Pakaraimoideae
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Pakaraimoideae
''Pakaraimaea'' is a genus of trees in the family Cistaceae. The genus contains a single species, ''Pakaraimaea dipterocarpacea'', from South America. The species is found in the western highlands of Guyana and in adjacent Bolivar State in Venezuela. It maintains strong ectomycorrhizal associations with a wide variety of fungal species. The trees can sometimes be seen forming large stands in the western Guyanas. As of APG IV The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was publishe ..., the species has been moved out of the Dipterocarpaceae (formerly in subfamily Pakaraimoideae) and is now placed within an expanded Cistaceae due to molecular evidence showing that it is sister to the remainder of Cistaceae. References Dipterocarpaceae Monotypic Malvales genera Cistaceae {{ ...
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Dipterocarpaceae
Dipterocarpaceae is a family of 16 genera and about 695 known species of mainly tropical lowland rainforest trees. The family name, from the type genus ''Dipterocarpus'', is derived from Greek (''di'' = two, ''pteron'' = wing and ''karpos'' = fruit) and refers to the two-winged fruit. The largest genera are ''Shorea'' (196 species), ''Hopea'' (104 species), ''Dipterocarpus'' (70 species), and ''Vatica'' (65 species).Ashton, P.S. Dipterocarpaceae. In ''Tree Flora of Sabah and Sarawak,'' Volume 5, 2004. Soepadmo, E., Saw, L. G. and Chung, R. C. K. eds. Government of Malaysia, Kuala Lumpur, Malaysia. Many are large forest-emergent species, typically reaching heights of 40–70 m, some even over 80 m (in the genera ''Dryobalanops'', ''Hopea'' and ''Shorea''), with the tallest known living specimen (''Shorea faguetiana'') 93.0 m tall. The species of this family are of major importance in the timber trade. Their distribution is pantropical, from northern South America to Africa, the S ...
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Plantae
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclude the fungi and some algae, as well as the prokaryotes (the archaea and bacteria). By one definition, plants form the clade Viridiplantae (Latin name for "green plants") which is sister of the Glaucophyta, and consists of the green algae and Embryophyta (land plants). The latter includes the flowering plants, conifers and other gymnosperms, ferns and their allies, hornworts, liverworts, and mosses. Most plants are multicellular organisms. Green plants obtain most of their energy from sunlight via photosynthesis by primary chloroplasts that are derived from endosymbiosis with cyanobacteria. Their chloroplasts contain chlorophylls a and b, which gives them their green color. Some plants are parasitic or mycotrophic and have lost the abilit ...
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Guyana
Guyana ( or ), officially the Cooperative Republic of Guyana, is a country on the northern mainland of South America. Guyana is an indigenous word which means "Land of Many Waters". The capital city is Georgetown. Guyana is bordered by the Atlantic Ocean to the north, Brazil to the south and southwest, Venezuela to the west, and Suriname to the east. With , Guyana is the third-smallest sovereign state by area in mainland South America after Uruguay and Suriname, and is the second-least populous sovereign state in South America after Suriname; it is also one of the least densely populated countries on Earth. It has a wide variety of natural habitats and very high biodiversity. The region known as " the Guianas" consists of the large shield landmass north of the Amazon River and east of the Orinoco River known as the "land of many waters". Nine indigenous tribes reside in Guyana: the Wai Wai, Macushi, Patamona, Lokono, Kalina, Wapishana, Pemon, Akawaio and Warao. ...
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APG IV System
The APG IV system of flowering plant classification is the fourth version of a modern, mostly molecular-based, system of plant taxonomy for flowering plants (angiosperms) being developed by the Angiosperm Phylogeny Group (APG). It was published in 2016, seven years after its predecessor the APG III system was published in 2009, and 18 years after the first APG system was published in 1998. In 2009, a linear arrangement of the system was published separately; the APG IV paper includes such an arrangement, cross-referenced to the 2009 one. Compared to the APG III system, the APG IV system recognizes five new orders (Boraginales, Dilleniales, Icacinales, Metteniusales and Vahliales), along with some new families, making a total of 64 angiosperm orders and 416 families. In general, the authors describe their philosophy as "conservative", based on making changes from APG III only where "a well-supported need" has been demonstrated. This has sometimes resulted in placements that ar ...
