The Late Triassic is the third and final
epoch
In chronology and periodization, an epoch or reference epoch is an instant in time chosen as the origin of a particular calendar era. The "epoch" serves as a reference point from which time is measured.
The moment of epoch is usually decided b ...
of the
Triassic
The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...
Period in the
geologic time scale
The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochro ...
, spanning the time between
Ma and Ma (million years ago). It is preceded by the
Middle Triassic Epoch and followed by the
Early Jurassic
The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...
Epoch. The corresponding
series of rock beds is known as the Upper Triassic. The Late Triassic is divided into the
Carnian
The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...
,
Norian
The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
and
Rhaetian
The Rhaetian is the latest age (geology), age of the Triassic period (geology), Period (in geochronology) or the uppermost stage (stratigraphy), stage of the Triassic system (stratigraphy), System (in chronostratigraphy). It was preceded by the N ...
ages.
Many of the first
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
s evolved during the Late Triassic, including ''
Plateosaurus'', ''
Coelophysis
''Coelophysis'' ( Traditional English pronunciation of Latin, traditionally; or , as heard more commonly in recent decades) is a genus of coelophysid Theropoda, theropod dinosaur that lived Approximation, approximately 215 to 201.4 million y ...
'', ''
Herrerasaurus'', and ''
Eoraptor''. The
Triassic–Jurassic extinction event
The Triassic–Jurassic (Tr-J) extinction event (TJME), often called the end-Triassic extinction, marks the boundary between the Triassic and Jurassic periods, . It represents one of five major extinction events during the Phanerozoic, profoundly ...
began during this epoch and is one of the five major mass extinction events of the Earth.
Etymology
The Triassic was named in 1834 by
Friedrich von Alberti, after a succession of three distinct rock layers (Greek meaning 'triad') that are widespread in southern
Germany
Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
: the lower
Buntsandstein
The Buntsandstein (German for ''coloured'' or ''colourful sandstone'') or Bunter sandstone is a lithostratigraphy, lithostratigraphic and allostratigraphy, allostratigraphic unit (a sequence of rock strata) in the Subsurface (geology), subsurface ...
(colourful sandstone'')'', the middle
Muschelkalk
The Muschelkalk (German for "shell-bearing limestone"; ) is a sequence of sedimentary rock, sedimentary rock strata (a lithostratigraphy, lithostratigraphic unit) in the geology of central and western Europe. It has a Middle Triassic (240 to 230 m ...
(shell-bearing limestone) and the upper
Keuper (coloured clay). The Late Triassic Series corresponds approximately to the middle and upper Keuper.
Dating and subdivisions
On the
geologic time scale
The geologic time scale or geological time scale (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochro ...
, the Late Triassic is usually divided into the Carnian, Norian, and Rhaetian ages, and the corresponding rocks are referred to as the Carnian, Norian, and Rhaetian stages.
Triassic
chronostratigraphy
Chronostratigraphy is the branch of stratigraphy that studies the ages of rock strata in relation to time.
The ultimate aim of chronostratigraphy is to arrange the sequence of deposition and the time of deposition of all rocks within a geological ...
was originally based on
ammonite
Ammonoids are extinct, (typically) coiled-shelled cephalopods comprising the subclass Ammonoidea. They are more closely related to living octopuses, squid, and cuttlefish (which comprise the clade Coleoidea) than they are to nautiluses (family N ...
fossils, beginning with the work of
Edmund von Mojsisovics in the 1860s. The base of the Late Triassic (which is also the base of the Carnian) is set at the first appearance of an ammonite, ''
Daxatina canadensis''. In the 1990s,
conodont
Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
s became increasingly important in the Triassic timescale, and the base of the Rhaetian is now set at the first appearance of a conodont, ''
Misikella posthernsteini''. , the base of the Norian has not yet been established, but will likely be based on conodonts.
The late Triassic is also divided into
land-vertebrate faunachrons. These are, from oldest to youngest, the
Berdyankian,
Otischalkian,
Adamanian,
Revueltian and
Apachean.
Late Triassic life
Following the
Permian–Triassic extinction event
The Permian–Triassic extinction event (also known as the P–T extinction event, the Late Permian extinction event, the Latest Permian extinction event, the End-Permian extinction event, and colloquially as the Great Dying,) was an extinction ...
