Udanoceratops
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''Udanoceratops'' (meaning "Udan's horned face") is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of large leptoceratopsid
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
that lived during the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
of
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
.


Discovery

''Udanoceratops'' was first named and described by
Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...
n
paleontologist Paleontology, also spelled as palaeontology or palæontology, is the scientific study of the life of the past, mainly but not exclusively through the study of fossils. Paleontologists use fossils as a means to classify organisms, measure geolo ...
Sergei Kurzanov Sergei Mikhailovich Kurzanov (Сергей Михайлович Курзанов, born 1947) is a Russian (formerly USSR, Soviet) paleontologist at the Paleontological Institute of the Russian Academy of Sciences. He is known mainly for his work in ...
in
1992 1992 was designated as International Space Year by the United Nations. Events January * January 1 – Boutros Boutros-Ghali of Egypt replaces Javier Pérez de Cuéllar of Peru as United Nations Secretary-General. * January 6 ** The Republ ...
and the
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Udanoceratops tschizhovi''. The
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
(PIN 3907/11) was collected during the 1980s at the Udan Sayr (also spelled Udyn Sayr or Üüden Sair) locality of the
Djadokhta Formation The Djadochta formation (sometimes transcribed and also known as Djadokhta, Djadokata, or Dzhadokhtskaya) is a highly fossiliferous geological formation in Central Asia, Gobi Desert, dating from the Late Cretaceous period, about 75 million ...
in
Ömnögovi Province Ömnögovi ( ''Ömnögovǐ'', ''South Gobi'') is an aimag (province) of Mongolia, located in the south of the country, in the Gobi Desert. Ömnögovi is Mongolia's largest aimag. The capital is Dalanzadgad. The province is rich in mineral dep ...
, dating to the
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
stage of the
Late Cretaceous The Late Cretaceous (100.5–66 Ma) is the more recent of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''cre ...
period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
. The generic name is derived from the name of the locality in which the holotype was found (Udan Sayr) and
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
''ceras/κέρας'' meaning "horn" and ''-ops/ωψ'' meaning "face". ''Udanoceratops'' is best known from the holotype specimen, which represents a large individual comprising a large, relatively well-preserved and almost complete
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
, and sparse body remains including vertebrae. In 1993 a large skull ("nearly 1 m long") assigned to ''Udanoceratops'' was reported from the nearby
Bayan Mandahu Formation The Bayan Mandahu Formation (also known as Wulansuhai Formation or Wuliangsuhai Formation) is a geological unit of "redbeds" located near the village of Bayan Mandahu in Inner Mongolia, China, in the Gobi Desert. It dates from the late Cretaceous ...
by paleontologist
Tomasz Jerzykiewicz Tomasz is a Polish given name, the equivalent of Thomas in English. Notable people with the given name include: *Tomasz Adamek (born 1976), Polish heavyweight boxer * Tomasz Arciszewski (1877–1955), Polish socialist politician and Prime Mini ...
. However, Polish paleontologist Łukasz Czepiński in 2020 pointed out that there are no referable specimens to ''Udanoceratops'' from the Bayan Mandahu collections, and it is most likely that these remains were confused with the concurring (and relatively large) ''
Protoceratops hellenikorhinus ''Protoceratops'' (; ) is a genus of small protoceratopsid dinosaurs that lived in Asia during the Late Cretaceous, around 75 to 71 million years ago. The genus ''Protoceratops'' includes two species: ''P. andrewsi'' and the larger ''P. hellenik ...
''. In 2004 Viktor S. Tereschhenko referred a juvenile specimen (PIN 4046/11) to ''Udanoceratops'' aff. ''tschizhovi'', from the Baga Tariach locality in
Dornogovi Province Dornogovi (, ''East Gobi'') is one of the 21 aimags (provinces) of Mongolia. It is located in the southeast of the country, bordering PR China's autonomous region of Inner Mongolia. Dornogovi is located in the Gobi Desert and frequent sand- and ...
, which Tereschhenko attributed to the Djadokhta Formation. Geological analyses carried out across fossiliferous localities of the
Gobi Desert The Gobi Desert (, , ; ) is a large, cold desert and grassland region in North China and southern Mongolia. It is the sixth-largest desert in the world. The name of the desert comes from the Mongolian word ''gobi'', used to refer to all of th ...
published by Mahito Watabe and team in 2010, indicates that this locality instead correlates best with the
Maastrichtian The Maastrichtian ( ) is, in the International Commission on Stratigraphy (ICS) geologic timescale, the latest age (geology), age (uppermost stage (stratigraphy), stage) of the Late Cretaceous epoch (geology), Epoch or Upper Cretaceous series (s ...
-stage
Barun Goyot Formation The Baruungoyot Formation (also known as Barun Goyot) is a geological formation dating to the Late Cretaceous Period. It is located within and is widely represented in the Gobi Desert Basin, in the Ömnögovi Province of Mongolia. Descriptio ...
. The assignment of this specimen, however, has varied since then from "''Udanoceratops''" sp., to ?''Udanoceratops'' sp.