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The Guianas
The Guianas, sometimes called by the Spanish loan-word ''Guayanas'' (''Las Guayanas''), is a region in north-eastern South America which includes the following three territories: * French Guiana, an overseas department and region of France * Guyana, formerly known as British Guiana from 1831 until 1966, after the colonies of Berbice, Essequibo, and Demerara, taken from the Netherlands in 1814, were merged into a single colony * Suriname, formerly Dutch Guiana, until 1814 together with Berbice, Essequibo and Demerara In the wider context, the Guianas also includes the following two territories: * Guayana Region in eastern Venezuela ( Amazonas, Bolívar, and Delta Amacuro states), formerly the Guayana Province, alternatively known as Spanish Guayana * State of Amapá in northern Brazil, known as Portuguese Guiana (or Brazilian Guiana) History Pre-colonial period Before the arrival of European colonials, the Guianas were populated by scattered bands of native Arawak ...
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Ectomycorrhiza
An ectomycorrhiza (from Greek ἐκτός ', "outside", μύκης ', "fungus", and ῥίζα ', "root"; pl. ectomycorrhizas or ectomycorrhizae, abbreviated EcM) is a form of symbiotic relationship that occurs between a fungal symbiont, or mycobiont, and the roots of various plant species. The mycobiont is often from the phyla Basidiomycota and Ascomycota, and more rarely from the Zygomycota. Ectomycorrhizas form on the roots of around 2% of plant species, usually woody plants, including species from the birch, dipterocarp, myrtle, beech, willow, pine and rose families. Research on ectomycorrhizas is increasingly important in areas such as ecosystem management and restoration, forestry and agriculture. Unlike other mycorrhizal relationships, such as arbuscular mycorrhiza and ericoid mycorrhiza, ectomycorrhizal fungi do not penetrate their host's cell walls. Instead they form an entirely intercellular interface known as the Hartig net, consisting of highly branched hyp ...
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Venezuela
Venezuela (; ), officially the Bolivarian Republic of Venezuela ( es, link=no, República Bolivariana de Venezuela), is a country on the northern coast of South America, consisting of a continental landmass and many islands and islets in the Caribbean Sea. It has a territorial extension of , and its population was estimated at 29 million in 2022. The capital and largest urban agglomeration is the city of Caracas. The continental territory is bordered on the north by the Caribbean Sea and the Atlantic Ocean, on the west by Colombia, Brazil on the south, Trinidad and Tobago to the north-east and on the east by Guyana. The Venezuelan government maintains a claim against Guyana to Guayana Esequiba. Venezuela is a federal presidential republic consisting of 23 states, the Capital District and federal dependencies covering Venezuela's offshore islands. Venezuela is among the most urbanized countries in Latin America; the vast majority of Venezuelans live in the cities of th ...
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South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the southern subregion of a single continent called America. South America is bordered on the west by the Pacific Ocean and on the north and east by the Atlantic Ocean; North America and the Caribbean Sea lie to the northwest. The continent generally includes twelve sovereign states: Argentina, Bolivia, Brazil, Chile, Colombia, Ecuador, Guyana, Paraguay, Peru, Suriname, Uruguay, and Venezuela; two dependent territories: the Falkland Islands and South Georgia and the South Sandwich Islands; and one internal territory: French Guiana. In addition, the ABC islands of the Kingdom of the Netherlands, Ascension Island (dependency of Saint Helena, Ascension and Tristan da Cunha, a British Overseas Territory), Bouvet Island ( dependency ...
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Angiosperms
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in ...
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Tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are usable as lumber or plants above a specified height. In wider definitions, the taller palms, tree ferns, bananas, and bamboos are also trees. Trees are not a taxonomic group but include a variety of plant species that have independently evolved a trunk and branches as a way to tower above other plants to compete for sunlight. The majority of tree species are angiosperms or hardwoods; of the rest, many are gymnosperms or softwoods. Trees tend to be long-lived, some reaching several thousand years old. Trees have been in existence for 370 million years. It is estimated that there are some three trillion mature trees in the world. A tree typically has many secondary branches supported clear of the ground by the trunk. This trunk typicall ...
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Peter Shaw Ashton
Peter Shaw Ashton (born 27 June 1934) is a British botanist. He is Charles Bullard Professor of Forestry at Harvard University, and director of the Arnold Arboretum there from 1978 to 1987. Born in Boscombe, Bournemouth, England, Ashton received his B.A. in Biology (1956), M.A. in Biology (1960) and Ph.D. Botany (1960) from the University of Cambridge. He has worked for many years on research projects to promote the conservation and sustainable use of tropical forests, and was instrumental in the project by the Center for Tropical Forest Science to formulate a network of Forest Dynamic Plots which are surveyed regularly to sample the health of the forest; he won the Japan Prize for this in 2007. In 1983, Ashton was elected a Fellow of the American Academy of Arts and Sciences The American Academy of Arts and Sciences (abbreviation: AAA&S) is one of the oldest learned societies in the United States. It was founded in 1780 during the American Revolution by John Adams, J ...
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