, surviving organisms diversified. On land,
archosauriforms, most notably the
dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
became an important faunal component in the Late Triassic. Likewise,
bony fishes diversified in aquatic environments, most notably the
Neopterygii, to which nearly all
extant
Extant or Least-concern species, least concern is the opposite of the word extinct. It may refer to:
* Extant hereditary titles
* Extant literature, surviving literature, such as ''Beowulf'', the oldest extant manuscript written in English
* Exta ...
species of fish belong. Among the neopterygians,
stem-group teleosts and the now
extinct
Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
Pycnodontiformes
Pycnodontiformes is an extinct order of primarily marine bony fish. The group first appeared during the Late Triassic and disappeared during the Eocene. The group has been found in rock formations in Africa, Asia, Europe, North and South America. ...
became more abundant in the Late Triassic.
Carnian Age
The Carnian is the first age of the Late Triassic, covering the time interval from 237 to 227 million years ago. The earliest true dinosaurs likely appeared during the Carnian and rapidly diversified.
They emerged in a world dominated by
crurotarsan archosaurs
Archosauria () or archosaurs () is a clade of diapsid sauropsid tetrapods, with birds and crocodilians being the only extant taxon, extant representatives. Although broadly classified as reptiles, which traditionally exclude birds, the cladistics ...
(ancestors of
crocodile
Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...
s), predatory
phytosaurs, herbivorous armored
aetosaurs
Aetosaurs () are heavily armored reptiles belonging to the extinct order (biology), order Aetosauria (; from Ancient Greek, Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized Omnivore, omnivorous or Herbivore, herbivor ...
, and giant carnivorous
rauisuchians, which the dinosaurs gradually began to displace.
The emergence of the first dinosaurs came at about the same time as the
Carnian pluvial episode, at 234 to 232 Ma. This was a humid interval in the generally arid Triassic. It was marked by high extinction rates in marine organisms, but may have opened niches for the radiation of the dinosaurs.
Norian Age
The Norian is the second age of the Late Triassic, covering the time interval from about 227 to 208.5 million years ago. During this age, herbiverous
sauropodomorphs diversified and began to displace the large herbivorous
therapsids
Therapsida is a clade comprising a major group of eupelycosaurian synapsids that includes mammals and their ancestors and close relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including li ...
, perhaps because they were better able to adapt to the increasingly arid climate. However crurotarsans continued to occupy more ecological niches than dinosaurs.
[ In the oceans, neopterygian fish proliferated at the expense of ceratitid ammonites.
The Manicouagan impact event occurred 214 million years ago. However, no extinction event is associated with this impact.][Ramezani, J., S. A. Bowring, M. S. Pringle, F. D. Winslow, III, and E. T. Rasbury (2005). "The Manicouagan impact melt rock: a proposed standard for intercalibration of U-Pb and 40Ar/39Ar isotopic systems". 15th V.M. Goldsmidt Conference Abstract Volume, p. A321.]
Rhaetian Age
The Rhaetian Age was the final age of the Late Triassic, following the Norian Age, and it included the last major disruption of life until the end-Cretaceous mass extinction. This age of the Triassic is known for its extinction of marine reptile
Marine reptiles are reptiles which have become secondarily adapted for an aquatic or semiaquatic life in a marine environment. Only about 100 of the 12,000 extant reptile species and subspecies are classed as marine reptiles, including mari ...
s, such as nothosaurs and shastasaurs with the ichthyosaurs, similar to today's dolphin
A dolphin is an aquatic mammal in the cetacean clade Odontoceti (toothed whale). Dolphins belong to the families Delphinidae (the oceanic dolphins), Platanistidae (the Indian river dolphins), Iniidae (the New World river dolphins), Pontopori ...
. This age was concluded with the disappearance of many species that removed types of plankton
Plankton are the diverse collection of organisms that drift in Hydrosphere, water (or atmosphere, air) but are unable to actively propel themselves against ocean current, currents (or wind). The individual organisms constituting plankton are ca ...
from the ocean, as well as some organisms known for reef
A reef is a ridge or shoal of rock, coral, or similar relatively stable material lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic component, abiotic (non-living) processes such as deposition (geol ...
-building, and the pelagic conodont
Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
s. In addition to these species that became extinct, the straight-shelled nautiloid
Nautiloids are a group of cephalopods (Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and species rich, with over 2,500 recorded species. Th ...
s, placodonts, bivalve
Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class (biology), class of aquatic animal, aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed b ...
s, and many types of reptile
Reptiles, as commonly defined, are a group of tetrapods with an ectothermic metabolism and Amniotic egg, amniotic development. Living traditional reptiles comprise four Order (biology), orders: Testudines, Crocodilia, Squamata, and Rhynchocepha ...
did not survive through this age.
