Zone-tailed Hawk
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The zone-tailed hawk (''Buteo albonotatus'') is a medium-sized
hawk Hawks are birds of prey of the family Accipitridae. They are very widely distributed and are found on all continents, except Antarctica. The subfamily Accipitrinae includes goshawks, sparrowhawks, sharp-shinned hawks, and others. This ...
of warm, dry parts of the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
. It is somewhat similar in plumage and flight style to a common scavenger, the
turkey vulture The turkey vulture (''Cathartes aura'') is the most widespread of the New World vultures. One of three species in the genus '' Cathartes'' of the family Cathartidae, the turkey vulture ranges from southern Canada to the southernmost tip of Sou ...
, and may benefit from being able to blend into groups of vultures. It feeds on small terrestrial
tetrapod A tetrapod (; from Ancient Greek :wiktionary:τετρα-#Ancient Greek, τετρα- ''(tetra-)'' 'four' and :wiktionary:πούς#Ancient Greek, πούς ''(poús)'' 'foot') is any four-Limb (anatomy), limbed vertebrate animal of the clade Tetr ...
s of all kinds.


Taxonomy

In 1844, the English zoologist
George Robert Gray George Robert Gray (8 July 1808 – 6 May 1872) was an English zoology, zoologist and author, and head of the Ornithology, ornithological section of the British Museum, now the Natural History Museum, London, Natural History Museum, London f ...
, in his ''List of the Specimens of Birds in the Collection of the British Museum'', mentioned the zone-tailed hawk under the common name "white spotted buzzard" and coined the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Buteo albonotatus''. As Gray omitted a
species description A species description is a formal scientific description of a newly encountered species, typically articulated through a scientific publication. Its purpose is to provide a clear description of a new species of organism and explain how it diff ...
, his binomial name is considered ''nomen nudum'', a naked name, and is not recognised. Instead, the German naturalist
Johann Jakob Kaup Johann Jakob von Kaup (10 April 1803 – 4 July 1873) was a German naturalist. A proponent of natural philosophy, he believed in an innate mathematical order in nature and he attempted biological classifications based on the Quinarian system. Kaup ...
is recognised as the
authority Authority is commonly understood as the legitimate power of a person or group of other people. In a civil state, ''authority'' may be practiced by legislative, executive, and judicial branches of government,''The New Fontana Dictionary of M ...
as in 1847, he provided a brief description and used Gray's name ''Buteo albonotatus''. The type locality is Mexico. The specific epithet ''albonotatus'' combines the Latin ''albus'' meaning "white" with ''notatus'' meaning "marked". The species is
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
: no
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
are recognised.


Description

The zone-tailed hawk is a fairly large but slender ''
Buteo ''Buteo'' is a genus of medium to fairly large, wide-ranging raptors with a robust body and broad wings. In the Old World, members of this genus are called " buzzards", but "hawk" is used in the New World (Etymology: ''Buteo'' is the Latin name ...
'' hawk. Grown birds are in length with a wingspan of about . The zone-tailed is comparable in length and wingspan to common large ''Buteos'' found to the north such as Swainson's and
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members of ...
, but may weigh considerably less. Their body mass can range from . In measurements, the sexes are close in size, but the female, at an average of , is much heavier and bulkier than the male, at an average of . Among standard measurements, the wing chord is , the fairly long tail is and the tarsus is . The adult
plumage Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
is mostly blackish. The notable exception is that the flight
feather Feathers are epidermal growths that form a distinctive outer covering, or plumage, on both avian (bird) and some non-avian dinosaurs and other archosaurs. They are the most complex integumentary structures found in vertebrates and an exa ...
s are barred with lighter gray, which can appear solid silver-gray from a distance. The tail has three or four bands (the "zones" of the common name), white from below and light gray from above, of which the one second from the tip is particularly broad and conspicuous. The
cere The beak, bill, or Rostrum (anatomy), rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for pecking, wikt:grasp#Verb, grasping, and holding (in wikt:probe ...
and legs are yellow, the lores are light gray and a light touch of white may be seen on the face. Immatures are similar except for small white spots on the breast and tails with narrow gray and black bands and a broad dark tip. The zone-tailed hawk adults resemble the common black hawk but are distinctly more slender in flight and overall small, and they have more white bars on the tail. Other ''Buteo'' hawks in their dark phase, especially the broad-winged hawk, may appear similar but often have more silvery coloration on the wings and are broader-winged. The call is a loud scream, a somewhat typical ''Buteo'' call, dropping in pitch at the end, ''kra kree-kree-kree-kree''. In at least some birds, there is an abrupt rise in pitch (like a break to a
falsetto Falsetto ( , ; Italian language, Italian diminutive of , "false") is the vocal register occupying the frequency range just above the modal voice register and overlapping with it by approximately one octave. It is produced by the vibration of the ...
voice) in the middle and an equally abrupt drop back down. They are most often heard vocalizing when engaging in breeding displays at the beginning of the mating season. When disturbed at the nest, they may utter a long, lower-pitched ''raaaaauu''.


