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''Quercus lobata'', commonly called the valley oak or roble, is the largest of the
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
oak An oak is a hardwood tree or shrub in the genus ''Quercus'' of the beech family. They have spirally arranged leaves, often with lobed edges, and a nut called an acorn, borne within a cup. The genus is widely distributed in the Northern Hemisp ...
s. It is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the state, growing in interior valleys and foothills from Siskiyou to
San Diego San Diego ( , ) is a city on the Pacific coast of Southern California, adjacent to the Mexico–United States border. With a population of over 1.4 million, it is the List of United States cities by population, eighth-most populous city in t ...
counties.
Deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
, it requires year-round
groundwater Groundwater is the water present beneath Earth's surface in rock and Pore space in soil, soil pore spaces and in the fractures of stratum, rock formations. About 30 percent of all readily available fresh water in the world is groundwater. A unit ...
, and may live up to 600 years. Its thick, ridged
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
(resembling
alligator An alligator, or colloquially gator, is a large reptile in the genus ''Alligator'' of the Family (biology), family Alligatoridae in the Order (biology), order Crocodilia. The two Extant taxon, extant species are the American alligator (''A. mis ...
hide) and deeply lobed leaves are characteristic, and assist in identification.


Description

The valley oak may surpass in height, with a sturdy trunk possibly exceeding in diameter. The "Henley Oak", in Covelo, California, is the tallest known valley oak, at . The branches have an irregular, spreading and arching appearance. During autumn, the
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
turn a yellow to light orange color but become brown later in the season. In advancing age, the branches droop. The trees have pewter-colored rippled
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
. Typically, the leaves are long and are roundly and deeply lobed. The leaf width is approximately one half its length. Each leaf is matte green with an underneath pale green appearance; moreover, the leaf is covered with abundant soft fuzz, yielding an almost velvety feeling. When a fresh leaf is rubbed or broken, an
aroma An odor (American English) or odour (Commonwealth English; see spelling differences) is a smell or a scent caused by one or more volatilized chemical compounds generally found in low concentrations that humans and many animals can perceive v ...
tic scent is exuded, evoking a forest
odor An odor (American English) or odour ( Commonwealth English; see spelling differences) is a smell or a scent caused by one or more volatilized chemical compounds generally found in low concentrations that humans and many animals can perceive ...
. The wood is a dull brown approaching yellow. The
acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...
s are medium to dark brown and range from in length. The caps have deep stippling and are found most often as singlets, but occasionally as doublets. The acorns ripen from October to November. Viable acorns germinate in their first winter, and none remain by mid-winter.


Taxonomy

Valley oak is of the
white oak ''Quercus'' subgenus ''Quercus'' is one of the two subgenera into which the genus ''Quercus'' was divided in a 2017 classification (the other being subgenus ''Cerris''). It contains about 190 species divided among five sections. It may be calle ...
evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...
ary lineage, which is officially known as the section ''Quercus'' in
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
''Quercus''. This section comprises numerous oaks from California and elsewhere, which species share similar leaves, acorns, bark and
wood pulp Pulp is a fibrous Lignocellulosic biomass, lignocellulosic material prepared by chemically, semi-chemically, or mechanically isolating the cellulose fiber, cellulosic fibers of wood, fiber crops, Paper recycling, waste paper, or cotton paper, rag ...
. Early
settlers A settler or a colonist is a person who establishes or joins a permanent presence that is separate to existing communities. The entity that a settler establishes is a Human settlement, settlement. A settler is called a pioneer if they are among ...
used a variety of common names for the valley oak including: white oak, bottom oak, swamp oak, water oak and mush oak. The
Spaniard Spaniards, or Spanish people, are a Romance languages, Romance-speaking Ethnicity, ethnic group native to the Iberian Peninsula, primarily associated with the modern Nation state, nation-state of Spain. Genetics, Genetically and Ethnolinguisti ...
s, because the tree looked like the white oaks in Europe, called the tree "roble". The Concow tribe call the acorns lō-ē’ ( Konkow language).


