Muscardinus Avellanarius Con Piccolo 2
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The hazel dormouse or common dormouse (''Muscardinus avellanarius'') is a small
dormouse A dormouse is a rodent of the family Gliridae (this family is also variously called Myoxidae or Muscardinidae by different taxonomists). Dormice are nocturnal animals found in Africa, Asia, and Europe. They are named for their long, dormant hibe ...
species native to Europe and the only living species in the
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Muscardinus''.


Distribution and habitat

The hazel dormouse is native to northern Europe and Asia Minor. It is the only
dormouse A dormouse is a rodent of the family Gliridae (this family is also variously called Myoxidae or Muscardinidae by different taxonomists). Dormice are nocturnal animals found in Africa, Asia, and Europe. They are named for their long, dormant hibe ...
native to the
British Isles The British Isles are an archipelago in the Atlantic Ocean, North Atlantic Ocean off the north-western coast of continental Europe, consisting of the islands of Great Britain, Ireland, the Isle of Man, the Inner Hebrides, Inner and Outer Hebr ...
, and is therefore often referred to simply as the "dormouse" in British sources, although the
edible dormouse ''Glis'' is a genus of rodent that contains two extant species, both known as edible dormice or fat dormice: the European edible dormouse ''(Glis glis'') and the Iranian edible dormouse (''Glis persicus''). It also contains a number of fossil spe ...
, ''Glis glis'', has been accidentally introduced and now has an established population in South East England. Though
Ireland Ireland (, ; ; Ulster Scots dialect, Ulster-Scots: ) is an island in the North Atlantic Ocean, in Northwestern Europe. Geopolitically, the island is divided between the Republic of Ireland (officially Names of the Irish state, named Irelan ...
has no native dormouse, the hazel dormouse was discovered in
County Kildare County Kildare () is a Counties of Ireland, county in Ireland. It is in the Provinces of Ireland, province of Leinster and is part of the Eastern and Midland Region. It is named after the town of Kildare. Kildare County Council is the Local gove ...
in 2010, and appears to be spreading rapidly, helped by the prevalence of
hedgerows A hedge or hedgerow is a line of closely spaced (3 feet or closer) shrubs and sometimes trees, planted and trained to form a barrier or to mark the boundary of an area, such as between neighbouring properties. Hedges that are used to separate ...
in the Irish countryside. The United Kingdom distribution of the hazel dormouse can be found on the
National Biodiversity Network website
A 2020 study found that hazel dormice in Britain have declined by 51% since 2000. Woodland habitat loss and management and a warming climate are seen as material threats to their future status.


Habitat

According to
English Nature English Nature was the Executive agency, United Kingdom government agency that promoted the Conservation (ethic), conservation of wildlife, geology and wild places throughout England between 1990 and 2006. It was a non-departmental public body ...
's ''Dormouse Conservation Handbook'', hazel dormice are "particularly associated with deciduous woodland" but also inhabit hedgerows and scrub. In relation to its habitat, the hazel dormouse abundance was "greater on sites with more honeysuckle, yew, and birch" (Goodwin et al., 2018). Dormice seldom travel more than 70 m from their nest.Paul Bright, Pat Morris & Tony Mitchell-Jones,
The Dormouse Conservation Handbook
' (2nd ed.:
English Nature English Nature was the Executive agency, United Kingdom government agency that promoted the Conservation (ethic), conservation of wildlife, geology and wild places throughout England between 1990 and 2006. It was a non-departmental public body ...
, 2006), p. 13.


Description

The hazel dormouse can reach a body length of about and a length of about if you consider the tail as well. It is long with a tail of . It weighs , although this increases to just before hibernation. This small mammal has reddish brown fur that can vary up to golden-brown or yellow-orange-brown becoming lighter in the lower part. Eyes are large and black. Ears are small and not very developed, while the tail is long and completely covered with hair. It is a
nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...
creature and spends most of its waking hours among the branches of trees looking for food. It will make long detours rather than come down to the ground and expose itself to danger. The hazel dormouse hibernates from October to April–May.


