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The tomato (, ), ''Solanum lycopersicum'', is a plant whose fruit is an edible
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
that is eaten as a vegetable. The tomato is a member of the
nightshade family Solanaceae (), commonly known as the nightshades, is a family of flowering plants in the order Solanales. It contains approximately 2,700 species, several of which are used as agricultural crops, medicinal plants, and ornamental plants. Many me ...
that includes
tobacco Tobacco is the common name of several plants in the genus '' Nicotiana'' of the family Solanaceae, and the general term for any product prepared from the cured leaves of these plants. More than 70 species of tobacco are known, but the ...
,
potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...
, and
chili pepper Chili peppers, also spelled chile or chilli ( ), are varieties of fruit#Berries, berry-fruit plants from the genus ''Capsicum'', which are members of the nightshade family Solanaceae, cultivated for their pungency. They are used as a spice to ...
s. It originated from and was domesticated in western
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...
. It was introduced to the
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
by the Spanish in the
Columbian exchange The Columbian exchange, also known as the Columbian interchange, was the widespread transfer of plants, animals, and diseases between the New World (the Americas) in the Western Hemisphere, and the Old World (Afro-Eurasia) in the Eastern Hemis ...
in the 16th century. Tomato plants are vines, largely
annual Annual may refer to: *Annual publication, periodical publications appearing regularly once per year **Yearbook **Literary annual *Annual plant *Annual report *Annual giving *Annual, Morocco, a settlement in northeastern Morocco *Annuals (band), a ...
and vulnerable to frost, though sometimes living longer in greenhouses. The flowers are able to self-fertilise. Modern varieties have been bred to ripen uniformly red, in a process that has impaired the fruit's sweetness and flavor. There are thousands of cultivars, varying in size, color, shape, and flavor. Tomatoes are attacked by many insect pests and nematodes, and are subject to diseases caused by viruses and by
mildew Mildew is a form of fungus. It is distinguished from its closely related counterpart, mold, largely by its colour: molds appear in shades of black, blue, red, and green, whereas mildew is white. It appears as a thin, superficial growth consisti ...
and
blight Blight is a specific symptom affecting plants in response to infection by a pathogenic organism. Description Blight is a rapid and complete chlorosis, browning, then death of plant tissues such as leaves, branches, twigs, or floral organs. A ...
fungi. The tomato has a strong savoury
umami Umami ( from ), or savoriness, is one of the five basic tastes. It is characteristic of broths and cooked meats. People taste umami through taste receptors that typically respond to glutamates and nucleotides, which are widely present in me ...
flavor, and is an important ingredient in cuisines around the world. It is used in
pizza Pizza is an Italian cuisine, Italian, specifically Neapolitan cuisine, Neapolitan, dish typically consisting of a flat base of Leavening agent, leavened wheat-based dough topped with tomato, cheese, and other ingredients, baked at a high t ...
s, pasta and other
sauces In cooking, a sauce is a liquid, cream, or semi- solid food, served on or used in preparing other foods. Most sauces are not normally consumed by themselves; they add flavour, texture, and visual appeal to a dish. ''Sauce'' is a French wor ...
, soups such as
gazpacho Gazpacho () or gaspacho (), also called Andalusian gazpacho (from Spanish ''gazpacho andaluz''), is a cold soup and drink made of raw, blended vegetables. It originated in the southern regions of the Iberian Peninsula and spread into other are ...
,
curries Curry is a dish with a sauce or gravy seasoned with spices, mainly derived from the interchange of Indian cuisine with European taste in food, starting with the Portuguese, followed by the Dutch and British, and then thoroughly internation ...
including
dhansak Dhansak is a popular Indian dish, originating among the Parsi community. It combines elements of Persian and Gujarati cuisine. Dhansak is made by cooking mutton or goat meat with a mixture of lentils and vegetables. This is served with ca ...
and
rogan josh Rogan josh ( English: /ˌroʊɡən ˈdʒɑʃ/);Rogan Josh
Oxford English Dictionary
), also spell ...
, as juice, and in Bloody Mary cocktails. Tomato festivals are held annually in
Buñol Buñol is a town and municipality in the province of Valencia, Spain. The municipality has an area of some 112 km2, and is situated approximately 38 km west of the provincial and autonomous community capital city, Valencia. It lies alon ...
, Spain, in
Reynoldsburg, Ohio Reynoldsburg is a city in Fairfield County, Ohio, Fairfield, Franklin County, Ohio, Franklin, and Licking County, Ohio, Licking counties in the U.S. state of Ohio. It is a suburban community in the Columbus, Ohio metropolitan area. The population ...
, and in
Närpes Närpes (Finland Swedish: ; ) is a town in Finland, located on the west coast of the country. Närpes is situated in Ostrobothnia, along the Gulf of Bothnia. The population of Närpes is approximately , while the sub-region has a population of ...
, Finland.


Naming


Etymology

The word ''tomato'' comes from the Spanish , which in turn comes from the
Nahuatl Nahuatl ( ; ), Aztec, or Mexicano is a language or, by some definitions, a group of languages of the Uto-Aztecan language family. Varieties of Nahuatl are spoken by about Nahuas, most of whom live mainly in Central Mexico and have smaller popul ...
word . The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
''lycopersicum'', meaning 'wolf peach', originated with
Galen Aelius Galenus or Claudius Galenus (; September 129 – AD), often Anglicization, anglicized as Galen () or Galen of Pergamon, was a Ancient Rome, Roman and Greeks, Greek physician, surgeon, and Philosophy, philosopher. Considered to be one o ...
, who used it to denote a plant that has never been identified.
Luigi Anguillara Luigi Anguillara, actually Luigi Squalermo, (born c. 1512 in Anguillara Sabazia, died September 1570 in Ferrara) was an Italian botanist. Life Little is known about Anguillara's early life. From 1539, he is attested at the private botanical gar ...
speculated in the 16th century that Galen's ''lycopersicum'' might be the tomato, and despite the impossibility of this identification, ''lycopersicum'' entered scientific use as a name for the fruit.