Distribution and habitat

Zone-tailed hawks range from parts of southern
Arizona Arizona is a U.S. state, state in the Southwestern United States, Southwestern region of the United States, sharing the Four Corners region of the western United States with Colorado, New Mexico, and Utah. It also borders Nevada to the nort ...
,
New Mexico New Mexico is a state in the Southwestern United States, Southwestern region of the United States. It is one of the Mountain States of the southern Rocky Mountains, sharing the Four Corners region with Utah, Colorado, and Arizona. It also ...
, and western
Texas Texas ( , ; or ) is the most populous U.S. state, state in the South Central United States, South Central region of the United States. It borders Louisiana to the east, Arkansas to the northeast, Oklahoma to the north, New Mexico to the we ...
almost throughout inland
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
and the central portions of
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
down into eastern
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
,
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
, and more sporadically, into
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
, southern
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
,
Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...
,
Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...
, and northern
Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...
. In winter, they generally withdraw from the
U.S. The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 states and a federal capital district, Washington, D.C. The 48 contiguous ...
and northernmost Mexico, with these populations wintering mostly in
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
and the
Yucatán Peninsula The Yucatán Peninsula ( , ; ) is a large peninsula in southeast Mexico and adjacent portions of Belize and Guatemala. The peninsula extends towards the northeast, separating the Gulf of Mexico to the north and west of the peninsula from the C ...
. They are also native to the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
island of
Trinidad Trinidad is the larger, more populous island of the Republic of Trinidad and Tobago, the country. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is the southernmost island in ...
. The hawks of Central America may be seasonally migratory, although their movements are not well known. Zone-tails sometimes wander out of their normal range, and the bird was once recorded in
Nova Scotia Nova Scotia is a Provinces and territories of Canada, province of Canada, located on its east coast. It is one of the three Maritime Canada, Maritime provinces and Population of Canada by province and territory, most populous province in Atlan ...
. They can adapt to various habitats across their broad range, including both closed and open ones and wet and dry ones. Often, the largest numbers are found in rocky areas with access to water. They often reside in
coniferous Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
or pine-oak forests as well as timbered canyonland, hilly riverine woods, dry open boscage and scrub, humid forests, and overgrown marshes. They may forage over ranches and even semi-desert, but always need at least scattered tree thickets for nesting. Furthermore, they may be distributed in elevation from sea level to , though are mainly found below in the north and in the southern reaches of the breeding range.


Behaviour and ecology

The bird's flight feathers closely resemble those of the
turkey vulture The turkey vulture (''Cathartes aura'') is the most widespread of the New World vultures. One of three species in the genus '' Cathartes'' of the family Cathartidae, the turkey vulture ranges from southern Canada to the southernmost tip of Sou ...
. Zone-tailed hawks soar with their wings held in a dihedral position (pointing slightly upwards), rocking from side to side, a flight style that parallels that of turkey vultures. Bird guides caution against confusing them with the much more common turkey vulture, but at a reasonable distance, one can distinguish them from vultures by their smaller size, the typical hawk shape of the wings and head, and the pale stripe on the tail. Since vultures frequently can be seen flying in numbers (groups are called "kettles"), zone-tailed hawks can mingle with them and are perhaps most often missed by the human eye in such kettles.


Breeding

The mating season of the species varies geographically but is almost always in the first half of the year. In the northern reaches of the range, the breeding season is mid-April through July, whereas in
Trinidad Trinidad is the larger, more populous island of the Republic of Trinidad and Tobago, the country. The island lies off the northeastern coast of Venezuela and sits on the continental shelf of South America. It is the southernmost island in ...
and
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
, it is February through June. Eggs have been found as late as August in
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
, implying only a loose breeding season in the true tropics. The mating pair performs a courtship display, which may include engaging in aerial loops, dives, and rolls with each other. The nest is typical of hawks: a big, bulky assemblage of sticks, lined with green leaves, usually built on the top or the main fork of a tree, in this case at above the ground. Typically, tall trees such as cottonwood or
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...
trees are selected, and the nest may be in the open or concealed by foliage. Occasionally, nests are found on cliffs. The clutch comprises one to three, typically two, white eggs, often marked with brown. Incubation lasts for around 28 to 35 days and typically the female incubates, while being fed by the male, although the male may occasionally incubate. The young are semi-
altricial Precocial species in birds and mammals are those in which the young are relatively mature and mobile from the moment of birth or hatching. They are normally nidifugous, meaning that they leave the nest shortly after birth or hatching. Altricial ...
at hatching and are covered in gray down. They grow slowly for the first 7 days of life and then considerably faster from 7 to 21 days old. As is common in raptors the older sibling often kills the younger one or outcompetes it for food; only occasionally do both survive to adulthood. The younger hatchling is sometimes referred to as the "spare" one since it may be tended to more directly if the first dies. The young fledge at 42 to 50 days, though are not typically self-assured fliers until around a week later. They may remain in their parents' care until the following breeding season, though in migratory populations, the young and adults often separate. There have been no extensive reports on longevity and mortality in the species.