Behaviour

Starting from the onset of colder weather (October or November), the hazel dormouse will hibernate in nests on the ground, at the base of old coppiced trees or hazel stools, under piles of leaves, or under log piles, as these situations are not subject to extreme variations in either temperature or humidity. Dormice are almost completely arboreal in habit but much less reluctant to cross open ground than was thought even recently. When it wakes up in spring (late April or early May), it builds woven nests of shredded
honeysuckle Honeysuckles are arching shrubs or Vine#Twining vines, twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely kno ...
bark, fresh leaves, and grasses in the undergrowth. If the weather is cold and wet and food is scarce, it saves energy by going into
torpor Torpor is a state of decreased physiological activity in an animal, usually marked by a reduced body temperature and metabolic rate. Torpor enables animals to survive periods of reduced food availability. The term "torpor" can refer to the ti ...
; it curls up into a ball and goes to sleep. The hazel dormouse, therefore, spends a large proportion of its life sleeping, either hibernating in winter or in torpor in summer. An examination of
hazelnut The hazelnut is the fruit of the hazel tree and therefore includes any of the nuts deriving from species of the genus '' Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or filberts according to ...
s may show a neat, round hole in the shell. This indicates it has been opened by a small
rodent Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal specie ...
, e.g. the dormouse,
wood mouse The wood mouse (''Apodemus sylvaticus'') is a Muridae, murid rodent native to Europe and northwestern Africa. It is closely related to the yellow-necked mouse (''Apodemus flavicollis'') but differs in that it has no band of yellow fur around the ...
, or
bank vole The bank vole (''Clethrionomys glareolus'') is a small vole with red-brown fur and some grey patches, with a tail about half as long as its body. A rodent, it lives in woodland areas and is around in length. The bank vole is found in much of Eu ...
. Other animals, such as
squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s or
jay Jays are a paraphyletic grouping of passerine birds within the family Corvidae. Although the term "jay" carries no taxonomic weight, most or all of the birds referred to as jays share a few similarities: they are small to medium-sized, usually ...
s, will either split the shell completely in half or make a jagged hole in it. Further examination reveals the cut surface of the hole has toothmarks which follow the direction of the shell. In addition, there will be toothmarks on the outer surface of the nut, at an angle of about 45 degrees to the cut surface. Woodmice and voles bite across the nutshell leaving clear parallel toothmarks from inside to outside. Woodmice also leave toothmarks on the outer surface of the nut but voles do not.


Diet

The hazel dormouse requires a variety of arboreal foods to survive. It eats berries and nuts and other fruit with
hazel Hazels are plants of the genus ''Corylus'' of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family, Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K ...
nuts being the main food for fattening up before hibernation. The dormouse also eats
hornbeam Hornbeams are hardwood trees in the plant genus ''Carpinus'' in the family Betulaceae. Its species occur across much of the temperateness, temperate regions of the Northern Hemisphere. Common names The common English name ''hornbeam'' derives ...
and
blackthorn ''Prunus spinosa'', called blackthorn or sloe, is an Old World species of flowering plant in the rose family, Rosaceae. It is locally naturalized in parts of the New World. The fruits are used to make sloe gin in Britain and patxaran in Basq ...
fruit where hazel is scarce. Other food sources are the buds of young leaves, and flowers which provide nectar and pollen. The dormouse also eats insects found on food-source trees, particularly
aphid Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...
s and
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s.


