''Zanthoxylum'' is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of about 250 species of

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

and

evergreen In botany, an evergreen is a plant which has Leaf, foliage that remains green and functional throughout the year. This contrasts with deciduous plants, which lose their foliage completely during the winter or dry season. Consisting of many diffe ...

trees, shrubs and climbers in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rueRUTACEAE
in Bo ...

that are native to warm

temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...

and

subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...

areas worldwide. It is the type genus of the tribe Zanthoxyleae in the subfamily

Rutoideae Rutoideae is a flowering plant subfamily in the family Rutaceae. The subfamily has had varying circumscriptions. In a 2012 classification of the family it was one of only two subfamilies and contained most of the genera, whereas in a 2021 classif ...

. Several of the species have yellow

heartwood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...

, to which their generic name alludes. Several species are cultivated for their use as spices, notably including

Sichuan pepper Sichuan pepper (, also known as Sichuanese pepper, Szechuan pepper, Chinese prickly ash, Chinese pepper, Mountain pepper, and ''mala'' pepper, is a spice commonly used in Sichuan cuisine in China, Bhutan and in northeast India. It is called mej ...

Description

Plants in the genus ''Zanthoxylum'' are typically

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

shrubs, trees or woody climbers armed with

trichomes Trichomes (; ) are fine outgrowths or appendages on plants, algae, lichens, and certain protists. They are of diverse structure and function. Examples are hairs, glandular hairs, scales, and papillae. A covering of any kind of hair on a plant ...

. The leaves are arranged alternately and are usually

pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...

trifoliate The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (that is, the leaf blade or 'lamina' is undivided) or compound (that is, the leaf blade is divided into two or more leaflets ...

. The flowers are usually arranged in

panicle In botany, a panicle is a much-branched inflorescence. (softcover ). Some authors distinguish it from a compound spike inflorescence, by requiring that the flowers (and fruit) be pedicellate (having a single stem per flower). The branches of a p ...

s and usually function as male or female flowers with four

sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...

s and four petals, the sepals remaining attached to the fruit. Male flowers have four

stamen The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...

s opposite the sepals. Female flowers have up to five, more or less free

carpels Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) ...

with the styles free or sometimes fused near the tip. The fruit is usually of up to four follicles fused at the base, each containing a single seed almost as large as the follicle.

Taxonomy

The genus ''Zanthoxylum'' was first formally described in 1753 by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in the first volume of ''

Species Plantarum ' (Latin for "The Species of Plants") is a book by Carl Linnaeus, originally published in 1753, which lists every species of plant known at the time, classified into genus, genera. It is the first work to consistently apply binomial nomenclature ...

''. The generic name is derived from

Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...

words (), meaning 'yellow', and (), meaning 'wood'. It is technically misspelled, as the ''z'' should be ''x'', but botanical nomenclature does not allow for spelling corrections. It refers to a yellow dye made from the roots of some species. The first species that Linnaeus described was ''Zanthoxylum trifoliatum'', now regarded as a

synonym A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...

of ''

Eleutherococcus trifoliatus ''Eleutherococcus'' is a genus of 38 species,p.40, citing Frodin, Govaerts 2003 of thorny shrubs and trees in the family Araliaceae. They are native to eastern Asia, from southeast Siberia and Japan to the Philippines and Vietnam. 18 species come ...

''. The once separate genus ''Fagara'' is now included in ''Zanthoxylum''.

Fossil record

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

seeds of the extinct ''Zanthoxylum kristinae'', from the early

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

, have been found in the Kristina Mine at

Hrádek nad Nisou Hrádek nad Nisou (; , ) is a town in Liberec District in the Liberec Region of the Czech Republic. It has about 8,000 inhabitants. The historic town centre is well preserved and is protected as an urban monument zone. Administrative division Hr� ...

North Bohemia North Bohemia (, ) is a region in the north of the Czech Republic. Location North Bohemia roughly covers the present-day NUTS regional unit of ''CZ04 Severozápad'' and the western part of ''CZ05 Severovýchod''. From an administrative perspec ...

, the Czech Republic.

Uses

Many ''Zanthoxylum'' species make excellent

bonsai Bonsai (; , ) is the Japanese art of Horticulture, growing and shaping miniature trees in containers, with a long documented history of influences and native Japanese development over a thousand years, and with unique aesthetics, cultural hist ...

and in temperate climates they can be grown quite well indoors. ''

Zanthoxylum beecheyanum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' and ''

Zanthoxylum piperitum ''Zanthoxylum piperitum'', also known as Japanese pepper or Japanese prickly-ash, is a deciduous aromatic spiny shrub or small tree of the citrus and rue family Rutaceae, native to Japan and Korea. It is called () in Japan and () in Korea. Bo ...

'' are two species commonly grown as

.Wilbur, C. Keith, MD. ''Revolutionary Medicine 1700-1800''. The Globe Pequot Press. Page 23. 1980.

Culinary use

Spice In the culinary arts, a spice is any seed, fruit, root, Bark (botany), bark, or other plant substance in a form primarily used for flavoring or coloring food. Spices are distinguished from herbs, which are the leaves, flowers, or stems of pl ...

s are made from a number of species in this genus, including: * ''

Zanthoxylum acanthopodium ''Zanthoxylum acanthopodium'', or andaliman, is a species of flowering plant in the family Rutaceae. Its range includes southern western China,Bangladesh, Bhutan, northern India and northeastern India, Nepal, Laos, Burma, northern Thailand, Vie ...

'' –

Andaliman ''Zanthoxylum acanthopodium'', or andaliman, is a species of flowering plant in the family Rutaceae. Its range includes southern western China,Bangladesh, Bhutan, northern India and northeastern India, Nepal, Laos, Burma, northern Thailand, Vie ...

* '' Zanthoxylum armatum'' – "green" or "vine"

* ''

Zanthoxylum bungeanum ''Zanthoxylum bungeanum '' is a species of plant in the family Rutaceae. It is one of the sources of the spice Sichuan pepper. The plant is native to North-Central China, South-Central China, Southeast China, East Himalayas The Himalayas, ...

'' – "red" or common Sichuan pepper * ''

Zanthoxylum gilletii ''Zanthoxylum gilletii'', the East African satinwood, is a tree species in the genus ''Zanthoxylum'' found in Africa. The fruits are used to produce the spice uzazi, which is the Nigerian (Igbo language) name of both the plant and the spice. It ...