Pronunciation

The usual pronunciations of ''tomato'' are (in
North American English North American English (NAmE) encompasses the English language as spoken in both the United States and Canada. Because of their related histories and cultures, plus the similarities between the pronunciations (accents), vocabulary, and grammar ...
) and (in
British English British English is the set of Variety (linguistics), varieties of the English language native to the United Kingdom, especially Great Britain. More narrowly, it can refer specifically to the English language in England, or, more broadly, to ...
). The word's dual pronunciations were immortalized in Ira and
George Gershwin George Gershwin (; born Jacob Gershwine; September 26, 1898 – July 11, 1937) was an American composer and pianist whose compositions spanned jazz, popular music, popular and classical music. Among his best-known works are the songs "Swan ...
's 1937 song "
Let's Call the Whole Thing Off "Let's Call the Whole Thing Off" is a song written by George Gershwin and Ira Gershwin for the 1937 film ''Shall We Dance'', where it was introduced by Fred Astaire and Ginger Rogers as part of a celebrated dance duet on roller skates. The shee ...
" ("You like and I like / You like and I like ").


History

The likely wild ancestor of the tomato, the red-fruited ''
Solanum pimpinellifolium ''Solanum pimpinellifolium'', commonly known as the currant tomato or pimp, is a wild species of tomato native to Ecuador and Peru but naturalized elsewhere, such as the Galápagos Islands. Its small fruits are edible, and it is commonly grown in ...
'', is native to western South America, where it was probably first domesticated. The resulting domesticated plant, ancestral to the modern large-fruited tomato varieties, was probably the cherry tomato, ''S. lycopersicum'' var. ''cerasiforme''. However, genomic analysis suggests that the domestication process may have been more complex than this. ''S. lycopersicon'' var. ''cerasiforme'' may have existed before domestication, while traits supposedly typical of domestication may have been reduced in that variety and then reselected (in a case of
convergent evolution Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last comm ...
) in the cultivated tomato. The analysis predicts that var. ''cerasiforme'' appeared around 78,000 years ago, while the cultivated tomato originated around 7,000 years ago (5,000 BCE), with substantial uncertainty, making it unclear how humans may have been involved in the process. The Spanish first introduced tomatoes to Europe, where they became used in Spanish food. Elsewhere in Europe, its first use was ornamental, not least because it was understood to be related to the
nightshade Solanaceae (), commonly known as the nightshades, is a family of flowering plants in the order Solanales. It contains approximately 2,700 species, several of which are used as agricultural crops, medicinal plants, and ornamental plants. Many me ...
s and assumed to be poisonous.


Mesoamerica

The exact date of domestication is unknown; by 500 BC, it was already being cultivated in southern
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
and probably other areas. The
Pueblo Pueblo refers to the settlements of the Pueblo peoples, Native American tribes in the Southwestern United States, currently in New Mexico, Arizona, and Texas. The permanent communities, including some of the oldest continually occupied settlement ...
people believed that tomato seeds could confer powers of
divination Divination () is the attempt to gain insight into a question or situation by way of an occultic ritual or practice. Using various methods throughout history, diviners ascertain their interpretations of how a should proceed by reading signs, ...
. The large, lumpy variety of tomato, a mutation from a smoother, smaller fruit, originated in Mesoamerica, and may be the direct ancestor of some modern cultivated tomatoes. The Aztecs raised several varieties of tomato, with red tomatoes called and green tomatoes (physalis) called (
tomatillo The tomatillo (''Physalis philadelphica'' and ''Physalis ixocarpa''), also known as the Mexican husk tomato, is a plant of the nightshade family bearing small, spherical, and green or green-purple fruit. Tomatillos originated in Mexico and were ...
).
Bernardino de Sahagún Bernardino de Sahagún ( – 5 February 1590) was a Franciscan friar, missionary priest and pioneering ethnographer who participated in the Catholic evangelization of colonial New Spain (now Mexico). Born in Sahagún, Spain, in 1499, he jour ...
reported seeing a great variety of tomatoes in the Aztec market at Tenochtitlán (Mexico City): "large tomatoes, small tomatoes, leaf tomatoes, sweet tomatoes, large serpent tomatoes, nipple-shaped tomatoes", and tomatoes of all colors from the brightest red to the deepest yellow. Sahagún mentioned Aztecs cooking various sauces, some with tomatoes of different sizes, serving them in city markets: "foods sauces, hot sauces; ... with tomatoes, ... sauce of large tomatoes, sauce of ordinary tomatoes, ..."