Climate and environment during the Triassic Period
During the beginning of the Triassic Period, the Earth consisted of a giant landmass known as Pangea, which covered about a quarter of Earth's surface. Towards the end of the period, continental drift occurred which separated Pangea. At this time, polar ice was not present because of the large differences between the equator and the poles. A single, large landmass similar to Pangea would be expected to have extreme seasons; however, evidence offers contradictions. Evidence suggests that there is arid climate as well as proof of strong precipitation. The planet's atmosphere and temperature components were mainly warm and dry, with other seasonal changes in certain ranges.
The Middle Triassic was known to have consistent intervals of high levels of humidity. The circulation and movement of these humidity patterns, geographically, are not known however. The major Carnian Pluvial Event stands as one focus point of many studies. Different hypotheses of the events occurrence include eruptions, monsoonal effects, and changes caused by plate tectonics. Continental deposits also support certain ideas relative to the Triassic Period. Sediments that include red beds, which are sandstones and shales of color, may suggest seasonal precipitation. Rocks also included dinosaur tracks, mudcracks, and fossils of crustaceans and fish, which provide climate evidence, since animals and plants can only live during periods of which they can survive through.
Evidence of environmental disruption and climate change
The Late Triassic is described as semiarid. Semiarid is characterized by light rainfall, having up to 10–20 inches of precipitation a year. The epoch had a fluctuating, warm climate in which it was occasionally marked by instances of powerful heat. Different basins in certain areas of Europe provided evidence of the emergence of the "Middle Carnian Pluvial Event." For example, the Western Tethys and German Basin was defined by the theory of a middle Carnian wet climate phase. This event stands as the most distinctive climate change within the Triassic Period. Propositions for its cause include:
* Different behaviors of atmospheric or oceanic circulation forced by plate tectonics that may have participated in modifying the carbon cycle
The carbon cycle is a part of the biogeochemical cycle where carbon is exchanged among the biosphere, pedosphere, geosphere, hydrosphere, and atmosphere of Earth. Other major biogeochemical cycles include the nitrogen cycle and the water cycl ...
and other scientific factors.
* heavy rains due to shifting of the earth
* sparked by eruptions, typically originating from an accumulation of igneous rocks, which could have included liquid rock or volcanic rock
Volcanic rocks (often shortened to volcanics in scientific contexts) are rocks formed from lava erupted from a volcano. Like all rock types, the concept of volcanic rock is artificial, and in nature volcanic rocks grade into hypabyssal and me ...
formations
Theories and concepts are supported universally, due to extensive areal proof of Carnian siliciclastic sediments. The physical positions as well as comparisons of that location to surrounding sediments and layers stood as basis for recording data. Multiple resourced and recurring patterns in results of evaluations allowed for the satisfactory clarification of facts and common conceptions on the Late Triassic. Conclusions summarized that the correlation of these sediments led to the modified version of the new map of Central Eastern Pangea, as well as that the sediment's relation to the "Carnian Pluvial Event" is greater than expected.
* High interest concerning the Triassic Period has fueled the need to uncover more information about the period's climate. The Late Triassic Epoch is classified as a phase entirely flooded with phases of monsoonal events. A monsoon affects large regions and brings heavy rains along with powerful winds. Field studies confirm the impact and occurrence of strong monsoonal circulation during this time frame. However, hesitations concerning climatic variability remains. Upgrading knowledge on the climate of a period is a difficult task to assess. Understanding of and assumptions of temporal and spatial patterns of the Triassic Period's climate variability still need revision. Diverse proxies hindered the flow of palaeontological evidence. Studies in certain zones are missing and could be benefited by collaborating the already existing but uncompared records of Triassic palaeoclimate.
* A specific physical piece of evidence was found. A fire scar on the trunk of a tree, found in southeast Utah, dates back to the Late Triassic. The feature was evaluated and paved the path to the conclusion of one fire's history. It was categorized through comparison of other modern tree scars. The scar stood as evidence of Late Triassic wildfire
A wildfire, forest fire, or a bushfire is an unplanned and uncontrolled fire in an area of Combustibility and flammability, combustible vegetation. Depending on the type of vegetation present, a wildfire may be more specifically identified as a ...
, an old climatic event.
Triassic–Jurassic extinction event
The extinction event that began during the Late Triassic resulted in the disappearance of about 76% of all terrestrial and marine life species, as well as almost 20% of taxonomic families. Although the Late Triassic Epoch did not prove to be as destructive as the preceding Permian
The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...
Period, which took place approximately 50 million years earlier and destroyed about 70% of land species, 57% of insect families as well as 95% of marine life
Marine life, sea life or ocean life is the collective ecological communities that encompass all aquatic animals, aquatic plant, plants, algae, marine fungi, fungi, marine protists, protists, single-celled marine microorganisms, microorganisms ...
, it resulted in great decreases in population sizes of many living organism populations.