Distribution and habitat

Valley oak tolerates cool wet winters and hot dry summers, but requires abundant water. It is most abundant in rich deep soils of valley floors below 600 meters (2000 feet) in elevation. Valley oak is found in dense
riparian A riparian zone or riparian area is the interface between land and a river or stream. In some regions, the terms riparian woodland, riparian forest, riparian buffer zone, riparian corridor, and riparian strip are used to characterize a ripar ...
forests, open foothill woodlands and valley
savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...
s. Commonly associated trees are coast live oak, interior live oak, blue oak, California black walnut, California sycamore and gray pine. The valley oak is widely distributed in: the California Central Valley; many smaller valleys such as the
San Fernando Valley The San Fernando Valley, known locally as the Valley, is an urbanized valley in Los Angeles County, Los Angeles County, California. Situated to the north of the Los Angeles Basin, it comprises a large portion of Los Angeles, the Municipal corpo ...
(original Spanish place-name from oak savannah),
Santa Clarita Valley The Santa Clarita Valley (SCV) is part of the upper watershed of the Santa Clara River in Southern California. The western portion of the valley was part of the Rancho San Francisco Mexican land grant. Located in Los Angeles County. The valle ...
,
Conejo Valley The Conejo Valley (Spanish language, Spanish: ''Valle del Conejo'', meaning "Valley of the Rabbit") is a region spanning both southeastern Ventura County, California, Ventura County and northwestern Los Angeles County, California, Los Angeles C ...
, and Santa Ynez Valley; the Inner Coast Ranges south of the Eel River; and the
Transverse Ranges The Transverse Ranges are a group of mountain ranges of Southern California, in the Pacific Coast Ranges physiographic region in North America. The Transverse Ranges begin at the southern end of the California Coast Ranges and lie within Santa Ba ...
from the
Tehachapi Mountains The Tehachapi Mountains (; Kawaiisu: ''Tihachipia'', meaning "hard climb") are a mountain range in the Transverse Ranges system of California in the Western United States. The range extends for approximately in southern Kern County and northwe ...
to the
Simi Hills The Simi Hills are a low rocky mountain range of the Transverse Ranges in eastern Ventura County, California, Ventura County and western Los Angeles County, California, Los Angeles County, of Southern California, United States. The range runs ma ...
,
Santa Susana Mountains The Santa Susana Mountains are a transverse range of mountains in Southern California, north of the city of Los Angeles, in the United States. The range runs east-west, separating the San Fernando and Simi valleys on its south from the Santa ...
. It is also present on
Santa Cruz Island Santa Cruz Island (Spanish language, Spanish: ''Isla Santa Cruz'', Chumashan languages, Chumash: ''Limuw'') is located off the southwestern coast of Ventura, California, United States. It is the largest island in California and largest of the ei ...
and Catalina Island in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...
. Some of the most picturesque stands are found in
Sonoma Valley Sonoma Valley is a valley located in southeastern Sonoma County, California, in the North Bay region of the San Francisco Bay Area. Known as the birthplace of the California wine industry, the valley is home to some of the earliest vineyards a ...
, Round Valley in
Mendocino County Mendocino County (; ''Mendocino'', Spanish language, Spanish for "of Antonio de Mendoza, Mendoza") is a County (United States), county located on the North Coast (California), North Coast of the U.S. state of California. As of the 2020 United S ...
and the southern
Salinas Valley The Salinas Valley (Spanish language, Spanish: ''Valle de Salinas'') is one of the major valleys and most productive Agriculture, agricultural regions in California. It is located west of the San Joaquin Valley and south of San Francisco Bay and ...
near the up-river reaches of the Salinas River.