Spanish distribution

The Spanish
conquistador Conquistadors (, ) or conquistadores (; ; ) were Spanish Empire, Spanish and Portuguese Empire, Portuguese colonizers who explored, traded with and colonized parts of the Americas, Africa, Oceania and Asia during the Age of Discovery. Sailing ...
Hernán Cortés Hernán Cortés de Monroy y Pizarro Altamirano, 1st Marquis of the Valley of Oaxaca (December 1485 – December 2, 1547) was a Spanish ''conquistador'' who led an expedition that caused the fall of the Aztec Empire and brought large portions o ...
's capture of Tenochtitlan in 1521 initiated the widespread cultural and biological interchange called the
Columbian exchange The Columbian exchange, also known as the Columbian interchange, was the widespread transfer of plants, animals, and diseases between the New World (the Americas) in the Western Hemisphere, and the Old World (Afro-Eurasia) in the Eastern Hemis ...
; certainly the tomato was being grown in Europe within a few years of that event. The earliest discussion of the tomato in European literature appeared in
Pietro Andrea Mattioli Pietro Andrea Gregorio Mattioli (; 12 March 1501– ) was a doctor and naturalist born in Siena. His most important work was a commentary on the medicinal plants of Pedanius Dioscorides first published in 1544 which was translated into several la ...
's 1544
herbal A herbal is a book containing the names and descriptions of plants, usually with information on their medicinal, Herbal tonic, tonic, culinary, toxic, hallucinatory, aromatic, or Magic (paranormal), magical powers, and the legends associated wi ...
. He suggested that a new type of
eggplant Eggplant (American English, US, Canadian English, CA, Australian English, AU, Philippine English, PH), aubergine (British English, UK, Hiberno English, IE, New Zealand English, NZ), brinjal (Indian English, IN, Singapore English, SG, Malays ...
had been brought to Italy. He stated that it was blood red or golden color when mature, and could be divided into segments and eaten like an eggplant—that is, cooked and seasoned with salt, black pepper, and oil. Ten years later Mattioli named the fruits in print as , or "golden apples". After the
Spanish colonization of the Americas The Spanish colonization of the Americas began in 1493 on the Caribbean island of Hispaniola (now Haiti and the Dominican Republic) after the initial 1492 voyage of Genoa, Genoese mariner Christopher Columbus under license from Queen Isabella ...
, the Spanish distributed the tomato throughout their colonies in the
Caribbean The Caribbean ( , ; ; ; ) is a region in the middle of the Americas centered around the Caribbean Sea in the Atlantic Ocean, North Atlantic Ocean, mostly overlapping with the West Indies. Bordered by North America to the north, Central America ...
. They brought it to the
Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
, from where it spread to
southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
and then the whole of Asia. The Spanish brought the tomato to Europe, where it grew easily in
Mediterranean climate A Mediterranean climate ( ), also called a dry summer climate, described by Köppen and Trewartha as ''Cs'', is a temperate climate type that occurs in the lower mid-latitudes (normally 30 to 44 north and south latitude). Such climates typic ...
s; cultivation began in the 1540s. It was probably eaten shortly after it was introduced, and was certainly being used as food by the early 17th century in Spain, as documented in the 1618 play ''La octava maravilla'' by
Lope de Vega Félix Lope de Vega y Carpio (; 25 November 156227 August 1635) was a Spanish playwright, poet, and novelist who was a key figure in the Spanish Golden Age (1492–1659) of Spanish Baroque literature, Baroque literature. In the literature of ...
with "lovelier than ... a tomato in season".


China

The tomato was introduced to China, likely via the Philippines or Macau, in the 16th century. It was given the name 番茄 (foreign eggplant), as the Chinese named many foodstuffs introduced from abroad, but referring specifically to early introductions.


Italy

In 1548, the house steward of Cosimo de' Medici, the grand duke of
Tuscany Tuscany ( ; ) is a Regions of Italy, region in central Italy with an area of about and a population of 3,660,834 inhabitants as of 2025. The capital city is Florence. Tuscany is known for its landscapes, history, artistic legacy, and its in ...
, wrote to the Medici private secretary informing him that the basket of tomatoes sent from the grand duke's Florentine estate at Torre del Gallo "had arrived safely". Tomatoes were grown mainly as ornamentals early on after their arrival in Italy. For example, the Florentine aristocrat Giovanvettorio Soderini wrote how they "were to be sought only for their beauty", and were grown only in gardens or flower beds. The tomato's ability to mutate and create new and different varieties helped contribute to its success and spread throughout Italy. However, in areas where the climate supported growing tomatoes, their habit of growing close to the ground suggested low status. They were not adopted as a staple of the peasant population because they were not as filling as other crops. Additionally, both toxic and inedible varieties discouraged many people from attempting to consume or prepare any other varieties. In certain areas of Italy, such as Florence, the fruit was used solely as a tabletop decoration, until it was incorporated into the local cuisine in the late 17th or early 18th century. The earliest discovered cookbook with tomato recipes was published in
Naples Naples ( ; ; ) is the Regions of Italy, regional capital of Campania and the third-largest city of Italy, after Rome and Milan, with a population of 908,082 within the city's administrative limits as of 2025, while its Metropolitan City of N ...
in 1692, though the author had apparently obtained these recipes from Spanish sources. Varieties were developed over the following centuries for drying, for sauce, for pizzas, and for long-term storage. These varieties are usually known for their place of origin as much as by a variety name. For example, there is the , the "hanging tomato of Vesuvius", and the well known and highly prized
San Marzano tomato San Marzano tomato is a variety of plum tomato originating in the Campania region of Italy. It is known for its flavour and quality as a canning tomato. San Marzano production is protected by a European protected designation of origin (PDO) cer ...
grown in that region, with a European
protected designation of origin The protected designation of origin (PDO) is a type of geographical indication of the European Union aimed at preserving the designations of origin of food-related products. The designation was created in 1992 and its main purpose is to designat ...
certification.


Britain

Tomatoes were not grown in England until the 1590s. One of the earliest cultivators was
John Gerard John Gerard (also John Gerarde, 1545–1612) was an English herbalist with a large garden in Holborn, now part of London. His 1,484-page illustrated ''Herball, or Generall Historie of Plantes'', first published in 1597, became a popular garde ...
, a
barber-surgeon The barber surgeon was one of the most common European medical practitioners of the Middle Ages, generally charged with caring for soldiers during and after battle. In this era, surgery was seldom conducted by physicians. Instead, barbers, who ...
. Gerard's ''Herbal'', published in 1597, and largely
plagiarized Plagiarism is the representation of another person's language, thoughts, ideas, or expressions as one's own original work.From the 1995 '' Random House Compact Unabridged Dictionary'': use or close imitation of the language and thoughts of anothe ...
from continental sources, is also one of the earliest discussions of the tomato in England. Gerard knew the tomato was eaten in Spain and Italy. Nonetheless, he believed it was poisonous. Gerard's views were influential, and the tomato was considered unfit for eating for many years in
Britain Britain most often refers to: * Great Britain, a large island comprising the countries of England, Scotland and Wales * The United Kingdom of Great Britain and Northern Ireland, a sovereign state in Europe comprising Great Britain and the north-eas ...
and its North American colonies. By 1820, tomatoes were described as "to be seen in great abundance in all our vegetable markets" and to be "used by all our best cooks", reference was made to their cultivation in gardens still "for the singularity of their appearance", while their use in cooking was associated with exotic Italian or Jewish cuisine. For example, in
Elizabeth Blackwell Elizabeth Blackwell (3 February 182131 May 1910) was an English-American physician, notable as the first woman to earn a medical degree in the United States, and the first woman on the Medical Register of the General Medical Council for the Un ...
's ''A Curious Herbal'', it is described under the name "Love Apple ()" as being consumed with oil and vinegar in Italy, similar to consumption of cucumbers in the UK. In 1963, ''
The New York Times ''The New York Times'' (''NYT'') is an American daily newspaper based in New York City. ''The New York Times'' covers domestic, national, and international news, and publishes opinion pieces, investigative reports, and reviews. As one of ...
'' gave an explanation of the name 'Love Apple' as a French misreading of the Italian ("the Moors' apple") as , ("apple of love").


