Xyloiuloidea is an extinct superfamily of
millipede
Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...
s that existed from the
Lower Devonian through the
Upper Pennsylvanian period in Europe and North America.
Description
Xyloiuloids are more or less cylindrical, with
sternites,
pleurites, and
tergites of each
body segment
Segmentation in biology is the division of some animal and plant body plans into a linear series of repetitive segments that may or may not be interconnected to each other. This article focuses on the segmentation of animal body plans, specifica ...
fused into a complete ring. Adults possess 40 to 50 body rings. The legs are no longer than half the height of the body. The body surface is marked by small parallel grooves (striations), which vary in surface coverage between xyloiuloid families.
Taxonomy
Xyloiuloidea comprises four families:
*†
Gaspestriidae
*†
Nyraniidae
*†
Plagiascetidae
*†
Xyloiulidae
The taxonomic history of Xyloiuloidea begins with
Orator F. Cook designating the family
Xyloiulidae in 1895. In 1969,
Richard L. Hoffman established the families
Nyraniidae and
Plagiascetidae, and placed all three extinct families in the extant (still-living) order
Spirobolida
Spirobolida is an Order (biology), order of "round-backed" millipedes containing approximately 500 species in 12 family (biology), families. Its members are distinguished by the presence of a "pronounced Suture (anatomy), suture that runs "verti ...
, as suborder "Xyloiulidea".
[Hoffman, Richard L., 1969. Myriapoda, exclusive of Insecta. R572–R606. In: Moore, R.C. (Ed.), ''Treatise on Invertebrate Paleontology'', Part R, Vol. 2. Geological Society of America and University of Kansas Press, Lawrence, KS.] In 2006, two new species were described and placed in the new family
Gaspestriidae, and group was reassigned as a superfamily of uncertain status (''
incertae sedis
or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'') within the
juliform millipedes, a group that includes the cylindrical, fused-bodied orders Spirobolida,
Spirostreptida
Spirostreptida is an order of long, cylindrical millipedes. There are approximately 1000 described species, making Spirostreptida the third largest order of millipedes after Polydesmida and Chordeumatida.
Description
Spirostreptida are generall ...
, and
Julida
Julida is an order of millipedes. Members are mostly small and cylindrical, typically ranging from in length. Eyes may be present or absent, and in mature males of many species, the first pair of legs is modified into hook-like structures. Addit ...
.
[
]
References
Millipede taxonomy
Arthropod superfamilies
Carboniferous myriapods
Devonian myriapods
Carboniferous arthropods of Europe
Carboniferous arthropods of North America
Devonian arthropods of Europe
Devonian arthropods of North America
Early Devonian first appearances
Pennsylvanian extinctions
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