'' - Uzazi * ''

'' – , * ''

Zanthoxylum nitidum ''Zanthoxylum nitidum'', commonly known as shiny-leaf prickly-ash, is a species of flowering plant in the family Rutaceae. It is a woody climber with prickles on the branchlets, thick, cone-shaped spines on the trunk and older branches, pinnate ...

'' * ''

Zanthoxylum rhetsa ''Zanthoxylum rhetsa'', commonly known as Indian prickly ash, is a species of flowering plant in the family Rutaceae and occurs from India east to the Philippines and south to northern Australia. It is a deciduous shrub or tree with cone-shaped s ...

'' – teppal, tirphal * ''

Zanthoxylum schinifolium ''Zanthoxylum schinifolium'', also called mastic-leaf prickly ash, is a species of flowering plant in the Rutaceae, the citrus family. It was first described and published in Abh. Math.-Phys. Cl. Königl. Bayer. Akad. Wiss. vol.4 (Issue 2) on pa ...

'' – , "green" Sichuan pepper * ''

Zanthoxylum simulans ''Zanthoxylum simulans'', the Chinese-pepper, Chinese prickly-ash or flatspine prickly-ash, is a flowering plant in the family Rutaceae, native to southern and central China, Taiwan, and South Korea. It is one of several species of ''Zanthoxylum' ...

'' – "red" or wild Sichuan pepper

Andaliman

Indonesia Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...

North Sumatra North Sumatra () is a Provinces of Indonesia, province of Indonesia located in the northern part of the island of Sumatra. Its capital and largest city is Medan on the east coast of the island. It borders Aceh to the northwest, Riau to the sou ...

province, ''

'' is harvested for ''andaliman''. In Indonesian Batak cuisine, ''

andaliman ''Zanthoxylum acanthopodium'', or andaliman, is a species of flowering plant in the family Rutaceae. Its range includes southern western China,Bangladesh, Bhutan, northern India and northeastern India, Nepal, Laos, Burma, northern Thailand, Vie ...

'' is ground and mixed with chilies and seasonings into a green

sambal Sambal is an Indonesian chili sauce or paste, typically made from a mixture of chillis with secondary ingredients such as shrimp paste (terasi), garlic, ginger, shallot, scallion, palm sugar, and lime juice. ''Sambal'' is an Indonesia ...

or chili paste.

Arsik Arsik is an Indonesian spicy fish dish of the Batak Toba and Mandailing people of North Sumatra, usually using the common carp (known in Indonesia as ''ikan mas'' or gold fish). Distinctively Batak elements of the dish are the use of torch gi ...

is a typical Indonesian dish containing ''andaliman''.

and

'' is harvested in Japan and Korea to produce () or (), which has numbing properties similar to those of Chinese Sichuan peppercorns. In Korean cuisine, is often used to accompany fish soups such as , whereas the plant's seeds are separated and used to make oil, and the oil is used as a medicine.

Sancho

The Korean is made from ''

'', which is slightly less bitter than . In

Korean cuisine Korean cuisine is the set of foods and culinary styles which are associated with Korean culture. This cuisine has evolved through centuries of social and political change. Originating from ancient Prehistoric Korea, agricultural and nomad ...

, is often used to accompany fish soups such as .

Sichuan pepper

The

fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...

of '' Zanthoxylum armatum'' and ''

'' species is used to make

by grinding the husks that surround the berries.

Triphal and teppal

In the states of

Maharashtra Maharashtra () is a state in the western peninsular region of India occupying a substantial portion of the Deccan Plateau. It is bordered by the Arabian Sea to the west, the Indian states of Karnataka and Goa to the south, Telangana to th ...

Karnataka Karnataka ( ) is a States and union territories of India, state in the southwestern region of India. It was Unification of Karnataka, formed as Mysore State on 1 November 1956, with the passage of the States Reorganisation Act, 1956, States Re ...

, and

Goa Goa (; ; ) is a state on the southwestern coast of India within the Konkan region, geographically separated from the Deccan highlands by the Western Ghats. It is bound by the Indian states of Maharashtra to the north, and Karnataka to the ...

in Western India, the dried berries of ''

'' are known as teppal or tirphal in

Marathi Marathi may refer to: *Marathi people, an Indo-Aryan ethnolinguistic group of Maharashtra, India **Marathi people (Uttar Pradesh), the Marathi people in the Indian state of Uttar Pradesh *Marathi language, the Indo-Aryan language spoken by the Mar ...

are added to foods such as legumes and fish. The name in both languages means 'three fruits' or 'three pods'. Because the trees bear fruit during the

monsoon A monsoon () is traditionally a seasonal reversing wind accompanied by corresponding changes in precipitation but is now used to describe seasonal changes in Atmosphere of Earth, atmospheric circulation and precipitation associated with annu ...

season, the berries are associated with the concurrent

Krishna Janmashtami Krishna Janmashtami (), also known simply as Krishnashtami, Janmashtami, or Gokulashtami, is an annual Hindu festival that celebrates the birth of Krishna, the eighth avatar of Vishnu. In certain Hindu texts, such as the '' Gita Govinda ...

festival. The fresh fruits are parrot green in color and are used as a flavouring agent in many curries made with a paste of coconut, chilis, and other spices. When dried, the flesh of the fruit hardens, turns a brownish black color and opens up to show the black seeds within. The seeds are discarded and the dried fruit is stored in containers for use around the year. Mostly used in fish preparations and a few vegetarian dishes, with coconut masala, this spice has a very strong woody aroma and is discarded at the time of eating the curry.

Chemistry

Plants in the genus ''Zanthoxylum'' contain the lignan

sesamin Sesamin is a lignan isolated from the bark of '' Fagara'' plants and from sesame oil Sesame oil is an edible vegetable oil derived from sesame seeds. The oil is one of the earliest-known crop-based oils. Worldwide mass modern production is ...

. Species identified in Nigeria contains several types of alkaloids including benzophenanthridines (

nitidine Nitidine is a benzophenanthridine alkaloid found in species of the genus '' Zanthoxylum '', notably in '' Zanthoxylum nitidum''. This compound has an anti-malarial activity.Bouquet, J., et al. (2012)Biological activities of nitidine, a potential a ...