Middle East

The tomato was introduced to cultivation in the Middle East by John Barker, British consul in
Aleppo Aleppo is a city in Syria, which serves as the capital of the Aleppo Governorate, the most populous Governorates of Syria, governorate of Syria. With an estimated population of 2,098,000 residents it is Syria's largest city by urban area, and ...
. Nineteenth century descriptions of its consumption are uniformly as an ingredient in a cooked dish. In 1881, it is described as only eaten in the region "within the last forty years".


United States

The earliest reference to tomatoes being grown in
British North America British North America comprised the colonial territories of the British Empire in North America from 1783 onwards. English colonisation of North America began in the 16th century in Newfoundland, then further south at Roanoke and Jamestown, ...
is from 1710, when herbalist
William Salmon William Salmon (2 June 1644–1713) was an English empiric doctor and a writer of medical texts. He advertised himself as a "Professor of Physick". Salmon held an equivocal place in the medical community. He led apothecaries in opposing atte ...
saw them in what is today
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
, perhaps introduced from the Caribbean. By the mid-18th century, they were cultivated on some Carolina plantations, and probably in other parts of the Southeast.
Thomas Jefferson Thomas Jefferson (, 1743July 4, 1826) was an American Founding Fathers of the United States, Founding Father and the third president of the United States from 1801 to 1809. He was the primary author of the United States Declaration of Indepe ...
, who ate tomatoes in Paris, sent some seeds back to America. Some early American advocates of the culinary use of the tomato included Michele Felice Cornè and
Robert Gibbon Johnson Robert Gibbon Johnson (July 23, 1771 – October 2, 1850), also known as Colonel Johnson, was an American gentleman farmer, historian, horticulturalist, judge, soldier, and statesman who lived in Salem, New Jersey. He is especially renowned for ...
. Many Americans considered tomatoes to be poisonous at this time and, in general, they were grown more as
ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
s than as food. In 1897, W. H. Garrison stated, "The belief was once transmitted that the tomato was sinisterly dangerous." He recalled in his youth tomatoes were dubbed "love-apples or wolf-apples" and shunned as "globes of the devil". When Alexander W. Livingston (1821–1898) began developing the tomato as a commercial crop, his aim had been to grow tomatoes smooth in contour, uniform in size, and sweet in flavor. He eventually developed over seventeen varieties. The U.S. Department of Agriculture's 1937 yearbook declared that "half of the major varieties were a result of the abilities of the Livingstons to evaluate and perpetuate superior material in the tomato." Livingston's first breed of tomato, the Paragon, was introduced in 1870. In 1875, he introduced the Acme, said to be in the parentage of most cultivars for the next twenty-five years. Other early breeders included Henry Tilden in Iowa and a Dr. Hand in Baltimore.Boswell, Victor R.
Improvement and Genetics of Tomatoes, Peppers, and Eggplant
" ''Yearbook of Agriculture, 1937,'' p. 179. U.S. Department of Agriculture. Accessed 25 May 2018/
Because of the tomato's need for heat and a long growing season, several states in the
Sun Belt The Sun Belt is a region of the United States generally considered stretching across the Southeast and Southwest. Another rough definition of the region is the area south of the Parallel 36°30′ north. Several climates can be found in the re ...
became major producers, particularly
Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...
and
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
. In California, tomatoes are grown under
irrigation Irrigation (also referred to as watering of plants) is the practice of applying controlled amounts of water to land to help grow crops, landscape plants, and lawns. Irrigation has been a key aspect of agriculture for over 5,000 years and has bee ...
for both the fresh market and for
canning Canning is a method of food preservation in which food is processed and sealed in an airtight container (jars like Mason jars, and steel and tin cans). Canning provides a shelf life that typically ranges from one to five years, although under ...
and processing. The
University of California, Davis The University of California, Davis (UC Davis, UCD, or Davis) is a Public university, public Land-grant university, land-grant research university in Davis, California, United States. It is the northernmost of the ten campuses of the University ...
's C.M. Rick Tomato Genetics Resource Center maintains a gene bank of wild relatives, monogenic mutants and genetic stocks. Research on processing tomatoes is also conducted by the California Tomato Research Institute in
Escalon, California Escalon (Spanish: ''Escalón'', meaning "Step") is a city in San Joaquin County, California, United States. The population was 7,472 at the 2020 census, up from 7,132 at the 2010 census and 5,963 at the 2000 census. Escalon is a Spanish word ...
. In California, growers have used a method of cultivation called dry-farming, especially with Early Girl tomatoes. This technique encourages the plant to send roots deep to find existing moisture.


