Vegasaurus Molyi
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''Vegasaurus'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
elasmosaurid Elasmosauridae, often called elasmosaurs or elasmosaurids, is an extinct family of plesiosaurs that lived from the Hauterivian stage of the Early Cretaceous to the Maastrichtian stage of the Late Cretaceous period (c. 130 to 66 mya). The taxo ...
plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...
known from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
(early
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
stage)
Snow Hill Island Formation The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
of
Vega Island Vega Island () is an island in Antarctica, long and wide, which is the northernmost of the James Ross Island group and lies in the west part of Erebus and Terror Gulf. It is separated from James Ross Island by Herbert Sound and from Trinit ...
,
Antarctic Peninsula The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martin in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctica. ...
. It contains a single species, ''Vegasaurus molyi''.


Description

''Vegasaurus'' is unique among
elasmosaurid Elasmosauridae, often called elasmosaurs or elasmosaurids, is an extinct family of plesiosaurs that lived from the Hauterivian stage of the Early Cretaceous to the Maastrichtian stage of the Late Cretaceous period (c. 130 to 66 mya). The taxo ...
s in having 54 neck vertebrae. ''Callawayasaurus'' with a similar count of 56 lacks the dumbbell-shaped articular faces that are present on the vertebrae of ''Vegasaurus''. Additional traits rarely seen in other elasmosaurid include: atlas-axis complex with a visible keel on its lower surface, a side ridge that is present on neck vertebrae 8 to about 42, a well developed notch on lower surface in neck vertebrae before the seventh but not in the last five vertebrae, a
coracoid A coracoid is a paired bone which is part of the shoulder assembly in all vertebrates except therian mammals (marsupials and placentals). In therian mammals (including humans), a coracoid process is present as part of the scapula, but this is n ...
with projection from the middle of its lower surface, a
femur The femur (; : femurs or femora ), or thigh bone is the only long bone, bone in the thigh — the region of the lower limb between the hip and the knee. In many quadrupeds, four-legged animals the femur is the upper bone of the hindleg. The Femo ...
with strongly convex capitulum (head), the lack of pectoral and pelvic bars, and a
humerus The humerus (; : humeri) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius (bone), radius and ulna, and consists of three sections. The humeral upper extrem ...
with front facing knee and a strong expansion to the back ending in an accessory back facet (only shared by '' Morenosaurus'' and '' Kaiwhekea''), among other traits. It was about long.


Discovery and naming

''Vegasaurus'' is known solely from the
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
MLP 93-I-5-1, a nearly complete well preserved
postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...
l
skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...
(lacking the tip of the tail) housed at the La Plata Museum in La Plata, Argentina. Elements include the whole neck with 54 complete
cervical vertebrae In tetrapods, cervical vertebrae (: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In saurop ...
, three
pectoral Pectoral may refer to: * The chest region and anything relating to it. * Pectoral cross, a cross worn on the chest * a decorative, usually jeweled version of a gorget * Pectoral (Ancient Egypt), a type of jewelry worn in ancient Egypt * Pectora ...
vertebrae, 17 back vertebrae, three
sacral vertebrae The sacrum (: sacra or sacrums), in human anatomy, is a triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part of the pelvic cavity, ...
, the front and middle tail vertebrae, pectoral and pelvic girdles, forelimbs and hindlimbs, ribs, and 45
gastrolith A gastrolith, also called a stomach stone or gizzard stone, is a rock held inside a gastrointestinal tract. Gastroliths in some species are retained in the muscular gizzard and used to grind food in animals lacking suitable grinding teeth. In ...
s associated with the dorsal region. MLP 93-I-5-1 was discovered in 1989, by Eduardo Olivero, Daniel Martinioni, Francisco Mussel and Jorge Lusky, at Cape Lamb of
Vega Island Vega Island () is an island in Antarctica, long and wide, which is the northernmost of the James Ross Island group and lies in the west part of Erebus and Terror Gulf. It is separated from James Ross Island by Herbert Sound and from Trinit ...
at the edge of the Antarctic Peninsula of James Ross Archipelago (northernmost part of
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
). Excavations took places during three Antarctic summer expeditions in 1993, 1999 and 2005. MLP 93-I-5-1 was collected from an area of 3 square meters in a semi-articulated state, at locality "Assemblage 10", which belongs to the middle section of Cape Lamb Member of the
Snow Hill Island Formation The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
, dating to the early
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
stage of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
, about 72
million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...
. ''Vegasaurus'' was first described and named by José P. O'Gorman, Leonardo Salgado, Eduardo B. Olivero and Sergio A. Marenssi in
2015 2015 was designated by the United Nations as: * International Year of Light * International Year of Soil __TOC__ Events January * January 1 – Lithuania officially adopts the euro as its currency, replacing the litas, and becomes ...
and the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Vegasaurus molyi''. The generic name is derived from ''Vega'', in reference to the
Vega Island Vega Island () is an island in Antarctica, long and wide, which is the northernmost of the James Ross Island group and lies in the west part of Erebus and Terror Gulf. It is separated from James Ross Island by Herbert Sound and from Trinit ...
where the holotype was found, and from
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
''saurus'', meaning "lizard", a common suffix for genus names of extinct reptile. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honors the Argentine
paleontological Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
technician Juan Jose Moly, for participating in 17 Antarctic field trips and in the collection of the holotype.


Classification

O’Gorman ''et al.'' (2015) tested the
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
position of ''Vegasaurus'' using a modified version of the Kubo ''et al.'' (2012) data-set. In the resultant topology, ''Vegasaurus'' nests with three other
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
South Pacific elasmosaurids, being '' Morenosaurus'', and two species from the
Weddell Sea The Weddell Sea is part of the Southern Ocean and contains the Weddell Gyre. Its land boundaries are defined by the bay formed from the coasts of Coats Land and the Antarctic Peninsula. The easternmost point is Cape Norvegia at Princess Martha C ...
region, the aristonectines ''
Aristonectes ''Aristonectes'' (meaning "best swimmer") is an extinct genus of large elasmosaurid plesiosaurs that lived during the Maastrichtian stage of the Late Cretaceous. Two species are known, ''A. parvidens'' and ''A. quiriquinensis'', whose fossil ...
'' and '' Kaiwhekea''. ''Vegasaurus'' differs from the two members of Aristonectinae by having elongated neck vertebrae, yet links them to taxa from the other side of the Pacific, e.g. '' Futabasaurus'' from
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
. The
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below follows their results, showing only the relationships within Elasmosauridae.


References

{{Taxonbar, from=Q21127348 Elasmosauridae Maastrichtian life Plesiosaurs of Antarctica Fossil taxa described in 2015 Sauropterygian genera