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''Trithemis annulata'', known commonly as the violet dropwing, violet-marked darter, purple-blushed darter or plum-coloured dropwing, is a species of
dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of true dragonfly are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
in the family
Libellulidae The skimmers or perchers and their relatives form the Libellulidae, the largest dragonfly family in the world. It is sometimes considered to contain the Corduliidae as the subfamily Corduliinae and the Macromiidae as the subfamily Macromiinae. Ev ...
. It is found in most of Africa, in the Middle East, in the Arabian Peninsula and southern Europe. These insects are called dropwings because of their habit of immediately lowering their wings after landing on a perch. Males of this species are violet-red with red veins in the wings while females are yellow and brown. Both sexes have red eyes.


Subspecies

The following
subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (morphology), but that can successfully interbreed. Not all species ...
are recognised as valid: *''Trithemis annulata annulata'' (Palisot de Beauvois, 1807) *''Trithemis annulata haematina'' (Rambur, 1842) *''Trithemis annulata scorteccii'' Nielsen, 1936


Description

''Trithemis annulata'' is a robust medium-sized species with a wingspan of . The mature male has a dark red head and a yellow labium with brown central spot. The eyes are red with white spots on the rear edge, and the frons is dark metallic purplish-red. The prothorax is violet with slightly darker longitudinal stripes. The membranous wings have distinctive red veins, the
pterostigma The pterostigma (plural: pterostigmata) is a group of specialized cells in the outer insect wing, wings of insects, which are often thickened or coloured, and thus stand out from other cells. It is particularly noticeable in dragonfly, dragonfli ...
is orange-brown and there is a large orange-brown splash at the base of the hind wings. The abdomen is fairly broad and is pinkish-violet, with purple markings on the top of each segment and blackish markings on the terminal three segments. Females are a similar size to males but the thorax is brownish and the abdomen is yellow with dark brown markings. The wings of females lack the red veins of males but have similar orange-brown patches. It is very similar in appearance to the red-veined dropwing (''Trithemis arteriosa''), but that species has a more slender abdomen and a wedge-shaped black area on either side of the tip of the abdomen.


Distribution

''Trithemis annulata'' is found in most of Africa and also in
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is geo ...
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Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders with ...
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India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the so ...
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Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
,
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
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Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical re ...
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Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
,
Kuwait Kuwait (; ar, الكويت ', or ), officially the State of Kuwait ( ar, دولة الكويت '), is a country in Western Asia. It is situated in the northern edge of Eastern Arabia at the tip of the Persian Gulf, bordering Iraq to the nort ...
,
Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus li ...
,
Mauritius Mauritius ( ; french: Maurice, link=no ; mfe, label=Mauritian Creole, Moris ), officially the Republic of Mauritius, is an island nation in the Indian Ocean about off the southeast coast of the African continent, east of Madagascar. It incl ...
,
Oman Oman ( ; ar, عُمَان ' ), officially the Sultanate of Oman ( ar, سلْطنةُ عُمان ), is an Arabian country located in southwestern Asia. It is situated on the southeastern coast of the Arabian Peninsula, and spans the mouth of t ...
,
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of ...
,
Qatar Qatar (, ; ar, قطر, Qaṭar ; local vernacular pronunciation: ), officially the State of Qatar,) is a country in Western Asia. It occupies the Qatar Peninsula on the northeastern coast of the Arabian Peninsula in the Middle East; it sh ...
,
Réunion Réunion (; french: La Réunion, ; previously ''Île Bourbon''; rcf, label= Reunionese Creole, La Rényon) is an island in the Indian Ocean that is an overseas department and region of France. It is located approximately east of the island ...
,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a list of transcontinental countries, transcontinental country located mainly on the Anatolia, Anatolian Peninsula in Western Asia, with ...
,
United Arab Emirates The United Arab Emirates (UAE; ar, اَلْإِمَارَات الْعَرَبِيَة الْمُتَحِدَة ), or simply the Emirates ( ar, الِْإمَارَات ), is a country in Western Asia (The Middle East). It is located at th ...
and
Yemen Yemen (; ar, ٱلْيَمَن, al-Yaman), officially the Republic of Yemen,, ) is a country in Western Asia. It is situated on the southern end of the Arabian Peninsula, and borders Saudi Arabia to the Saudi Arabia–Yemen border, north and ...
. It was also recorded in the Maltese islands in 2005 and found breeding there in 2007. With the advent of global warming, it is increasing its range northwards in Europe and in the late 1990s advanced from southern Spain and southern Italy into France, and from Greece and Turkey into Central Europe. One such place where it has become established is the floodplain of the River Ebro in northeastern Spain.


Ecology

''Trithemis annulata'' is an adaptable species, and the adults are able to tolerate a range of habitats including semi-arid rangeland. They can be seen flying near sluggish rivers, in marshes and also beside still-water ponds. They are sometimes seen in brackish water habitats, although it is unclear whether they actually breed in salty water. The
larva A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle. The ...
e develop rapidly, so these dragonflies are able to make use of temporary water bodies for breeding. Males are often to be seen perching on the twigs of waterside shrubs and on rocks in the sunshine, but in the evening or when the sun is obscured, they move into trees. The female ''T. annulata'' is thought to deposit her eggs by flying over the surface and dipping the tip of her abdomen in the water. The larvae develop in water where they are aggressive
predators Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill the ...
. The adults are also predators, using their excellent eyesight to detect prey and their legs to hold and carry their victims.


Status

''Trithemis annulata'' is a very common species throughout its wide range which includes most of Africa, the Mediterranean area, the Arabian Peninsula and the Near East. Although it may be threatened locally by habitat loss, in general it seems to be increasing in abundance, and the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
has assessed its conservation status as being of
least concern A least-concern species is a species that has been categorized by the International Union for Conservation of Nature (IUCN) as evaluated as not being a focus of species conservation because the specific species is still plentiful in the wild. T ...
.


References


External links

* *
Text for violet dropwing from South African Dragonfly Atlas
{{Taxonbar, from=Q947217 Dragonflies of Europe Insects described in 1807 annulata Taxonomy articles created by Polbot