Description

''Udanoceratops'' was a large ceratopsian, estimated at to have reached nearly long with a weight of . It is the largest leptoceratopsid known so far. The skull had a short frill and no horns over the eyes or nose. Its skull was about long. The lower jaw was distinctively robust.


Classification

''Udanoceratops'' belonged to the Ceratopsia (the name is derived
Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...
meaning 'horned face'), a group of herbivorous dinosaurs with
parrot Parrots (Psittaciformes), also known as psittacines (), are birds with a strong curved beak, upright stance, and clawed feet. They are classified in four families that contain roughly 410 species in 101 genus (biology), genera, found mostly in ...
-like beaks which thrived in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
and
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
during the
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
Period. It is placed within the
Leptoceratopsidae Leptoceratopsidae is an extinct family (biology), family of neoceratopsian dinosaurs from Asia, North America and possibly Europe. Leptoceratopsids resembled, and were closely related to, other neoceratopsians, such as the family (biology), famil ...
, as the only Asian representative at the time, along with the North American ''
Leptoceratops ''Leptoceratops'' (meaning 'small horn face') is a genus of ceratopsian dinosaur from the Late Cretaceous of North America. First found in Alberta in 1910, the type species ''Leptoceratops gracilis'' was named in 1914 by Barnum Brown for a parti ...
'', ''
Montanoceratops ''Montanoceratops'' is an extinct genus of small ceratopsian dinosaur that lived approximately 70 million years ago during the latter part of the Cretaceous Period (geology), Period in what is now Montana and Alberta. ''Montanoceratops'' was a s ...
'' and ''
Prenoceratops ''Prenoceratops'', (meaning 'bent or prone-horned face' and derived from Greek ''prene-/πρηνη-'' meaning 'bent forwards' or 'prone', ''cerat-/κερατ-'' meaning 'horn' and ''-ops/ωψ'' meaning 'face') is a genus of herbivorous ceratopsi ...
''.


Paleobiology

''Udanoceratops'', like all ceratopsians, was a
herbivore A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat ...
. The short, deep jaws would have given the animal a powerful bite. The toothless beak would have served to grasp and crop stems or leaves, and as in other leptoceratopsids, the teeth would have met with an action that combined shearing and crushing. The feeding adaptations seen in leptoceratopsids suggest a diet of relatively tough food items, however little is known about the plants that grew in the Gobi Desert during the Cretaceous. The type specimen does bear an injury to its face, possibly caused by the biting of another ''Udanoceratops''. This might also explain its hard bite as having some sort of social or defensive function.


See also

*
Timeline of ceratopsian research This timeline of ceratopsian research is a chronological listing of events in the History of paleontology, history of paleontology focused on the ceratopsians, a group of herbivorous marginocephalian dinosaurs that evolved parrot-like beaks, b ...


References


External links

* * {{Taxonbar, from=Q132581 Leptoceratopsidae Dinosaur genera Campanian dinosaurs Djadochta Formation Taxa named by Sergei Kurzanov Fossil taxa described in 1992 Dinosaurs of Mongolia