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In human anatomy, the third trochanter is a bony projection occasionally present on the proximal
femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...
near the
superior Superior may refer to: *Superior (hierarchy), something which is higher in a hierarchical structure of any kind Places * Superior (proposed U.S. state), an unsuccessful proposal for the Upper Peninsula of Michigan to form a separate state *Lak ...
border of the
gluteal tuberosity The gluteal tuberosity is the lateral one of the three upward prolongations of the linea aspera of the femur, extending to the base of the greater trochanter. It serves as the principal insertion site for the gluteus maximus muscle. Structure ...
. When present, it is oblong, rounded, or conical in shape and sometimes continuous with the gluteal ridge. It generally occurs bilaterally without significant side to side dimorphism. A structure of minor importance in humans, the incidence of the third trochanter varies from 17 to 72% between ethnic groups and it is frequently reported as more common in females than in males. Structures analogous to the third trochanter are present in other mammals, including some primates, but also in horses. It is called the third trochanter in reference to the greater and
lesser trochanter In human anatomy, the lesser trochanter is a conical, posteromedial, bony projection from the shaft of the femur. It serves as the principal insertion site of the iliopsoas muscle. Structure The lesser trochanter is a conical posteromedial p ...
s that are always present on the femur.


Function

Its function is to provide an attachment for the ascending tendon of the
gluteus maximus muscle The gluteus maximus is the main extensor muscle of the hip in humans. It is the largest and outermost of the three gluteal muscles and makes up a large part of the shape and appearance of each side of the hips. It is the single largest muscle i ...
. It may function as (1) a reinforcement mechanism for the proximal femoral
diaphysis The diaphysis (: diaphyses) is the main or midsection (shaft) of a long bone. It is made up of cortical bone and usually contains bone marrow and adipose tissue (fat). It is a middle tubular part composed of compact bone which surrounds a centr ...
in response to increased ground reaction force and (2) to increase the attachment surface area for the gluteal musculature and thereby providing greater efficiency of contracture. Mechanical load from the gluteus maximus may, on the other hand, affect the morphology of the proximal femur, similar to how
quadriceps The quadriceps femoris muscle (, also called the quadriceps extensor, quadriceps or quads) is a large muscle group that includes the four prevailing muscles on the front of the thigh. It is the sole extensor muscle of the knee, forming a large ...
determines the size and shape of the
tibial tuberosity The tuberosity of the tibia, tibial tuberosity or tibial tubercle is an elevation on the proximal, anterior aspect of the tibia, just below where the anterior surfaces of the lateral and medial tibial condyles end. Structure The tuberosity of ...
, and thereby the shape of the third trochanter. However, no certain conclusions regarding the biomechanical role of the third trochanter can be made.


Anthropology

The third trochanter is associated with short, robust femora, and is frequently present and well developed in
Neanderthal Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinction, extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle Pleistocene, Middle to Late Plei ...
s (that were shorter and more robust than
anatomically modern humans Early modern human (EMH), or anatomically modern human (AMH), are terms used to distinguish ''Homo sapiens'' ( sometimes ''Homo sapiens sapiens'') that are anatomically consistent with the range of phenotypes seen in contemporary humans, from ...
) but generally absent in
higher primates The simians, anthropoids, or higher primates are an infraorder (Simiiformes ) of primates containing all animals traditionally called monkeys and apes. More precisely, they consist of the parvorders Platyrrhini (New World monkeys) and Catarrh ...
. Studying fossils of ''
Ardipithecus ramidus ''Ardipithecus ramidus'' is a species of australopithecine from the Afar region of Early Pliocene Ethiopia 4.4 million years ago (mya). The species ''A. ramidus'' is the type species for the genus ''Ardipithecus''. There is an older species in t ...
'' (a
chimpanzee–human last common ancestor The chimpanzee–human last common ancestor (CHLCA) is the last common ancestor shared by the extant ''Homo'' (human) and '' Pan'' (chimpanzee and bonobo) genera of Hominini. Estimates of the divergence date vary widely from thirteen to five mill ...
candidate), noted that in this species—as well as in ''
Proconsul A proconsul was an official of ancient Rome who acted on behalf of a Roman consul, consul. A proconsul was typically a former consul. The term is also used in recent history for officials with delegated authority. In the Roman Republic, military ...
'', '' Nacholapithecus'', and ''
Dryopithecus ''Dryopithecus'' is a genus of extinct great apes from the middle–late Miocene boundary of Europe 12.5 to 11.1 million years ago (mya). Since its discovery in 1856, the genus has been subject to taxonomic turmoil, with numerous new species b ...
'' (
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
primates)—
homologs Homologous chromosomes or homologs are a set of one maternal and one paternal chromosome that pair up with each other inside a cell during meiosis. Homologs have the same genes in the same loci, where they provide points along each chromosome th ...
to the third trochanter and the hypotrochanteric fossa are present while both traits are absent in extant apes. They concluded that the
hominid The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
morphotype In biology, polymorphism is the occurrence of two or more clearly different morphs or forms, also referred to as alternative ''phenotypes'', in the population of a species. To be classified as such, morphs must occupy the same habitat at the s ...
is primitive and the femoral shape of apes is derived contrary to what was previously thought.


See also

*
Fourth trochanter The fourth trochanter is a shared characteristic common to archosaurs. It is a protrusion on the posterior-medial side of the middle of the femur shaft that serves as a muscle attachment, mainly for the '' musculus caudofemoralis longus'', the m ...


Notes


References

* * * {{Authority control Bones of the lower limb Femur