionally consi ...
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Cephalerpeton
''Cephalerpeton'' is an extinct genus of " protorothyridid" eureptile known from the Late Carboniferous (late Westphalian stage) of Illinois. It is known from the holotype YPM 796, a partial skeleton. It was collected in the Mazon Creek site, from the Francis Creek Shale Member of the Carbondale Formation. It was first named by R. L. Moodie in 1912 as an amphibamid amphibian and the type species is ''Cephalerpeton ventriarmatum''. It was first assigned to Protorothyrididae Protorothyrididae is an extinct family of small, lizard-like reptiles belonging to Eureptilia. Their skulls did not have fenestrae, like the more derived diapsids. Protorothyridids lived from the Late Carboniferous to Early Permian periods, ... by Robert L. Carroll and Donald Baird in 1972 and this placement has been widely accepted. References Prehistoric romeriids Fossil taxa described in 1912 Carboniferous reptiles of North America Prehistoric reptile genera {{carboniferous- ...
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Eureptile
Eureptilia ("true reptiles") is one of the two major subgroups of the clade Sauropsida, the other one being Parareptilia. Eureptilia includes Diapsida (the clade containing all modern reptiles and birds), as well as a number of primitive Permo-Carboniferous forms previously classified under Anapsida, in the old (no longer recognised) order "Cotylosauria". Eureptilia is characterized by the skull having greatly reduced supraoccipital, tabular, and supratemporal bones that are no longer in contact with the postorbital. Aside from Diapsida, the group notably contains Captorhinidae, a diverse and long lived (Late Carboniferous-Late Permian) clade of initially small carnivores that later evolved into large herbivores. Other primitive eureptiles such as ''Hylonomus'' and " protorothyrids" were all small, superficially lizard-like forms, that were probably insectivorous. One primitive eureptile, the Late Carboniferous "protorothyrid" '' Anthracodromeus,'' is the oldest known climbin ...
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Eureptilia
Eureptilia ("true reptiles") is one of the two major subgroups of the clade Sauropsida, the other one being Parareptilia. Eureptilia includes Diapsida (the clade containing all modern reptiles and birds), as well as a number of primitive Permo-Carboniferous forms previously classified under Anapsida, in the old (no longer recognised) order "Cotylosauria". Eureptilia is characterized by the skull having greatly reduced supraoccipital, tabular, and supratemporal bones that are no longer in contact with the postorbital. Aside from Diapsida, the group notably contains Captorhinidae, a diverse and long lived (Late Carboniferous-Late Permian) clade of initially small carnivores that later evolved into large herbivores. Other primitive eureptiles such as ''Hylonomus'' and "protorothyrids" were all small, superficially lizard-like forms, that were probably insectivorous. One primitive eureptile, the Late Carboniferous "protorothyrid" ''Anthracodromeus,'' is the oldest known climbing ...
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Captorhinids
Captorhinidae (also known as cotylosaurs) is an extinct family of tetrapods, traditionally considered primitive reptiles, known from the late Carboniferous to the Late Permian. They had a cosmopolitan distribution across Pangea. Description Captorhinids are a clade of small to very large lizard-like reptiles that date from the late Carboniferous through the Permian. Their skulls were much stronger than those of their relatives, the Protorothyrididae, and had teeth that were better able to deal with tough plant material. The postcranial skeleton is very similar to that of advanced reptiliomorph amphibians, so much in fact that the amphibian Seymouriamorpha and Diadectomorpha were thought to be reptiles and grouped together in "Cotylosauria" as the first reptiles in the early 20th century. Captorhinids have broad, robust skulls that are generally triangular in shape when seen in dorsal view. The premaxillae are characteristically downturned. The largest captorhinid, the herbivor ...
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Amniote
Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are distinguished from the other tetrapod clade — the amphibians — by the development of three extraembryonic membranes ( amnion for embryoic protection, chorion for gas exchange, and allantois for metabolic waste disposal or storage), thicker and more keratinized skin, and costal respiration (breathing by expanding/constricting the rib cage). All three main features listed above, namely the presence of an amniotic buffer, water-impermeable cutes and a robust respiratory system, are very important for amniotes to live on land as true terrestrial animals – the ability to reproduce in locations away from water bodies, better homeostasis in drier environments, and more efficient air respiration to power terrestrial locomotions, althou ...