Botany


Description

Tomato plants are vines, becoming
decumbent This glossary of botanical terms is a list of definitions of terms and concepts relevant to botany and plants in general. Terms of plant morphology are included here as well as at the more specific Glossary of plant morphology and Glossary ...
, and can grow up to ; bush varieties are generally no more than tall. They are tender perennials, often grown as annuals. Tomato plants are
dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, ...
s. They grow as a series of branching stems, with a terminal bud at the tip that does the actual growing. When the tip eventually stops growing, whether because of pruning or flowering, lateral buds take over and grow into new, fully functional, vines. Tomato vines are typically pubescent, meaning covered with fine short hairs. The hairs facilitate the vining process, turning into roots wherever the plant is in contact with the ground and moisture, especially if the vine's connection to its original root has been damaged or severed. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are long, odd
pinnate Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and ...
, with five to nine leaflets on petioles, each leaflet up to long, with a serrated margin; both the stem and leaves are densely glandular-hairy. Tomato flowers are bisexual and are able to self fertilize. As tomatoes were moved from their native areas, their traditional
pollinator A pollinator is an animal that moves pollen from the male anther of a flower to the female carpel, stigma of a flower. This helps to bring about fertilization of the ovules in the flower by the male gametes from the pollen grains. Insects are ...
s (probably a species of halictid bee) did not move with them. The trait of self-fertility became an advantage, and domestic
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s of tomato have been selected to maximize this trait. This is not the same as
self-pollination Self-pollination is a form of pollination in which pollen arrives at the stigma of a flower (in flowering plants) or at the ovule (in gymnosperms) of the same plant. The term cross-pollination is used for the opposite case, where pollen from ...
, despite the common claim that tomatoes do so. That tomatoes pollinate themselves poorly without outside aid is clearly shown in
greenhouse A greenhouse is a structure that is designed to regulate the temperature and humidity of the environment inside. There are different types of greenhouses, but they all have large areas covered with transparent materials that let sunlight pass an ...
situations, where pollination must be aided by artificial wind, vibration of the plants, or by cultured
bumblebee A bumblebee (or bumble bee, bumble-bee, or humble-bee) is any of over 250 species in the genus ''Bombus'', part of Apidae, one of the bee families. This genus is the only Extant taxon, extant group in the tribe Bombini, though a few extinct r ...
s. The flowers develop on the apical
meristem In cell biology, the meristem is a structure composed of specialized tissue found in plants, consisting of stem cells, known as meristematic cells, which are undifferentiated cells capable of continuous cellular division. These meristematic c ...
. They have the
anther The stamen (: stamina or stamens) is a part consisting of the male reproductive organs of a flower. Collectively, the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament ...
s fused along the edges, forming a column surrounding the
pistil Gynoecium (; ; : gynoecia) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl (botany), whorl of a flower; it consists ...
's style. The anthers bend into a cone-like structure, surrounding the stigma. The flowers are across, yellow, with five pointed lobes on the corolla; they are borne in a cyme of three to twelve together. The fruit develops from the ovary of the plant after fertilization, its flesh comprising the pericarp walls. The fruit contains
locule A locule (: locules) or loculus (; : loculi) is a small cavity or compartment within an organ or part of an organism (animal, plant, or fungus). In angiosperms (flowering plants), the term ''locule'' usually refers to a chamber within an ovary ...
s, hollow spaces full of seeds. These vary among cultivated varieties. Some smaller varieties have two locules; globe-shaped varieties typically have three to five; beefsteak tomatoes have a great number of small locules; and
plum tomato A plum tomato, also known in the United States as a processing tomato or paste tomato, is a type of tomato cultivated for sauce and packing purposes. It is generally oval or cylindrical in shape, with significantly fewer locules (seed compartment ...
es have very few, very small locules. For propagation, the seeds need to come from a mature fruit, and must be lightly fermented to remove the gelatinous outer coating and then dried before use. The tomato has a mutualistic relationship with
arbuscular mycorrhiza An arbuscular mycorrhiza (AM) (plural ''mycorrhizae'') is a type of mycorrhiza in which the symbiont fungus (''Arbuscular mycorrhizal fungi'', or AMF) penetrates the cortical cells of the roots of a vascular plant forming arbuscules. Arbuscul ...
l fungi such as ''
Rhizophagus irregularis ''Rhizophagus irregularis'' (previously known as ''Glomus intraradices'') is an arbuscular mycorrhizal fungus used as a soil inoculant in agriculture and horticulture. ''Rhizophagus irregularis'' is also commonly used in scientific studies of the ...
''. Scientists use the tomato as a
model species A model organism is a non-human species that is extensively studied to understand particular biological phenomena, with the expectation that discoveries made in the model organism will provide insight into the workings of other organisms. Mode ...
for investigating such symbioses. File:Germinating tomatos.jpg , Seedlings 7 days after planting File:Tomato 27 days from planting seeds.jpg , 27 days after planting File:Solanum lycopersicum - Flor tomaca 057.jpg, Flower File:Tomato fruit and flowers at day 52.jpg , 52-day-old plant, first fruits File:Green Tomato.jpg, Unripe fruit on the vine


Phylogeny

Like the
potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...
, tomatoes belong to the genus ''
Solanum ''Solanum'' is a large and diverse genus of flowering plants, which include three food crops of high economic importance: the potato, the tomato and the eggplant (aubergine, brinjal). It is the largest genus in the nightshade family Solana ...
'', which is a member of the nightshade family, the
Solanaceae Solanaceae (), commonly known as the nightshades, is a family of flowering plants in the order Solanales. It contains approximately 2,700 species, several of which are used as agricultural crops, medicinal plants, and ornamental plants. Many me ...
. That is a diverse family of flowering plants, often poisonous, that includes the mandrake (''Mandragora (genus), Mandragora''), deadly nightshade (''Atropa''), and tobacco (''Nicotiana''), as shown in the outline phylogenetic tree (many branches omitted).