Food and feeding

Unlike turkey vultures that do not normally prey on live animals, zone-tailed hawks are active predators. Therefore, some ornithologists believe that this
mimicry In evolutionary biology, mimicry is an evolved resemblance between an organism and another object, often an organism of another species. Mimicry may evolve between different species, or between individuals of the same species. In the simples ...
tricks potential prey animals into not being alarmed when a zone-tail flies overhead. This hawk mainly preys on small birds and mammals (including
bat Bats are flying mammals of the order Chiroptera (). With their forelimbs adapted as wings, they are the only mammals capable of true and sustained flight. Bats are more agile in flight than most birds, flying with their very long spread-out ...
s,
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
s,
mice A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus' ...
,
squirrel Squirrels are members of the family Sciuridae (), a family that includes small or medium-sized rodents. The squirrel family includes tree squirrels, ground squirrels (including chipmunks and prairie dogs, among others), and flying squirrel ...
s),Buteo albonotatus (Zone-tailed Hawk)
/ref> but reptiles can be locally favored, including virtually any type of
lizard Lizard is the common name used for all Squamata, squamate reptiles other than snakes (and to a lesser extent amphisbaenians), encompassing over 7,000 species, ranging across all continents except Antarctica, as well as most Island#Oceanic isla ...
, such as the
common collared lizard The common collared lizard (''Crotaphytus collaris''), also Common name, commonly called eastern collared lizard,Robert C. Stebbins, Stebbins RC (2003). ''A Field Guide to Western Reptiles and Amphibians, Third Edition''. The Peterson Field Guid ...
, Yarrow's spiny lizard, the crevice spiny lizard, and the mesquite lizard. Among the rats preyed upon are the tawny-bellied cotton rat and the white-eared cotton rat. In the north, California quail, along with possibly other quail species,
chipmunk Chipmunks are small, striped rodents of subtribe Tamiina. Chipmunks are found in North America, with the exception of the Siberian chipmunk which is found primarily in Asia. Taxonomy and systematics Chipmunks are classified as four genera: '' ...
s, and other
ground squirrel Ground squirrels are rodents of the squirrel family (Sciuridae) that generally live on the ground or in burrows, rather than in trees like the tree squirrels. The term is most often used for the medium-sized ground squirrels, as the larger ones ar ...
s seemed to be the favorite prey. Other birds preyed upon include red-shafted flickers, acorn woodpeckers, eared quetzals,
Steller's jay Steller's jay (''Cyanocitta stelleri'') is a bird native to western North America and the mountains of Central America, closely related to the blue jay (''C. cristata'') found in eastern North America. It is the only crest (feathers), crested jay ...
s, yellow-rumped caciques, tropical kingbirds, and even
eastern screech owl The eastern screech owl (''Megascops asio'') or eastern screech-owl, is a small typical owl, owl that is relatively common in Eastern North America, from Mexico to Canada. This species resides in most types of woodland habitats across its range, ...
s. Zone-tails also eat various young birds, having been observed preying on nestlings and fledglings of species as varied as
heron Herons are long-legged, long-necked, freshwater and coastal birds in the family Ardeidae, with 75 recognised species, some of which are referred to as egrets or bitterns rather than herons. Members of the genus ''Botaurus'' are referred to as bi ...
s and
passerine A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
s. Zone-tailed hawks snatch young birds from trees or the ground without landing. Second-hand reports of predation on
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
s, other
amphibian Amphibians are ectothermic, anamniote, anamniotic, tetrapod, four-limbed vertebrate animals that constitute the class (biology), class Amphibia. In its broadest sense, it is a paraphyletic group encompassing all Tetrapod, tetrapods, but excl ...
s, and
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
es may be cases of misidentification of common black-hawks. Zone-tailed hawks are very active foragers, hunting almost exclusively by transects and random quartering in low flight at around over the ground. When approaching the prey, the hawk may try to use obstructing cover such as trees until it is within of the prey, an easy striking distance. Outside the breeding pair bond, these hawks are wholly solitary and are not known to hunt in pairs.


References


Further reading

* * * Also available fro
Researchgate
*


External links


Xeno-canto: audio recordings of the Zone-tailed Hawk
{{Taxonbar, from=Q595114 Buteo Birds of prey Birds of Central America Birds of Mexico Birds of South America Birds of Trinidad and Tobago Birds of the Caribbean Native birds of the Southwestern United States Hawks Birds described in 1847 Taxa named by Johann Jakob Kaup