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Protorothyris Archeri
''Protorothyris'' is an extinct genus of Early Permian protorothyridid known from Texas and West Virginia of the United States. It was first named by Llewellyn Ivor Price in 1937 and the type species is ''Protorothyris archeri''. ''P. archeri'' is known from the holotype MCZ 1532, a three-dimensionally preserved skull and from the referred specimens, which come from four additional individuals, MCZ 2147-2150. All specimens were collected in the Cottonwood Creek site, from the Archer City Formation, Texas, dating to the Asselian stage of the Cisuralian epoch, about 299–294.6 million years ago. A second species, ''P. morani'', was first named by Alfred Sherwood Romer in 1952 with its own generic name, ''Melanothyris''. In 1973, J. Clark and Robert L. Carroll recombined ''P. morani'' as a ''Protorothyris'' species. It is known from the holotype CM 8617, a three-dimensionally preserved skull. It was collected in the Blacksville site, from the Washington Formatio ...
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Anthracodromeus
''Anthracodromeus'' is an extinct genus of Late Carboniferous (late Westphalian stage) " protorothyridid" eureptile known from Ohio. It is known from the holotype AMNH 6940, a nearly complete skeleton. It was collected in the Linton site in Jefferson County, from the upper Freeport Coal Member. ''A. longipes'' was first assigned by Edward Drinker Cope in 1875 to a species of '' Sauropleura''. The genus was first named by Robert L. Carroll and Donald Baird in 1972 and the type species is ''Anthracodromeus longipes''. It is amongst the oldest known tetrapods to display adaptations for climbing. Features and environment ''Anthracodromeus'' was between 15 and 20 cm long when alive, with about 5 cm of this being tail. It had a long body, with 27 vertebrae between the five cervical vertebrae and the sacrum. Each vertebra had a short pair of ribs attached to it. The limbs were quite short, but each foot was at least 2 cm long - as long as the rest of the limb pu ...
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Protorothyris
''Protorothyris'' is an extinct genus of Early Permian protorothyridid known from Texas and West Virginia of the United States. It was first named by Llewellyn Ivor Price in 1937 and the type species is ''Protorothyris archeri''. ''P. archeri'' is known from the holotype MCZ 1532, a three-dimensionally preserved skull and from the referred specimens, which come from four additional individuals, MCZ 2147-2150. All specimens were collected in the Cottonwood Creek site, from the Archer City Formation, Texas, dating to the Asselian stage of the Cisuralian epoch, about 299–294.6 million years ago. A second species, ''P. morani'', was first named by Alfred Sherwood Romer in 1952 with its own generic name, ''Melanothyris''. In 1973, J. Clark and Robert L. Carroll recombined ''P. morani'' as a ''Protorothyris'' species. It is known from the holotype CM 8617, a three-dimensionally preserved skull. It was collected in the Blacksville site, from the Washington Formation ...
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Hylonomus
''Hylonomus'' (; ''hylo-'' "forest" + ''nomos'' "dweller") is an extinct genus of reptile that lived 312 million years ago during the Late Carboniferous period. It is the earliest unquestionable reptile (''Westlothiana'' is older, but in fact it may have been an amphibian, and '' Casineria'' is rather fragmentary). The only species is the type species ''Hylonomus lyelli''. Despite being amongst the oldest known reptiles, it is not the most primitive member of group, being a eureptile more derived than either parareptiles or captorhinids. Description ''Hylonomus'' was long (including the tail). Most of them are 20 cm long and probably would have looked rather similar to modern lizards. It had small sharp teeth and it likely ate small invertebrates such as millipedes or early insects. Fossils of ''Hylonomus'' have been found in the remains of fossilized club moss stumps in the Joggins Formation, Joggins, Nova Scotia, Canada. It is supposed that, after harsh weather, the ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, in the ICS geologic timescale, the younger of two subperiods (or upper of two subsystems) of the Carboniferous Period. It lasted from roughly . As with most other geochronologic units, the rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal-productive beds of this age are widespread. The division between Pennsylvanian and Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian and the Permian. In parts of Europe, the Mississippian and Pennsylvanian are one more-or-less continuous sequence of lowland continental deposits and are grouped t ...
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Paraphyletic
In taxonomy, a group is paraphyletic if it consists of the group's last common ancestor and most of its descendants, excluding a few monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia ( reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancestor, including all extant reptiles as well as the extinct synaps ...
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Monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic groups are typically characterised by shared derived characteristics ( synapomorphies), which distinguish organisms in the clade from other organisms. An equivalent term is holophyly. The word "mono-phyly" means "one-tribe" in Greek. Monophyly is contrasted with paraphyly and polyphyly as shown in the second diagram. A ''paraphyletic group'' consists of all of the descendants of a common ancestor minus one or more monophyletic groups. A '' polyphyletic group'' is characterized by convergent features or habits of scientific interest (for example, night-active primates, fruit trees, aquatic insects). The features by which a polyphyletic group is differentiated from others are not inherited from a common ancestor. These definitions have ta ...
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