Taxonomy

In 1753, Carl Linnaeus, Linnaeus placed the tomato in the genus ''Solanum'' (alongside the potato) as ''Solanum lycopersicum''. In 1768, Philip Miller moved it to its own genus, naming it ''Lycopersicon esculentum''. The name came into wide use, but was technically in breach of the International Code of Botanical Nomenclature, plant naming rules because Linnaeus's species name ''lycopersicum'' still had priority. Although the name ''Lycopersicum lycopersicum'' was suggested by Karsten (1888), it is not used because it violates the International Code of Nomenclature barring the use of tautonyms in botanical nomenclature. The corrected name ''Lycopersicon lycopersicum'' (Nicolson 1974) was technically valid, because Miller's genus name and Linnaeus's species name differ in exact spelling. As ''Lycopersicon esculentum'' has become so well known, it was officially listed as a ''nomen conservandum'' in 1983, and would be the correct name for the tomato in classifications which do not place the tomato in the genus ''Solanum''. Genetics, Genetic evidence shows that Linnaeus was correct to put the tomato in the genus ''Solanum'', making ''S. lycopersicum'' the correct name. Both names, however, will probably be found in the literature for some time. Two of the major reasons for considering the genera separate are the leaf structure (tomato leaves are markedly different from any other ''Solanum''), and the biochemistry (many of the alkaloids common to other ''Solanum'' species are conspicuously absent from the tomato). On the other hand, hybrids of tomato and diploid
potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...
can be created in the lab by somatic fusion, and are partially fertile, providing evidence of the close relationship between these species.


Plant breeding


Genetics

An international consortium of researchers from 10 countries began sequencing the tomato genome in 2004. A prerelease version of the genome was made available in December 2009. The complete genome for the cultivar Heinz 1706 was published on 31 May 2012 in ''Nature (journal), Nature''. The latest reference genome published in 2021 had 799 MB and encodes 34,384 (predicted) proteins, spread over 12 chromosomes. The first commercially available genetically modified food was Genetically modified tomato, a tomato called Flavr Savr, which was engineered to have a longer shelf life. It could be vine ripened without compromising shelf life. However, the product was not commercially successful, and was sold only until 1997.


Breeding of modern commercial varieties

The poor taste and lack of sugar in modern garden and commercial tomato varieties resulted from breeding tomatoes to ripen uniformly red. This change occurred after discovery of a mutant "u" phenotype in the mid-20th century, so named because the fruits ripened uniformly. This was widely cross-bred to produce red fruit without the typical green ring around the stem on un-crossbred varieties. Before this, most tomatoes produced more sugar during ripening, and were sweeter and more flavorful. 10–20% of the total Biological carbon fixation, carbon fixed in the fruit can be produced by photosynthesis in the developing fruit of the normal U phenotype. The u mutation encodes a factor that produces defective chloroplasts with lower density in developing fruit, making them a lighter green, and reducing sugar in the resulting ripe fruit by 10–15%. Perhaps more importantly, the fruit chloroplasts are remodelled during ripening into chlorophyll-free chromoplasts that synthesize and accumulate the carotenoids lycopene, beta-Carotene, β-carotene, and other metabolites that are sensory and nutritional assets of the ripe fruit. The potent chloroplasts in the dark-green shoulders of the "U" phenotype are beneficial here, but have the disadvantage of leaving green shoulders near the stems of the ripe fruit, and even cracked yellow shoulders. This is apparently because of oxidative stress due to overload of the photosynthetic chain in direct sunlight at high temperatures. Hence, genetic design of a commercial variety that combines the advantages of types "u" and "U" requires fine tuning, but may be feasible. Breeders strive to produce tomato plants with improved yield, shelf life, size, and resistance to environmental pressures, including disease. These efforts have yielded unintended negative consequences on various fruit attributes. For instance, linkage drag, the introduction of an undesired trait during backcrossing, has altered the metabolism of the fruit. This trait is physically close to the desired allele along the chromosome. Breeding for traits like larger fruit has thus unintentionally altered nutritional value and flavor. Breeders have turned to wild tomato species as a source of alleles to introduce beneficial traits into modern varieties. For example, wild relatives may possess higher amounts of fruit solids (associated with greater sugar content), or resistance to diseases such as to the early blight pathogen ''Alternaria solani''. However, this tactic has limitations, since selection for traits such as pathogen resistance can negatively impact other favorable traits such as fruit production.


Cultivation

The tomato is grown worldwide for its edible fruits, with thousands of
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s. File:Tomatoes in a market in France.jpg, Heirloom varieties in a French market File:Heirlooms.jpg, Heirloom cultivars Brandywine (tomato), Brandywine (biggest red), Black Krim (tomato), Black Krim (lower left) and Green Zebra (top left) File:Tomatoes for sale in France.jpg, Modern varieties in France File:Yellow cherry tomatoes.jpg, Yellow cherry tomatoes


Hydroponic and greenhouse cultivation

Greenhouse tomato production in large-acreage commercial greenhouses and owner-operator stand-alone or multiple-bay greenhouses is increasing, providing fruit during those times of the year when field-grown fruit is not readily available. Smaller fruit (cherry and grape), or cluster tomatoes (fruit-on-the-vine) are the fruit of choice for the large commercial greenhouse operators while the beefsteak varieties are the choice of owner-operator growers. Tomatoes are also grown using hydroponics. File:Tomato flower and young fruit.jpg, Flower and young fruit File:Tomato scanned.jpg, Flowers and ripe fruit can be present simultaneously. File:Tomato P5260299b.jpg, Hydroponic cultivation File:Greenhouses near El Ejido.jpg, Greenhouse cultivation in Andalusia


Picking and ripening

To facilitate transportation and storage, tomatoes are often picked unripe (green) and ripened in storage with ethylene as a plant hormone, the plant hormone ethylene. At industrial scale, such as for canning, tomatoes are picked mechanically. The machine cuts the whole vine and uses sensors to separate ripe tomatoes from the rest of the plant, which is returned to the farm for use either as green manure or to be grazed by livestock.