Plants of value to dormice

*
Hazel Hazels are plants of the genus ''Corylus'' of deciduous trees and large shrubs native to the temperate Northern Hemisphere. The genus is usually placed in the birch family, Betulaceae,Germplasmgobills Information Network''Corylus''Rushforth, K ...
is the principal food source, supports insects, forms an understory of poles, especially when
coppiced Coppicing is the traditional method in woodland management of cutting down a tree to a stump, which in many species encourages new shoots to grow from the stump or roots, thus ultimately regrowing the tree. A forest or grove that has been su ...
, which makes it useful for its
arboreal Arboreal locomotion is the locomotion of animals in trees. In habitats in which trees are present, animals have evolved to move in them. Some animals may scale trees only occasionally (scansorial), but others are exclusively arboreal. The hab ...
activity. The hazel dormouse's Latin name ''avellanarius'' means "hazel". *
Oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
s supply insect and flower food; the acorns are of little value. *
Honeysuckle Honeysuckles are arching shrubs or Vine#Twining vines, twining vines in the genus ''Lonicera'' () of the family Caprifoliaceae. The genus includes 158 species native to northern latitudes in North America, Eurasia, and North Africa. Widely kno ...
bark is their primary nesting material, and flowers and fruit are used for food. *
Bramble ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. I ...
flowers and fruits provide food over a long period. The thorns give protection for nests. Dormice thrive on blackberries. *
Alder buckthorn ''Frangula alnus'', commonly known as alder buckthorn, glossy buckthorn, or breaking buckthorn, is a tall deciduous shrub in the family Rhamnaceae. Unlike other "buckthorns", alder buckthorn does not have thorns. It is native to Europe, northern ...
– in parts of the dormouse range where hazel is scarce or absent, berries of alder buckthorn are the principal food source and vital for the accumulation of fat reserves in autumn prior to hibernation. *
Willow Willows, also called sallows and osiers, of the genus ''Salix'', comprise around 350 species (plus numerous hybrids) of typically deciduous trees and shrubs, found primarily on moist soils in cold and temperate regions. Most species are known ...
– unripe seeds in early spring. *
Birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' contains 3 ...
– seeds. * Hawthorn flowers are an important food in the spring. The fruit is eaten occasionally. *
Blackthorn ''Prunus spinosa'', called blackthorn or sloe, is an Old World species of flowering plant in the rose family, Rosaceae. It is locally naturalized in parts of the New World. The fruits are used to make sloe gin in Britain and patxaran in Basq ...
– fruits (blackthorn fruit is called "sloe"). * Ash – seed keys whilst they are still on the tree. *
Sycamore Sycamore is a name which has been applied to several types of trees, but with somewhat similar leaf forms. The name derives from the Ancient Greek () meaning . Species of otherwise unrelated trees known as sycamore: * ''Acer pseudoplatanus'', a ...
supplies insects and pollen, and a habitat. However, they cast a dense shade which decreases the understory. *
Hornbeam Hornbeams are hardwood trees in the plant genus ''Carpinus'' in the family Betulaceae. Its species occur across much of the temperateness, temperate regions of the Northern Hemisphere. Common names The common English name ''hornbeam'' derives ...
– seeds. *Wayfaring tree (''
Viburnum lantana ''Viburnum lantana'', the wayfarer or wayfaring tree, is a species of ''Viburnum'', native to central, southern and western Europe (north to Yorkshire in England), northwest Africa, and southwestern Asia.Blamey, M. & Grey-Wilson, C. (1989). ''Fl ...
'') – fruits and flowers. *
Broom A broom (also known as a broomstick) is a cleaning tool, consisting of usually stiff fibers (often made of materials such as plastic, hair, or corn husks) attached to, and roughly parallel to, a cylindrical handle, the broomstick. It is thus a ...
– flowers (in early summer). *
Yew Yew is a common name given to various species of trees. It is most prominently given to any of various coniferous trees and shrubs in the genus '' Taxus'': * European yew or common yew (''Taxus baccata'') * Pacific yew or western yew ('' Taxus ...
– fruits are a favoured food. *
Sweet chestnut The sweet chestnut (''Castanea sativa''), also known as the Spanish chestnut or European chestnut, is a species of tree in the family Fagaceae, native to Southern Europe and Asia Minor, and widely cultivated throughout the temperate world. A ...
provides an excellent foodsource, and the flowers are eaten, as well.