, dihydronitidine, oxynitidine,

fagaronine Fagaronine is a benzophenanthridine alkaloid found in ''Zanthoxylum zanthoxyloides'' and other species in the genus ''Zanthoxylum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the ...

, dihydro

avicine Avicine, tested and developed by AVI BioPharma, and also known as CTP-37 was trialled as a possible cancer vaccine to treat a number of different cancers. These included colorectal cancer, pancreatic cancer and prostate cancer. The treatment was t ...

chelerythrine Chelerythrine is a benzophenanthridine alkaloid present in the plant '' Chelidonium majus'' (greater celandine). It is a potent, selective, and cell-permeable protein kinase C inhibitor ''in vitro''. And an efficacious antagonist of G-protein-co ...

, dihydrochelerythrine, methoxychelerythrine, norchelerythrine, oxychelerythrine, decarine and fagaridine),

furoquinoline Furoquinoline alkaloids are a group of alkaloids with simple structure. Distribution of this group of alkaloids is essentially limited to plant family Rutaceae. The simplest member of this group is dictamnine and most widespread member is skimmia ...

s ( dictamine, 8-methoxydictamine,

skimmianine Skimmianine is a furoquinoline alkaloid found in '' Skimmia japonica'', a flowering plant in family Rutaceae that is native to Japan and China. It is a known acetylcholinesterase inhibitor. Biosynthesis The biosynthesis of skimmianine starts f ...

, 3-dimethylallyl-4-methoxy-2-quinolone),

carbazole Carbazole is an aromatic Heterocyclic compound, heterocyclic organic compound. It has a tricyclic structure, consisting of two six-membered benzene rings fused on either side of a five-membered nitrogen-containing ring. The compound's structure is ...

s (3-methoxycarbazole, glycozoline),

aporphine Aporphine is an alkaloid with the chemical formula . It is the core chemical substructure of the aporphine alkaloids, a subclass of quinoline alkaloids. It can exist in either of two enantiomeric forms, (''R'')-aporphine and (''S'')-aporphine. ...

s (

berberine Berberine is a Quaternary ammonium cation, quaternary ammonium salt from the protoberberine group of benzylisoquinoline alkaloids, occurring naturally as a secondary metabolite in some plants including species of ''Berberis'', from which its name ...

, tembetarine,

magnoflorine ''(S)-''Magnoflorine is a quaternary benzylisoquinoline alkaloid (BIA) of the aporphine structural subgroup which has been isolated from various species of the family Menispermaceae, such as ''Pachygone ovata,'' '' Sinomenium acutum,'' and '' Ci ...

, M-methyl- corydine), canthinones (6-canthinone),

acridone Acridone is an organic compound based on the acridine skeleton, with a carbonyl group at the ''9'' position. Synthesis and structure The molecule is planar. Optical spectra reveal that the keto tautomer predominates in the gas-phase and in ethanol ...

s (1-hydroxy-3-methoxy-10-methylacridon-9-one, 1-hydroxy-10-methylacridon-9-one, zanthozolin), and aromatic and

aliphatic amide Fatty acid amides (FAAs) are amides formed from a fatty acid and an amine. In nature, many FAAs have ethanolamine as the amine component. Also known as ''N''-acylethanolamines, they contain the functionality RC(O)N(H)CH2CH2OH. A well known examp ...

s.The Nigerian Zanthoxylum; Chemical and biological values. S. K. Adesina, Afr. J. Trad. CAM, 2005, volume 2, issue 3, pages 282-301
article
) Hydroxy-alpha sanshool is a bioactive component of plants from the genus ''Zanthoxylum'', including the Sichuan pepper. It provides the characteristic numbness.

Ecology

''Zanthoxylum'' species are used as food plants by the

larvae A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect developmental biology, development such as insects, some arachnids, amphibians, or cnidarians typical ...

of some

Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...

species including the engrailed (''Ectropis crepuscularia'').

Species

The following is a list of species accepted by

Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...

as of August 2020: * ''

'' D.R.Simpson (southern Asia) * ''

Zanthoxylum aculeatissimum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

Engl. Engl or Engl. may refer to: *England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more ...

(Bolivia) * '' Zanthoxylum acuminatum'' Sw. (Cuba, Jamaica) * ''

Zanthoxylum ailanthoides ''Zanthoxylum ailanthoides'', also called ailanthus-like prickly ash, (, lit. "Ailanthus-leaved pepper",; the name is obviously adaptation of latin ailanthoides "ailanthus-like" , lit. " Yue pepper",, p.503 食茱萸 ''shi zhu yu'',, 「澳名に ...

'' Siebold & Zucc. – ailanthus-like prickly ash * ''

Zanthoxylum albiflorum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

Baker f. Edmund Gilbert Baker (1864–1949) was a British plant collector and botanist. He was the son of John Gilbert Baker John Gilbert Baker (13 January 1834 – 16 August 1920) was an England, English botanist. His son was the botanist Edmund G ...

(New Caledonia) * '' Zanthoxylum albuquerquei'' D.C. (Peru) * ''

Zanthoxylum amamiense ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

Ohwi was a Japanese Botany, botanist. He was a distinguished member of the Faculty of Science of Kyoto University, Kyoto Imperial University. He is perhaps best known for his 1953 . Species named after Ohwi * (Cyperaceae) ''Carex ohwii'' Genkei Masa ...

(

Ryukyu Islands The , also known as the or the , are a chain of Japanese islands that stretch southwest from Kyushu to Geography of Taiwan, Taiwan: the Ryukyu Islands are divided into the Satsunan Islands (Ōsumi Islands, Ōsumi, Tokara Islands, Tokara and A ...

) * ''

Zanthoxylum amapaense ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' (Albuq.) P.G.Waterm. (Brazil, Venezuela) * ''

Zanthoxylum americanum ''Zanthoxylum americanum'', the common prickly-ash, common pricklyash, common prickly ash or northern prickly-ash (also sometimes called toothache tree, yellow wood, or suterberry), is an aromatic shrub or small tree native to central and easter ...

'' Mill. – northern prickly ash, prickly ash, toothache tree (Canada, United States) * ''

Zanthoxylum anadenium ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' (

Urb. Ignatz Urban (7 January 1848 – 7 January 1931) was a German botany, botanist. He is known for his contributions to the flora of the Caribbean and Brazil, and for his work as curator of the Botanical Garden in Berlin, Berlin Botanical Garden. B ...

