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''Vegasaurus'' is an
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...
of elasmosaurid
plesiosaur The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared ...
known from the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
(early
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
stage)
Snow Hill Island Formation The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
of
Vega Island Vega Island is a small island to the northwest of James Ross Island, on the Antarctic Peninsula. It is separated from James Ross Island by Herbert Sound. The island was named by Otto Nordenskjold, leader of the Swedish Antarctic Expedition (190 ...
,
Antarctic Peninsula The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martín in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctic ...
. It contains a single species, ''Vegasaurus molyi''.


Description

''Vegasaurus'' is unique among elasmosaurids in having 54 neck vertebrae. ''Callawayasaurus'' with a similar count of 56 lacks the dumbbell-shaped articular faces that are present on the vertebrae of ''Vegasaurus''. Additional traits rarely seen in other elasmosaurid include: atlas-axis complex with a visible keel on its lower surface, a side ridge that is present on neck vertebrae 8 to about 42, a well developed notch on lower surface in neck vertebrae before the seventh but not in the last five vertebrae, a coracoid with projection from the middle of its lower surface, a
femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...
with strongly convex capitulum (head), the lack of pectoral and pelvic bars, and a
humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...
with front facing knee and a strong expansion to the back ending in an accessory back facet (only shared by ''
Morenosaurus ''Morenosaurus'' is an extinct genus of plesiosaur from the Cretaceous of what is now California. The type species is ''Morenosaurus stocki'', first named by Samuel Welles in 1943, in honor of Dr. Chester Stock.Hilton, Richard P., ''Dinosaurs ...
'' and '' Kaiwhekea''), among other traits. It was about long and weighed .


Discovery and naming

''Vegasaurus'' is known solely from the
holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...
MLP 93-I-5-1, a nearly complete well preserved
postcrania Postcrania (postcranium, adjective: postcranial) in zoology and vertebrate paleontology is all or part of the skeleton apart from the skull. Frequently, fossil remains, e.g. of dinosaurs or other extinct tetrapods, consist of partial or isolated sk ...
l
skeleton A skeleton is the structural frame that supports the body of an animal. There are several types of skeletons, including the exoskeleton, which is the stable outer shell of an organism, the endoskeleton, which forms the support structure inside ...
(lacking the tip of the tail) housed at the La Plata Museum in La Plata, Argentina. Elements include the whole neck with 54 complete
cervical vertebrae In tetrapods, cervical vertebrae (singular: vertebra) are the vertebrae of the neck, immediately below the skull. Truncal vertebrae (divided into thoracic and lumbar vertebrae in mammals) lie caudal (toward the tail) of cervical vertebrae. In ...
, three pectoral vertebrae, 17 back vertebrae, three
sacral vertebrae The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30. The sacrum situates at the upper, back part ...
, the front and middle
tail vertebrae The vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordate ...
, pectoral and pelvic girdles, forelimbs and hindlimbs, ribs, and 45 gastroliths associated with the dorsal region. MLP 93-I-5-1 was discovered in 1989, by Eduardo Olivero, Daniel Martinioni, Francisco Mussel and Jorge Lusky, at Cape Lamb of
Vega Island Vega Island is a small island to the northwest of James Ross Island, on the Antarctic Peninsula. It is separated from James Ross Island by Herbert Sound. The island was named by Otto Nordenskjold, leader of the Swedish Antarctic Expedition (190 ...
at the edge of the Antarctic Peninsula of James Ross Archipelago (northernmost part of
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest contine ...
). Excavations took places during three Antarctic summer expeditions in 1993, 1999 and 2005. MLP 93-I-5-1 was collected from an area of 3 square meters in a semi-articulated state, at locality "Assemblage 10", which belongs to the middle section of Cape Lamb Member of the
Snow Hill Island Formation The Snow Hill Island Formation is an Maastrichtian, Early Maastrichtian geologic Formation (geology), formation found on James Ross Island, James Ross Island group, Antarctica. Remains of a Paraves, paravian Theropoda, theropod ''Imperobator anta ...
, dating to the early
Maastrichtian The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interval from ...
stage of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
, about 72
million years ago The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). ...
. ''Vegasaurus'' was first described and named by José P. O'Gorman,
Leonardo Salgado Leonardo Salgado is an Argentinean palaeontologist with a special interest in dinosaurs of the Cretaceous period and other investigations of the palaeobiology of fossil bearing geological formations. Salgado is the leading or coauthor of several t ...
, Eduardo B. Olivero and Sergio A. Marenssi in
2015 File:2015 Events Collage new.png, From top left, clockwise: Civil service in remembrance of November 2015 Paris attacks; Germanwings Flight 9525 was purposely crashed into the French Alps; the rubble of residences in Kathmandu following the Apri ...
and the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen ...
is ''Vegasaurus molyi''. The generic name is derived from ''Vega'', in reference to the
Vega Island Vega Island is a small island to the northwest of James Ross Island, on the Antarctic Peninsula. It is separated from James Ross Island by Herbert Sound. The island was named by Otto Nordenskjold, leader of the Swedish Antarctic Expedition (190 ...
where the holotype was found, and from Greek ''saurus'', meaning "lizard", a common suffix for genus names of extinct reptile. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honors the Argentine paleontological technician Juan Jose Moly, for participating in 17 Antarctic field trips and in the collection of the holotype.


Classification

O’Gorman ''et al.'' (2015) tested the
phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ...
position of ''Vegasaurus'' using a modified version of the Kubo ''et al.'' (2012) data-set. In the resultant topology, ''Vegasaurus'' nests with three other
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', the ...
South Pacific The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the continen ...
elasmosaurids, being ''
Morenosaurus ''Morenosaurus'' is an extinct genus of plesiosaur from the Cretaceous of what is now California. The type species is ''Morenosaurus stocki'', first named by Samuel Welles in 1943, in honor of Dr. Chester Stock.Hilton, Richard P., ''Dinosaurs ...
'', and two species from the Weddell Sea region, the
aristonectine Aristonectinae is a clade of plesiosaurs in the family Elasmosauridae. It includes the Late Cretaceous plesiosaurs ''Aristonectes'' and '' Kaiwhekea'', traditionally grouped with the Late Jurassic ''Tatenectes'' and ''Kimmerosaurus'' in the fami ...
s ''
Aristonectes ''Aristonectes'' (meaning 'best swimmer') is an extinct genus of plesiosaur from the Late Cretaceous Paso del Sapo Formation of what is now Argentina, the Quiriquina Formation of Chile and the Lopez de Bertodano Formation of Antarctica. The ...
'' and '' Kaiwhekea''. ''Vegasaurus'' differs from the two members of Aristonectinae by having elongated neck vertebrae, yet links them to taxa from the other side of the Pacific, e.g. ''
Futabasaurus ''Futabasaurus'' is a genus of plesiosaur from the Late Cretaceous of Fukushima, Japan. It was described and named in 2006, and was assigned to the family Elasmosauridae. The genus contains one species, ''F. suzukii''. Description The size of ...
'' from
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the north ...
. The cladogram below follows their results, showing only the relationships within Elasmosauridae.


References

{{Taxonbar, from=Q21127348 Elasmosaurids Maastrichtian life Plesiosaurs of Antarctica Fossil taxa described in 2015 Sauropterygian genera