Production

In 2022, world production of tomatoes was 186 million tonnes, with China accounting for 37% of the total, followed by India, Turkey, and the United States as major producers (table). The world dedicated 4.8 million hectares in 2012 for tomato cultivation and the total production was about 161.8 million tonnes. The average world farm yield for tomato was 33.6 tonnes per hectare in 2012. Tomato farms in the Netherlands were the most productive in 2012, with a nationwide average of 476 tonnes per hectare, followed by Belgium (463 tonnes per hectare) and Iceland (429 tonnes per hectare)."FAOSTAT: Production-Crops, 2012 data"
''Food and Agriculture Organization of the United Nations'', August 2014


Pests and diseases


Pests

Common tomato pests include the Engytatus modestus, tomato bug, Brown marmorated stink bug, stink bugs, cutworms, tomato hornworms and tobacco hornworms, aphids, cabbage loopers, whitefly, whiteflies, tomato fruitworms, flea beetles, red spider mite, ''Tuta absoluta'' (tomato leafminer), slugs, and Colorado potato beetles. The tomato russet mite, ''Aculops lycopersici'', feeds on foliage and young fruit of tomato plants, causing shrivelling and necrosis of leaves, flowers, and fruit, possibly killing the plant. After an insect attack tomato plants produce systemin, a plant peptide hormone. This activates defensive mechanisms, such as the production of protease inhibitor (biology), protease inhibitors to slow the growth of insects. The hormone was first identified in tomatoes. File:Engytatus modestus closeup 2.jpg, Engytatus modestus, Tomato bug feeding on plant sap File:Tomato fruitworm.jpg, Tomato fruitworm feeding on unripe fruit File:Tomato hornworm.jpg, Tomato hornworm larva on stem File:20230811 Aculops lycopersici 08 D.jpg, Aculops lycopersici, Tomato russet mites on greenhouse plant


Diseases

Tomato cultivars vary widely in their resistance to disease. Modern Hybrid (biology), hybrids focus on improving disease resistance over the heirloom plants. A common tomato disease is tobacco mosaic virus. Handling cigarettes and other infected tobacco products can transmit the virus to tomato plants. A serious disease is curly top, carried by the beet leafhopper, which interrupts the lifecycle. As the name implies, it has the symptom of making the top leaves of the plant wrinkle up and grow abnormally. Tomato bacterial wilt, Bacterial wilt is another common disease impacting yield. Wang ''et al.'', 2019 found phage therapy, phage combination therapies to reduce the impact of bacterial wilt, sometimes by reducing bacterial abundance and sometimes by selection (biology), selecting for phage resistance, resistant but slow growing genetics. File:Tomato with Phytophthora infestans (late blight).jpg, Late blight, caused by the oomycete ''Phytophthora infestans'' File:Ralstonis wilt symptom.jpg, Wilt caused by the Bacteria, bacterium
''Ralstonia solanacearum'' File:Tomaquera amb Fusarium HV.JPG, Wilt caused by ''Fusarium oxysporum'' File:Tomato (Solanum lycopersicum)- Root knot nematodes - 27421750599.jpg, Nematode root-knot caused by ''Meloidogyne incognita''


As food


Culinary

Tomatoes, with their
umami Umami ( from ), or savoriness, is one of the five basic tastes. It is characteristic of broths and cooked meats. People taste umami through taste receptors that typically respond to glutamates and nucleotides, which are widely present in me ...
flavor, are extensively used in Mediterranean cuisine as a key ingredient in
pizza Pizza is an Italian cuisine, Italian, specifically Neapolitan cuisine, Neapolitan, dish typically consisting of a flat base of Leavening agent, leavened wheat-based dough topped with tomato, cheese, and other ingredients, baked at a high t ...
and many pasta sauces. Tomatoes are used in Spanish cuisine, Spanish
gazpacho Gazpacho () or gaspacho (), also called Andalusian gazpacho (from Spanish ''gazpacho andaluz''), is a cold soup and drink made of raw, blended vegetables. It originated in the southern regions of the Iberian Peninsula and spread into other are ...
and Catalan cuisine, Catalan . The tomato is a crucial and ubiquitous part of Middle Eastern cuisine, served fresh in salads (e.g., Arab salad, Israeli salad, Shirazi salad and Çoban salatası, Turkish salad), grilled with kebabs and other dishes, made into sauces, and so on. Tomatoes were gradually incorporated into Indian curry dishes after Europeans introduced them. A Kashmiri cuisine, Kashmiri curry,
rogan josh Rogan josh ( English: /ˌroʊɡən ˈdʒɑʃ/);Rogan Josh
Oxford English Dictionary
), also spell ...
, often contains tomato; it may originally have been colored red with chili pepper, and tomatoes may characterize the Punjabi cuisine, Punjabi version of the dish. The modern British curry tikka masala often has a tomato and cream sauce. File:Tomato soup with bread.jpg, Tomato soup with croutons File:Rogan josh02.jpg, Rogan josh, a curry often made with tomatoes Bloody Mary Coctail with celery stalk - Evan Swigart.jpg, Bloody Mary (cocktail), Bloody Mary, a tomato cocktail File:Pan tumaca cortado (cropped).jpg, , Catalan tomato bread File:Tomates farcies végétariennes.jpg, Tomatoes stuffed with egg and Parmigiano-Reggiano, Parmesan cheese


Storage

Tomatoes keep best unwashed at room temperature and out of direct sunlight, rather than in a refrigerator. Storing stem down can prolong shelf life. Unripe tomatoes can be kept in a paper bag to ripen. Tomatoes can be preserved by canning, freezing, drying, or cooking down to a paste or puree.


Nutrition

A raw tomato is 95% water, 4% carbohydrates, and less than 1% each of fat and protein (nutrient), protein (table). In a reference amount of , raw tomatoes supply 18 calories and 16% of the Daily Value of vitamin C, but otherwise have low micronutrient content (table).