Ekman Ekman is a Swedish surname. Notable people with the surname include: *Carl Gustaf Ekman (1872–1945), Swedish politician *Carl Daniel Ekman (1845–1904), Swedish chemical engineer *Elisabeth Ekman (1862–1936), Swedish botanist *Erik Leonard Ekm ...

) J.Jiménez Alm. (Haiti) * '' Zanthoxylum andamanicum''

Kurz Kurz is a German-language surname. Notable people with the surname include: * Heinz D. Kurz (born 1946), Austrian economics professor * Hermann Kurz (1813–1873), German poet and novelist * Isolde Kurz (1853–1944), German poet and short story ...

(

Andaman Islands The Andaman Islands () are an archipelago, made up of 200 islands, in the northeastern Indian Ocean about southwest off the coasts of Myanmar's Ayeyarwady Region. Together with the Nicobar Islands to their south, the Andamans serve as a mari ...

) * ''

Zanthoxylum andinum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the Family (biology), family Rutaceae that are native to warm temperate and Subtropics, subtropical areas worldwide. It is the type gen ...

'' Reynel (Ecuador) * ''

Zanthoxylum anison ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

L.O.Williams Louis Otho (Otto) Williams (1908 – 1991) was a botanist from Wyoming. He received his BA and MA from the University of Wyoming then a PhD from Washington University in St. Louis. He went on to be editor of the ''American Orchid Society#Publicati ...

(Guatemala) * ''

Zanthoxylum anthyllidifolium ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

Guillaumin Guillaumin is a French surname. Notable people with the surname include: *André Guillaumin (1885–1974), French botanist *Armand Guillaumin Armand Guillaumin (; February 16, 1841 – June 26, 1927) was a French Impressionist painter and l ...

(Vietnam) * ''

Zanthoxylum apiculatum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the Family (biology), family Rutaceae that are native to warm temperate and Subtropics, subtropical areas worldwide. It is the type gen ...

'' ( Sandwith) P.G.Waterman (South America) * ''

Zanthoxylum arborescens ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

Rose A rose is either a woody perennial plant, perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred Rose species, species and Garden roses, tens of thousands of cultivar ...

(Mexico, Nicaragua) * '' Zanthoxylum armatum''

DC. Augustin Pyramus (or Pyrame) de Candolle (, , ; 4 February 17789 September 1841) was a Swiss botanist. René Louiche Desfontaines launched de Candolle's botanical career by recommending him at a herbarium. Within a couple of years de Candolle ...

– winged prickly ash (Indian subcontinent to temperate east Asia and Malesia) * '' Zanthoxylum atchoum'' ( Aké Assi) P.G.Waterman (

Côte d'Ivoire Ivory Coast, also known as Côte d'Ivoire and officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital city of Yamoussoukro is located in the centre of the country, while its largest city and ...

) * '' Zanthoxylum austrosinense'' C.C.Huang (southern China) * ''

Zanthoxylum avicennae ''Zanthoxylum avicennae'' is a woody plant in the family Rutaceae. Description ''Z. avicennae'' is a deciduous tree that reaches 15 meters high. The trunk has chicken-foot-like thorns, the thorn base is oblate and thickened, shaped like a bulge, ...

'' ( Lam.) DC. (southern China to western and central Malesia) * ''

Zanthoxylum backeri ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' (

Bakh.f. Reinier Cornelis Bakhuizen van den Brink Jr. (11 September 1911, Panjinangan, Sukabumi Regency, Java – 1 May 1987, Leiden) was a Dutch botanist. He was the son of Djahini of Tjiampea and Dutch botanist Reinier Cornelis Bakhuizen van den Br ...

)

T.G.Hartley Thomas Gordon Hartley (9 January 1931 in Beaumont, Texas – 8 March 2016 in Canberra, Australia) was an American botanist. Biography In 1955 Hartley graduated in botany with the academic degree Bachelor of Science at the University of Wisconsin-E ...

* ''

Zanthoxylum beechyanum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

K.Koch Karl Heinrich Emil Koch (6 June 1809 – 25 May 1879) was a German botanist. He is best known for his botanical explorations in the Caucasus region, including northeast Turkey. Most of his collections have today been lost. He is also known as t ...

(Bonin Islands, Ryukya Islands) * ''

Zanthoxylum bifoliolatum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

Leonard Leonard or ''Leo'' is a common English language, English masculine given name and a surname. The given name and surname originate from the Old High German ''Leonhard'' containing the prefix ''levon'' ("lion") from the Greek wikt:Λέων#Greek, Λ ...

– Maricao prickly ash (Hispaniola to Puerto Rico) * ''

Zanthoxylum bissei ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' Beurton (Cuba) * '' Zanthoxylum bonifaziae''

Cornejo Cornejo is a Spanish surname meaning someone who lived by a Dogwood tree or who lived in one of several places called Cornejo. It may refer to: * Aldo Cornejo (born 1955), Chilean politician * Alfredo Cornejo (boxer) (1933–2021), Chilean boxer * ...

& Reynel (Ecuador) * '' Zanthoxylum bouetense'' (

Pierre Pierre is a masculine given name. It is a French form of the name Peter. Pierre originally meant "rock" or "stone" in French (derived from the Greek word πέτρος (''petros'') meaning "stone, rock", via Latin "petra"). It is a translation ...

ex Letouzey) P.G.Waterman (Congo, Gabon) * ''

Zanthoxylum brachyacanthum ''Zanthoxylum brachyacanthum'', known as thorny yellow-wood, satinwood, satin tree or scrub mulga, is a species of flowering plant in the family Rutaceae and is endemic to north-eastern Australia. It is a rainforest shrub or tree with thick, cone ...

F.Muell. Baron Sir Ferdinand Jacob Heinrich von Mueller, (; 30 June 1825 – 10 October 1896) was a German-Australian physician, geographer, and most notably, a botanist. He was appointed government botanist for the then colony of Victoria (state), Vic ...