The environment of the Late Triassic had negative effects on the conodont
Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard ...
s and ammonoid groups. These groups once served as vital index fossil
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock strata by using the fossil assemblages contained within them.Hine, Robert. "Biostratigraphy." ''Oxford Reference: Dictionary of Biology ...
s, which made it possible to identify feasible life span to multiple strata of the Triassic strata. These groups were severely affected during the epoch, and conodonts became extinct soon after (in the earliest Jurassic). Despite the large populations that withered away with the coming of the Late Triassic, many families, such as the pterosaur
Pterosaurs are an extinct clade of flying reptiles in the order Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 million to 66 million years ago). Pterosaurs are the earli ...
s, crocodile
Crocodiles (family (biology), family Crocodylidae) or true crocodiles are large, semiaquatic reptiles that live throughout the tropics in Africa, Asia, the Americas and Australia. The term "crocodile" is sometimes used more loosely to include ...
s, mammal
A mammal () is a vertebrate animal of the Class (biology), class Mammalia (). Mammals are characterised by the presence of milk-producing mammary glands for feeding their young, a broad neocortex region of the brain, fur or hair, and three ...
s and fish were very minimally affected. However, such families as the bivalves, gastropod
Gastropods (), commonly known as slugs and snails, belong to a large Taxonomy (biology), taxonomic class of invertebrates within the phylum Mollusca called Gastropoda ().
This class comprises snails and slugs from saltwater, freshwater, and fro ...
s, marine reptiles and brachiopod
Brachiopods (), phylum (biology), phylum Brachiopoda, are a phylum of animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear e ...
s were greatly affected and many species became extinct during this time.
Causes of the extinction
Most of the evidence suggests the increase of volcanic activity was the main cause of the extinction. As a result of the rifting of the super continent Pangea
Pangaea or Pangea ( ) was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia (continent), Siberia during the Carboniferous period ...
, there was an increase in widespread volcanic activity which released large amounts of carbon dioxide. At the end of the Triassic Period, massive eruptions occurred along the rift zone
A rift zone is a feature of some volcanoes, especially shield volcanoes, in which a set of linear cracks (or rifts) develops in a volcanic edifice, typically forming into two or three well-defined regions along the flanks of the vent. Believed ...
, known as the Central Atlantic Magmatic Province, for about 500,000 years. These intense eruptions were classified as flood basalt
A flood basalt (or plateau basalt) is the result of a giant volcanic eruption or series of eruptions that covers large stretches of land or the ocean floor with basalt lava. Many flood basalts have been attributed to the onset of a hotspot (geolo ...
eruptions, which are a type of large scale volcanic activity that releases a huge volume of lava in addition to sulfur dioxide
Sulfur dioxide (IUPAC-recommended spelling) or sulphur dioxide (traditional Commonwealth English) is the chemical compound with the formula . It is a colorless gas with a pungent smell that is responsible for the odor of burnt matches. It is r ...
and carbon dioxide. The sudden increase in carbon dioxide levels is believed to have enhanced the greenhouse effect
The greenhouse effect occurs when greenhouse gases in a planet's atmosphere insulate the planet from losing heat to space, raising its surface temperature. Surface heating can happen from an internal heat source (as in the case of Jupiter) or ...
, which acidified the oceans and raised average air temperature. As a result of the change in biological conditions in the oceans, 22% of marine families became extinct. In addition, 53% of marine genera and about 76–86% of all species became extinct, which vacated ecological niches; thus, enabling dinosaurs to become the dominant presence in the Jurassic Period. While the majority of the scientists agree that volcanic activity was the main cause of the extinction, other theories suggest the extinction was triggered by the impact of an asteroid, climate change, or rising sea levels.
Biological impact
The impacts that the Late Triassic had on surrounding environments and organisms were wildfire destruction of habitats and prevention of photosynthesis. Climatic cooling also occurred due to the soot in the atmosphere. Studies also show that 103 families of marine invertebrates became extinct at the end of the Triassic, but another 175 families lived on into the Jurassic. Marine and extant species were hit fairly hard by extinctions during this epoch. Almost 20% of 300 extant families became extinct; bivalves, cephalopods, and brachiopods suffered greatly. 92% of bivalves were wiped out episodically throughout the Triassic.
The end of the Triassic also brought about the decline of corals and reef builders during what is called a "reef gap". The changes in sea levels brought this decline upon corals, particularly the calcisponges and scleractinian corals. However, some corals would make a resurgence during the Jurassic Period. 17 brachiopod species were also wiped out by the end of the Triassic. Furthermore, conulariids became extinct.
References
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Further reading
GeoWhen Database – Late Triassic
{{Use dmy dates, date=June 2019
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Geological epochs
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