Ecology

Like many oaks, valley oaks can tolerate wildfires. Although smaller individuals may be top-killed, most resprout from the root crown. A variety of
mammal A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s and
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
s eat the acorns, including the acorn woodpecker,
California scrub jay The California scrub jay (''Aphelocoma californica'') is a species of scrub jay native to western North America. It ranges from southern British Columbia throughout California and western Nevada near Reno to west of the Sierra Nevada (U.S.), Sie ...
, yellow-billed magpie, and
California ground squirrel The California ground squirrel (''Otospermophilus beecheyi''), also known as the Beechey ground squirrel, is a common and easily observed ground squirrel of the western United States and the Baja California Peninsula; it is common in Oregon and ...
. The acorns are also attacked by bruchid beetles, but can survive moderate levels of infestation. Globular
gall Galls (from the Latin , 'oak-apple') or ''cecidia'' (from the Greek , anything gushing out) are a kind of swelling growth on the external tissues of plants. Plant galls are abnormal outgrowths of plant tissues, similar to benign tumors or war ...
s up to several centimeters in diameter are frequently attached to twigs of mature specimens of valley oak. These house the
larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...
l stage of small indigenous
wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...
s '' Andricus quercuscalifornicus''. A related wasp species, '' Feron kingi'', produces small galls shaped like Hershey's kisses on leaf surfaces. The valley oak is the only known food plant of '' Chionodes petalumensis'' caterpillars.


Uses

The
acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...
s are sweet and edible; Native Americans including the
Southern Paiute people The Southern Paiute people () are a tribe of Native Americans who have lived in the Colorado River basin of southern Nevada, northern Arizona, and southern Utah. Bands of Southern Paiute live in scattered locations throughout this territory an ...
roasted them and ground the edible portion into meal to make into bread and mush. Difficulties in acquiring valley oak wood as well as issues stemming from its drying such as cracking and warping have shifted its consumption from a general purpose lumber to a primarily niche product. Valley oak wood has a small, but significant market for use in cabinetry though, and is also suitable for hardwood flooring. Tyloses present in the pores of valley oak wood increase its impermeability to fluids allowing it to be used in the production of water-tight vessels. Such vessels include wine barrels where valley oak wood sees limited role in the composition of and where it has similar properties to other white oaks such as a reduced tannin load compared to the red oaks and an open grain that allows for an increased transfer of oxygen.


Observational history

In 1792, the English explorer
George Vancouver Captain (Royal Navy), Captain George Vancouver (; 22 June 1757 – 10 May 1798) was a Royal Navy officer and explorer best known for leading the Vancouver Expedition, which explored and charted North America's northwestern West Coast of the Uni ...
noted on his expedition through the
Santa Clara Valley The Santa Clara Valley (Spanish language, Spanish: ''Valle de Santa Clara'') is a geologic trough in Northern California that extends south–southeast from San Francisco to Hollister, California, Hollister. The longitudinal valley is bordered ...
, after seeing an expanse of valley oaks: In the year 1861, William Henry Brewer, the chief botanist for the first
California Geological Survey The California Geological Survey, previously known as the California Division of Mines and Geology, is the California state geology, geologic agency. History Although it was not until 1880 that the California State Mining Bureau, predecessor to ...
wrote of the valley oaks that he saw in Monterey County: The Hooker Oak of
Chico, California Chico ( ; Spanish language, Spanish for "little") is the most populous city in Butte County, California, United States. Located in the Sacramento Valley region of Northern California, the city had a population of 101,475 in the 2020 United Sta ...
, was once considered the largest-known valley oak. When it fell on May 1, 1977, it was nearly and in circumference at from the ground.


See also

*
California oak woodland California oak woodland is a plant community found throughout the California chaparral and woodlands ecoregion of California in the United States and northwestern Baja California in Mexico. Oak woodland is widespread at lower elevations in coast ...
*
List of Quercus species The genus ''Quercus'' contains about 500 known species, plus about 180 hybrids between them. The genus, as is the case with many large genera, is divided into subgenera and sections. Traditionally, the genus ''Quercus'' was divided into the tw ...


References


External links

* * {{Authority control
lobata Lobata is an order of transparent marine invertebrates belonging to the phylum of Ctenophora in the class Tentaculata, and are commonly referred to as comb jellies or sea gooseberries. There are currently 19 extant known species in the order of ...
Endemic flora of California Trees of Northern America Natural history of the California chaparral and woodlands Natural history of the California Coast Ranges Natural history of the Central Valley (California) Natural history of the Channel Islands of California Natural history of the San Francisco Bay Area Natural history of the Santa Monica Mountains Natural history of the Transverse Ranges Trees of Mediterranean climate Plants described in 1801 Garden plants of North America Drought-tolerant trees Ornamental trees