– thorny yellowwood (New South Wales, Queensland) * ''

Zanthoxylum brisasanum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' ( Cuatrec.) P.G.Waterman (Colombia to Peru) * ''

Zanthoxylum buesgenii ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the Family (biology), family Rutaceae that are native to warm temperate and Subtropics, subtropical areas worldwide. It is the type gen ...

'' (

) P.G.Waterman (southern Nigeria to west central tropical Africa) * ''

Maxim. Carl Johann Maximovich, also Karl Ivanovich Maximovich (Russian: Карл Иванович Максимович; 23 November 1827 – 16 February 1891) was a Russian botanist. Maximovich spent most of his life studying the flora of the countries ...

– Chinese/Sichuan pepper (Himalaya to China) * '' Zanthoxylum burkillianum'' Babu * '' Zanthoxylum calcicola'' C.C.Huang (China) * ''

Zanthoxylum campicola ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' Reynel (Colombia) * ''

Zanthoxylum canalense ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' (Guillaumin) P.G.Waterman * '' Zanthoxylum capense'' ( Thunb.) Harv. (southern Africa) * ''

Zanthoxylum caribaeum ''Zanthoxylum caribaeum'' is a species of plant in the family Rutaceae. It is found in Belize, Guatemala, and Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, ...

'' Lam. – yellow prickly ash (Mexico to tropical America) * ''

Zanthoxylum caudatum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' Alston (Sri Lanka) * ''

Zanthoxylum celebicum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' Koord. (Sulawesi) * ''

Zanthoxylum chalybeum ''Zanthoxylum chalybeum'' is an aromatic deciduous shrub or tree within the family Rutaceae. It is also known as the lemon scented knobwood. Description A shrub or tree that can grow up to 12 m in height with a large crown; the Trunk (botany), ...

'' Engl. (

Belize Belize is a country on the north-eastern coast of Central America. It is bordered by Mexico to the north, the Caribbean Sea to the east, and Guatemala to the west and south. It also shares a maritime boundary with Honduras to the southeast. P ...

Guatemala Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...

Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Ocean at the Gulf of Fonseca, ...

) * ''

Zanthoxylum chevalieri ''Zanthoxylum chevalieri'' is a species of plant in the family Rutaceae. It is endemic to Ghana. It is threatened by habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longe ...

'' P.G.Waterman (western tropical Africa) * ''

Zanthoxylum chuquisaquense ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' Reynel (Bolivia) * '' Zanthoxylum ciliatum'' Engl. (Mexico to northern Venezuela) * '' Zanthoxylum claessensii'' ( De Wild.) P.G.Waterman (west central tropical Africa) * ''

Zanthoxylum clava-herculis ''Zanthoxylum clava-herculis'', the Hercules' club, Hercules-club, pepperwood, or southern prickly ash, is a spiny tree or shrub native to the southeastern United States. It grows to 10–17 m tall and has distinctive spined thick, corky lumps 2� ...

'' L. – Hercules' club, pepperbark, southern prickly ash, West Indian yellowwood (central and southeastern United States to eastern Mexico) * ''

Zanthoxylum coco ''Zanthoxylum coco'' (also known as ''Fagara coco'') is an evergreen tree of the family Rutaceae, native to Argentina and Bolivia where it grows in the wild, mostly in spiniferous forests of the low mountain ranges of the western Chaco. It is c ...

Gillies Gillies is both a Scottish surname and a given name shared by several notable people: Surname uses Politicians * Duncan Gillies (1834–1903), Australian colonial and state politician * James McPhail Gillies (1924–2015), Canadian national po ...

Hook.f. Sir Joseph Dalton Hooker (30 June 1817 – 10 December 1911) was a British botanist and explorer in the 19th century. He was a founder of geographical botany and Charles Darwin's closest friend. For 20 years he served as director of the Ro ...

f. &

Arn. George Arnott Walker Arnott of Arlary (6 February 1799 – 17 April 1868) was a Scottish botanist. He collaborated with botanists from around the world and served as a regius professor of botany at the University of Glasgow. An orchid genus ' ...

– coco, cochucho, smelly sauco (

Argentina Argentina, officially the Argentine Republic, is a country in the southern half of South America. It covers an area of , making it the List of South American countries by area, second-largest country in South America after Brazil, the fourt ...

Bolivia Bolivia, officially the Plurinational State of Bolivia, is a landlocked country located in central South America. The country features diverse geography, including vast Amazonian plains, tropical lowlands, mountains, the Gran Chaco Province, w ...

Paraguay Paraguay, officially the Republic of Paraguay, is a landlocked country in South America. It is bordered by Argentina to the Argentina–Paraguay border, south and southwest, Brazil to the Brazil–Paraguay border, east and northeast, and Boli ...

) * ''

Zanthoxylum collinsiae ''Zanthoxylum collinsiae'' is a woody plant in the family Rutaceae The Rutaceae () is a family (biology), family, commonly known as the rue

'' Craib (China to Indochina) * ''

Zanthoxylum comosum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' (

Herzog (; feminine ; masculine plural ; feminine plural ) is a German hereditary title held by one who rules a territorial duchy, exercises feudal authority over an estate called a duchy, or possesses a right by law or tradition to be referred to ...

) P.G.Waterman (central Bolivia) * '' Zanthoxylum compactum'' (

Huber Huber is a German-language surname. It derives from the German word meaning hide, a unit of land a farmer might possess, granting them the status of a free tenant. It is in the top ten most common surnames in the German-speaking world, especial ...

ex Albuq.) P.G.Waterman (Venezuela to western South America) * '' Zanthoxylum conspersipunctatum''

Merr. Elmer Drew Merrill (October 15, 1876 – February 25, 1956) was an American botanist and taxonomist. He spent more than twenty years in the Philippines where he became a recognized authority on the flora of the Asia-Pacific region. Through ...

Perry Perry or pear cider is an alcoholic beverage made from fermented pears, traditionally in England (particularly Gloucestershire, Herefordshire, and Worcestershire), parts of South Wales, France (especially Normandy and Anjou), Canada, Austral ...

(New Guinea) * ''

Zanthoxylum cucullatipetalum ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' Guillaumin (Vietnam) * '' Zanthoxylum davyi'' (

I.Verd. Inez Clare Verdoorn (15 June 1896 – 2 April 1989) was a South African botanist and taxonomist, noted for her major revisions of plant families and genera. She is also a niece of Eugène Marais, Eugene Nielen Marais, lawyer, Natural history, na ...