Threats

* Predation from
fox Foxes are small-to-medium-sized omnivorous mammals belonging to several genera of the family Canidae. They have a flattened skull; upright, triangular ears; a pointed, slightly upturned snout; and a long, bushy tail ("brush"). Twelve species ...
es,
wild boar The wild boar (''Sus scrofa''), also known as the wild swine, common wild pig, Eurasian wild pig, or simply wild pig, is a Suidae, suid native to much of Eurasia and North Africa, and has been introduced to the Americas and Oceania. The speci ...
s,
weasel Weasels are mammals of the genus ''Mustela'' of the family Mustelidae. The genus ''Mustela'' includes the least weasels, polecats, stoats, ferrets, and European mink. Members of this genus are small, active predators, with long and slend ...
s,
owl Owls are birds from the order Strigiformes (), which includes over 200 species of mostly solitary and nocturnal birds of prey typified by an upright stance, a large, broad head, binocular vision, binaural hearing, sharp talons, and feathers a ...
s and
domestic cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small Domestication, domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have sh ...
s.Verbeylen, Goedele
"How small should the entrance be? Is it possible to let common dormice Muscardinus avellanarius enter nest boxes and exclude other species.
(2017): 35-39.
* Being dug up or disturbed during hibernation by
badger Badgers are medium-sized short-legged omnivores in the superfamily Musteloidea. Badgers are a polyphyletic rather than a natural taxonomic grouping, being united by their squat bodies and adaptions for fossorial activity rather than by the ...
s. * Lack of food source, e.g., from too frequent hedge-trimming, or competition from other species, e.g.,
squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s, other dormouse species, mice. * Destruction of
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
and hedgerow habitats, or their diverse range of species, as a broad spectrum of food is required across the calendar year. * Reduction in traditional forest management. * A warming climate.


Protection status

The hazel dormouse is protected by and in UK under the
Wildlife and Countryside Act The Wildlife and Countryside Act 1981 (c. 69) is an act of Parliament (UK), act of Parliament in the United Kingdom implemented to comply with European Council Birds Directive, Directive 79/409/EEC on the conservation of wild birds. In short, the ...
. At European level, the hazel dormouse is classed as In Need of Strict Protection under annex IV of the
Habitats Directive The Habitats Directive (more formally known as Council Directive 92/43/EEC on the Conservation of natural habitats and of wild fauna and flora) is a directive adopted by the European Community in 1992 as a response to the Berne Convention. The ...
92/43/EEC "Gliridae - All species except Glis glis and Eliomys quercinus"


Evolutionary history

The oldest fossils of the genus ''Muscardinus'' date to the
Serravallian The Serravallian is, in the geologic timescale, an List of time periods, age or a stage (stratigraphy), stage in the middle Miocene epoch (geology), Epoch/series (stratigraphy), Series, which spans the time between 13.82 annum, Ma and 11.63 Ma (m ...
stage of the Middle
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
approximately 13.8 to 11.6 million years ago in what is now Spain. The oldest fossils of the modern species date to the
Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...
.


References


External links


The Mammal Society
site with a Hazel dormouse fact sheet. There is also a book entitled The Dormouse available, by Pat Morris.
Peoples Trust for Endangered Species
site describing the hazel dormouse and its conservation
Information and images
from the
BBC The British Broadcasting Corporation (BBC) is a British public service broadcaster headquartered at Broadcasting House in London, England. Originally established in 1922 as the British Broadcasting Company, it evolved into its current sta ...

Extensive information and picturesA lot of facts, links and book reviews about the dormouseDormouse nest-box construction
{{Authority control Dormice Mammals of Asia Rodents of Europe Mammals of Russia Mammals of Turkey Mammals described in 1758 Taxa named by Carl Linnaeus