Effects on health

The US Food and Drug Administration has determined there is little credible evidence that tomatoes or tomato-based foods reduce the risk of various types of cancer. In a 2011 scientific review, the European Food Safety Authority concluded that lycopene did not favorably influence DNA, skin exposed to ultraviolet radiation, heart function or vision.


Toxins

The leaves, stem, and green unripe fruit of the tomato plant contain small amounts of the alkaloid tomatine. They contain small amounts of solanine, a toxic alkaloid found in larger amounts in
potato The potato () is a starchy tuberous vegetable native to the Americas that is consumed as a staple food in many parts of the world. Potatoes are underground stem tubers of the plant ''Solanum tuberosum'', a perennial in the nightshade famil ...
leaves and other members of the
nightshade family Solanaceae (), commonly known as the nightshades, is a family of flowering plants in the order Solanales. It contains approximately 2,700 species, several of which are used as agricultural crops, medicinal plants, and ornamental plants. Many me ...
. Tomato plants can be toxic to dogs if they eat large amounts of the fruit, or chew plant material. Small amounts of tomato foliage are sometimes used for flavoring, and the green fruit of unripe red tomato varieties is sometimes used for cooking, particularly as fried green tomatoes.


''Salmonella'' outbreaks

Tomatoes have been linked to multiple ''Salmonella'' food poisoning outbreaks in the US. 2008 United States salmonellosis outbreak, One in 2008 caused the temporary removal of tomatoes from stores and restaurants across the United States and parts of Canada. In 2022 and 2023, an outbreak of Salmonella Senftenberg ST14 affected the US and 12 countries in Europe.


In popular culture


Celebrations

A massive "tomato tree" in the Walt Disney World Resort's experimental greenhouses in Lake Buena Vista, Florida may have been the largest single tomato plant. It yielded a harvest of more than 32,000 tomatoes, together weighing .The country's only single vine "tomato tree" growing in The Land pavilion at Epcot
Walt Disney World News
The town of
Buñol Buñol is a town and municipality in the province of Valencia, Spain. The municipality has an area of some 112 km2, and is situated approximately 38 km west of the provincial and autonomous community capital city, Valencia. It lies alon ...
, Spain, annually celebrates Tomatina, La Tomatina, a festival centered on an enormous tomato fight. On 30 August 2007, as many as 40,000 Spaniards gathered to throw of tomatoes at each other in the festival. Some US states have adopted the tomato as a state fruit or vegetable. Arkansas took both sides by declaring the ''South Arkansas Vine Ripe Pink Tomato'' both the state fruit and the state vegetable in the same law, citing both its culinary and botanical classifications. In 2009, the state of Ohio passed a law making the tomato the state's official fruit, while tomato juice has been the state's official beverage since 1965. Livingston's plant breeding is commemorated in his home town of Reynoldsburg, Ohio, Reynoldsburg with an annual Tomato Festival; it calls itself "The Birthplace of the Tomato".About Reynoldsburg
ci.reynoldsburg.oh.us
In Finland, the ''Tomatkarnevalen'' is held annually in the town of
Närpes Närpes (Finland Swedish: ; ) is a town in Finland, located on the west coast of the country. Närpes is situated in Ostrobothnia, along the Gulf of Bothnia. The population of Närpes is approximately , while the sub-region has a population of ...
. Tomatoes are sometimes thrown in public protests. Embracing it for this connotation, the Socialist Party (Netherlands), Dutch Socialist party adopted the tomato as their logo. The same meaning is evoked in the name of the American review aggregator, review-aggregation website for film and television, "Rotten Tomatoes", though its founder mentions a scene in the 1992 movie ''Léolo, Leolo'' as the immediate source of the name. File:Arrojando tomates desde un camión - La Tomatina 2010.jpg, Throwing tomatoes from a truck during the Spanish ''La Tomatina, Tomatina'' festival File:Tomatkarnevalen i Närpes 1993c.jpg, (The Tomato Carnival) in
Närpes Närpes (Finland Swedish: ; ) is a town in Finland, located on the west coast of the country. Närpes is situated in Ostrobothnia, along the Gulf of Bothnia. The population of Närpes is approximately , while the sub-region has a population of ...
, Finland, in 1993


Fruit or vegetable

Although the tomato is List of culinary fruits, cooked and eaten as a vegetable, Fruit#Botanic fruit and culinary fruit, botanically, a tomato is a fruit, specifically a
berry A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit although many pips or seeds may be present. Common examples of berries in the cul ...
, consisting of the ovary (plants), ovary, together with its seeds, of a flowering plant. The issue has led to legal dispute in the United States. In 1887, U.S. tariff laws that imposed a Duty (economics), duty on vegetables, but not on fruit, caused the tomato's status to become a matter of legal importance. In ''Nix v. Hedden'', the Supreme Court of the United States, U.S. Supreme Court settled the controversy on 10 May 1893, by declaring that for the purposes of the Tariff of 1883 only, the tomato is a vegetable, based on the popular definition that classifies vegetables by use—they are generally served with dinner and not dessert..


See also

* La Tomatina, world's largest tomato food fight * List of countries by tomato production * List of tomato dishes * Marglobe, an early attempt at breeding a disease-resistant tomato * Ring culture * Physalis, a similar fruit also used in cooking * Tomato effect * Tomato jam * Nightshades ** Potato ** Eggplant ** Tomatillo, a similar fruit from the related genus


References


Sources

* *


Further reading

* David Gentilcore. ''Pomodoro! A History of the Tomato in Italy'' (Columbia University Press, 2010), scholarly history *


External links

* * * * *
Tomato Genome Sequencing Project
– Sequencing of the twelve tomato chromosomes.
Tomato core collection database
– Phenotypes and images of 7,000 tomato cultivars {{Authority control Tomatoes, Tomatoes Crops originating from Mexico Crops originating from indigenous Americans Crops originating from South America Fruit vegetables Fruits originating in North America Fruits originating in South America Plants described in 1753 Solanum