) P.G.Waterman – forest knobwood (Zimbabwe to South Africa) * ''

Zanthoxylum decaryi ''Zanthoxylum'' is a genus of about 250 species of deciduous and evergreen trees, shrubs and climbers in the family Rutaceae that are native to warm temperate and subtropical areas worldwide. It is the type genus of the tribe Zanthoxyleae i ...

'' H.Perrier (Madagascar) * ''

Zanthoxylum delagoense ''Zanthoxylum delagoense'' is a species of plant in the family Rutaceae. It is endemic to Mozambique Mozambique, officially the Republic of Mozambique, is a country located in Southeast Africa bordered by the Indian Ocean to the east, Tan ...

'' P.G.Waterman (southern Mozambique) * ''Zanthoxylum deremense'' (Engl.) John Ongayo Kokwaro, Kokwaro (Malawi, Tanzania) * ''Zanthoxylum dimorphophyllum'' Hemsl. (southern China to Indochina, Taiwan) * ''Zanthoxylum dinklagei'' (Engl.) P.G.Waterman (southern Nigeria to western central tropical Africa) * ''Zanthoxylum dipetalum'' H.Mann – Kāwau (Hawaii) * ''Zanthoxylum dissitum'' Hemsl. (central and southern China to Hainan) * ''Zanthoxylum diversifolium'' Warb. (New Guinea) * ''Zanthoxylum djalma-batistae'' (Albuq.) P.G.Waterman (Brazil) * ''Zanthoxylum domingense'' (Karl (Carl) Wilhelm Leopold Krug, Krug & Urb.) J.Jiménez Alm. (Haiti) * ''Zanthoxylum dumosum'' A.Rich. (Cuba) * ''Zanthoxylum echinocarpum'' Hemsl. (southern China) * ''Zanthoxylum ekmanii'' (Urb.) Alain (southern Mexico to southern tropical America, western Cuba) * ''Zanthoxylum eliasii'' Duncan MacNair Porter, D.M.Porter (Panama to Peru) * ''Zanthoxylum engleri'' Waterm. * ''Zanthoxylum esquirolii'' Augustin Abel Hector Léveillé, H.Lév. (southern central China) * ''Zanthoxylum fagara'' (L.) Charles Sprague Sargent, Sarg. – Lime prickly ash (tropical and subtropical America) * ''Zanthoxylum fauriei'' (Takenoshin Nakai, Nakai) Jisaburo Ohwi, Ohwi – lesser ailanthus-like prickly ash (South Korea, Japan) * ''Zanthoxylum finlaysonianum'' Nathaniel Wallich, Wall. (Thailand) * ''Zanthoxylum flavum'' Martin Vahl, Vahl – West Indian satinwood (Caribbean) * ''Zanthoxylum foliolosum'' Donn.Sm. (Mexico to Nicaragua) * ''Zanthoxylum forbesii'' T.G.Hartley (Sumatra) * ''Zanthoxylum formiciferum'' ( Cuatrec.) P.G.Waterman (Colombia to northern Peru) * ''Zanthoxylum gardneri'' Engl. (Brazil) * ''Zanthoxylum gentryi'' Reynel (western Colombia) * ''Zanthoxylum ghisbreghtii'' Turcz. (Gulf of Mexico) * ''Zanthoxylum gillespieanum'' (A.C.Sm) A.C.Sm. (Fiji) * ''

'' (De Wild.) P.G.Waterman (tropical Africa) * ''Zanthoxylum glomeratum'' C.C.Huang (China) * ''Zanthoxylum grandifolium'' Tul. (Colombia) * ''Zanthoxylum haitiense'' (Urb.) J.Jiménez Alm. (Haiti) * ''Zanthoxylum hamadryadicum'' José Rubens Pirani, Pirani (north-eastern Brazil) * ''Zanthoxylum harrisii'' Percy Wilson (botanist), P.Wilson (Jamaica) * ''Zanthoxylum hartii'' [(Krug & Urb.) P.Wilson (Jamaica) * ''Zanthoxylum hawaiiense'' William Hillebrand, Hillebr. – a'e, Hawai'i prickly ash (Hawaii) * ''Zanthoxylum heitzii'' (Aubrév. & Pellegr.) P.G.Waterm. (west central tropical Africa * ''Zanthoxylum heterophyllum'' ( Lam.) Sm. (Mascarenes) * ''Zanthoxylum holtzianum'' (Engl.) P.G.Waterman (southern Somalia to northern Mozambique) * ''Zanthoxylum huberi'' P.G.Waterman (South America) * ''Zanthoxylum humile'' (Eileen Adelaide Bruce, E.A.Bruce P.G.Waterman (Zimbabwe to Mozambique and Mpumalanga) * ''Zanthoxylum impressinervium'' Reynel (Colombia) * ''Zanthoxylum integrifoliolum'' Adolph Daniel Edward Elmer, Elmer (Taiwan to the Philippines) * ''Zanthoxylum iwahigense'' Reynel (the Philippines) * ''Zanthoxylum jamaicense'' P.Wilson (Jamaica) * ''Zanthoxylum juniperinum'' Poepp. (Central America) * ''Zanthoxylum kauaense'' Asa Gray, A.Gray – Kauai prickly ash (Hawaii) * ''Zanthoxylum khasianum''

(Assam to China (W. Yunnan)) * ''Zanthoxylum kleinii'' (R.S.Cowan) P.G.Waterman (Brazil) * ''Zanthoxylum kwangsiense'' (Hand.-Mazz.) Woon Young Chun, Chun ex C.C.Huang (China) * ''Zanthoxylum laetum'' Emmanuel Drake del Castillo, Drake (China to Vietnam) * ''Zanthoxylum laurentii'' (De Wild.) P.G.Waterman (Congo to Democratic Republic of Congo) * ''Zanthoxylum leiboicum'' C.C.Huang (China) * ''Zanthoxylum lemairei'' (De Wild.) P.G.Waterman (western tropical Africa to Uganda) * ''Zanthoxylum lenticellosum'' (Urb. & Ekman) J.Jiménez Alm. (Dominican Republic) * ''Zanthoxylum lenticulare'' Reynel (Colombia to Venezuela) * ''Zanthoxylum lepidopteriphilum'' Reynel (Ecuador to Peru, north-west Venezuela) * ''Zanthoxylum leprieurii'' Guill. & Perr. (tropical and southern Africa) * ''Zanthoxylum leratii'' Guillaumin (New Caledonia) * ''Zanthoxylum liboense'' C.C.Huang (China) * ''Zanthoxylum limoncello'' Planch. & Oerst. (Mexico to Colombia) * ''Zanthoxylum lindense'' (Engl.) Kokwaro (Tanzania) * ''Zanthoxylum macranthum'' (Hand.-Mazz.) C.C. Huang (south-east Tibet to southern China) * ''Zanthoxylum madagascariense'' John Gilbert Baker, Baker (Madagascar) * ''Zanthoxylum magnifasciculatum'' Reynel (Venezuela) * ''Zanthoxylum mananarense'' H.Perrier (Madagascar) * ''Zanthoxylum mantaro'' (J.F.Macbr.) J.F.Macbr. (Peru) * ''Zanthoxylum maranionense'' Reynel (Peru) * ''Zanthoxylum martinicense'' ( Lam.) DC. – white prickly ash * ''Zanthoxylum mayu'' Carlo Luigi Giuseppe Bertero, Bertero (Juan Fernández Islands) * ''Zanthoxylum megistophyllum'' (B.L.Burtt) T.G.Hartley (Solomon Islands) * ''Zanthoxylum melanostictum'' Schltdl. & Cham. (Mexico to north-west Venezuela and Peru) * ''Zanthoxylum mezoneurispinosum'' ( Aké Assi) William D. Hawthorne, W.D.Hawth. (Liberia to Ivory Coast) * ''Zanthoxylum micranthum'' Hemsl. (China) * ''Zanthoxylum mildbraedii'' (Engl.) P.G.Waterman (east central tropical Africa to south-west Kenya * ''Zanthoxylum minahassae'' Koord. (Sulawesi) * ''Zanthoxylum molle'' Alfred Rehder, Rehder (China) * ''Zanthoxylum mollissimum'' (Engl.) P.Wilson (Mexico to Colombia) * ''Zanthoxylum monogynum'' Augustin Saint-Hilaire, A.St.-Hil. (eastern Bolivia to Brazil) * ''Zanthoxylum motuoense'' C.C.Huang * ''Zanthoxylum multijugum'' Franch. (China) * ''Zanthoxylum myriacanthum'' Wall. ex Hook.f. (eastern Himalaya to southern China and western and central Malesia) * ''Zanthoxylum myrianthum'' (A.C.Sm.) P.G.Waterman (Fiji) * ''Zanthoxylum nadeaudii'' Emmanuel Drake del Castillo, Drake (French Polynesia) * ''Zanthoxylum nashii'' P.Wilson (Cuba to Hispaniola) * ''Zanthoxylum nebuletorum'' (Herzog) P.G.Waterman (Bolivia) * ''Zanthoxylum nemorale'' Mart. (eastern Brazil) * ''Zanthoxylum neocaledonicum'' Baker f. (New Caledonia) * ''Zanthoxylum nigrum'' Mart. (Brazil) * ''

'' (Roxb.) DC. – shiny-leaf prickly ash (tropical and subtropical Asia, Queensland) * ''Zanthoxylum novoguineense'' T.G.Hartley (New Guinea) * ''Zanthoxylum oahuense'' Hillebr. – Oahu prickly ash (Hawaii) * ''Zanthoxylum ovalifolium'' Robert Wight, Wight (Andaman Island, Bangladesh, Myanmar, Queensland) * ''Zanthoxylum ovatifoliolatum'' (Engl.) Finkelstein (Angola to north-west Namibia) * ''Zanthoxylum oxyphyllum'' Edgew. (Nepal to China and Myanmar) * ''Zanthoxylum panamense'' P.Wilson (Mexico to Central America) * ''Zanthoxylum pancheri'' P.S.Green (New Caledonia) * ''Zanthoxylum paniculatum'' Balf.f. (Rodrigues in Mauritius) * ''Zanthoxylum paracanthum'' (Mildbr.) Kokwaro (south-east Kenya to central Tanzania) * ''Zanthoxylum paulae'' (Albuq.) P.G. Waterman (Guianas to northern Brazil) * ''Zanthoxylum pentandrum'' (Aubl.) Richard Alden Howard, R.A.Howard (Trinidad to northern Brazil) * ''Zanthoxylum petenense'' Cyrus Longworth Lundell, Lundell (Mexico to Guatemala) * ''Zanthoxylum petiolare'' A.St.-Hil. & Tul. (Bolivia to Brazil and northern Argentina) * ''Zanthoxylum piasezkii'' Maxim. (China) * ''Zanthoxylum pilosiusculum'' (Engl.) P.G.Waterman (Democratic Republic of Congo to Angola) * ''Zanthoxylum pilosulum'' Rehder & E.H.Wilson (south-central China) * ''Zanthoxylum pimpinelloides'' ( Lam.) DC. (Cuba to Hispaniola) * ''Zanthoxylum pinnatum'' (J.R.Forst. & G.Forst.) W.R.B.Oliv. (Norfolk Island, Lord Howe Island, south-west Pacific.) * ''

'' (L.) DC. (China to South Korea, Japan) * ''Zanthoxylum phyllopterum'' (Griseb.) Wright (Cuba) * ''Zanthoxylum piasezkii'' Maxim. * ''Zanthoxylum pilosiusculum'' (Engl.) P.G.Waterm. * ''Zanthoxylum pilosulum'' Rehder & E.H. Wilson * ''Zanthoxylum pimpinelloides'' ( Lam.) DC. * ''Zanthoxylum pinnatum'' Druce * ''

'' (L.) DC. – Japanese pepper, chopi, Korean pepper (China, Japan, Korea) * ''Zanthoxylum pluviatile'' T.G.Hartley (New Guinea and Solomon Islands) * ''Zanthoxylum poggei'' (Engl.) P.G.Waterman (western central tropical Africa) * ''Zanthoxylum psammophilum'' (Aké Assi) P.G.Waterman (Ivory Coast) * ''Zanthoxylum pseudoxyphyllum'' Babu (Ivory Coast) * ''Zanthoxylum pteracanthum'' Rehder & E.H.Wilson (China) * ''Zanthoxylum pucro'' D.M Porter (Panama to north-west Colombia) * ''Zanthoxylum punctatum'' Vahl – dotted prickly ash (Caribbean) * ''Zanthoxylum quassiifolium'' (Donn.Sm.) Standl. & Steyerm. (Mexico to Guatemala.) * ''Zanthoxylum quinduense'' Tul. (western South America to north-west Venezuela) * ''Zanthoxylum renieri'' (G.C.C.Gilbert) P.G.Waterman (Democratic Republic of Congo to Uganda) * ''Zanthoxylum retroflexum'' T.G.Hartley (Sumatra) * ''Zanthoxylum retusum'' (Albuq.) P.G.Waterman (Brazil) * ''

'' (Roxb.) DC. – Cape yellowwood, Indian ivy-rue, Indian pepper (tropical Asia, Queensland) * ''Zanthoxylum rhodoxylum'' (Urb.) P.Wilson (Jamaica) * ''Zanthoxylum rhoifolium'' Lam. (tropical and subtropical America) * ''Zanthoxylum rhombifoliolatum'' C.C.Huang (China) * ''Zanthoxylum riedelianum'' Engl. (Brazil to north-east Argentina) * ''Zanthoxylum rigidum'' Humb. & Bonpl. ex Willd. (western South America to Paraguay) * ''Zanthoxylum rubescens'' Planch. ex Hook. (western tropical Africa to south-west Kenya and Angola) * ''Zanthoxylum sapindifolium'' Wall. (Myanmar) * ''Zanthoxylum sarasinii'' Guillaumin (New Caledonia) * ''Zanthoxylum scandens'' Blume (southern China to Nansei-shoto and western Malesia) * ''

'' Philipp Franz von Siebold, Siebold & Zucc. – mastic-leaf prickly ash, Sichuan pepper (central and eastern China to temperate eastern Asia) * ''Zanthoxylum schlechteri'' Guillaumin (New Caledonia) * ''Zanthoxylum schreberi'' (J.F.Gmel.) Reynel ex C.Nelson (Mexico to Venezuela and Bolivia, Caribbean) * ''Zanthoxylum setulosum'' P.Wilson (Costa Rica to north-west Venezuela) * ''

'' Henry Fletcher Hance, Hance (China, South Korea, Taiwan) * ''Zanthoxylum spinosum'' (Sw.) Sw. (Florida to Caribbean) * ''Zanthoxylum spondiifolium'' Wall. (Myanmar) * ''Zanthoxylum sprucei'' Engl. (Ecuador to Peru and western Brazil) * ''Zanthoxylum stelligerum'' Turcz. (central and eastern Brazil) * ''Zanthoxylum stenophyllum'' Hemsl. (central and eastern central China) * ''Zanthoxylum subspicatum'' H. Perrier (Madagascar) * ''Zanthoxylum syncarpum'' (Tul.) Tul. ex Benjamin Daydon Jackson, B.D.Jacks. (Venezuela, eastern Brazil) * ''Zanthoxylum taediosum'' A.Rich. (Cuba) * ''Zanthoxylum tahitense'' (Nadeaud) Jacques Florence, J.Florence & Nicolas Hallé, N.Hallé (Society Islands) * ''Zanthoxylum tambopatense'' Reynel (Peru) * ''Zanthoxylum tetraphyllum'' (urb. & Ekman) J.Jiménez Alm. (Haiti) * ''Zanthoxylum tetraspermum'' Wight &

(southern India, Sri Lanka) * ''Zanthoxylum thomense'' (Engl.) A.Chev. ex P.G.Waterman (Gabon and Gulf of Guinea Island) * ''Zanthoxylum thornycroftii'' (I.Verd.) P.G.Waterman (Limpopo to Mpumalanga) * ''Zanthoxylum thouventii'' H.Perrier (Madagascar]) * ''Zanthoxylum tidorense'' Miq. (Maluku Islands) * ''Zanthoxylum timoriense'' Span. (Lesser Sunda Islands) * ''Zanthoxylum tingoassuiba'' A.St.-Hil. (eastern and southern Brazil) * ''Zanthoxylum tomentellum'' Hook.f. (Nepal to China and Myanmar) * ''Zanthoxylum tragodes'' (L.) DC. – Niaragato (Caribbean) * ''Zanthoxylum trijugum'' (Harvey Lawrence Dunkley, Dunkley) P.G.Waterman (central Tanzania to Zambia) * ''Zanthoxylum tsihanimposa'' H.Perrier (Madagascar) * ''Zanthoxylum undulatifolium'' Hemsl. (China) * ''Zanthoxylum unifoliatum'' Milton Groppo, Groppo & Pirani (Brazil) * ''Zanthoxylum usambarense'' (Engl.) Kokwara (Ethiopia to Rwanda) * ''Zanthoxylum usitatum'' Pierre ex Laness. (Vietnam) * ''Zanthoxylum venosum'' Leonard (Hispaniola) * ''Zanthoxylum verrucosum'' (Cuatrec.) P.G.Waterman (Colombia to Ecuador) * ''Zanthoxylum vinkii'' T.G.Hartley (New Guinea) * ''Zanthoxylum viride'' (A.Chev.) T.G.Hartley (west tropical Africa to Congo) * ''Zanthoxylum vitiense'' A.C.Sm. (Fiji) * ''Zanthoxylum wutaiense'' Ih Sheng Chen, I.S.Chen (Taiwan) * ''Zanthoxylum xichouense'' C.C.Huang (China) * ''Zanthoxylum yakumontanum'' (Junichi Sugimoto, Sugim.) Hidetoshi Nagamasu, Nagam. (Japan) * ''Zanthoxylum yuanjiangensis'' C.C.Huang (China) * ''Zanthoxylum zanthoxyloides'' ( Lam.) Bernhard Zepernick, Zepern. & Friedrich Karl Timler, Timler – Senegal prickly ash (western tropical Africa)

Unplaced species

The genus ''Fagara'' has been sunk into ''Zanthoxylum'', but , no name seemed to have been provided for ''Fagara externa'', which was regarded as an unplaced name by

References

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PDF
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External links

* * {{Authority control Zanthoxylum, Peppers Zanthoxyloideae genera Taxa named by Carl Linnaeus Dioecious plants