This timeline of ichthyosaur research is a chronological listing of events in the
history
History (derived ) is the systematic study and the documentation of the human activity. The time period of event before the History of writing#Inventions of writing, invention of writing systems is considered prehistory. "History" is an umbr ...
of
paleontology
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fos ...
focused on the ichthyosauromorphs, a group of secondarily aquatic marine reptiles whose later members superficially resembled dolphins, sharks, or swordfish. Scientists have documented ichthyosaur fossils at least as far back as the late 17th century. At that time, a scholar named
Edward Lhwyd
Edward Lhuyd FRS (; occasionally written Llwyd in line with modern Welsh orthography, 1660 – 30 June 1709) was a Welsh naturalist, botanist, linguist, geographer and antiquary. He is also named in a Latinate form as Eduardus Luidius.
Lif ...
published a book on British fossils that misattributed some ichthyosaur vertebrae to actual fishes; their true nature was not recognized until the 19th century. In
1811
Events
January–March
* January 8 – An unsuccessful slave revolt is led by Charles Deslondes, in St. Charles and St. James Parishes, Louisiana.
* January 17 – Mexican War of Independence – Battle of Calderón B ...
, a boy named Joseph Anning discovered the first ichthyosaur fossils that would come to be scientifically recognized as such.His sister Mary would later find the rest of its skeleton and would go on to become a respected fossil collector and paleontologist in her own right.
Early researchers recognized ichthyosaurs as marine reptiles, but major aspects of their anatomy and behavior needed to be resolved. They were frequently portrayed as leaving the water to bask on rocks and with straight tails. Although a bend in ichthyosaurs' tail vertebrae was seen from the earliest specimens, scholars assumed the bend reflected damage incurred to the animal's carcass after death. This bend was so common, however, that scholars eventually realized that it was natural and supported a shark-like tail fin. Scientists came to realize that ichthyosaurs were too adapted to leave the water even to lay eggs. Evidence for live birth in ichthyosaurs dates back as far as 1846, when Chaning Pierce reported an apparent fossil '' Ichthyosaurus''
embryo
An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sperm ...
to Sir Richard Owen.
Ichthyosaur discoveries continued to be made into the 20th century. In 1928, Simeon Muller discovered the remains of 40 gigantic ichthyosaurs in
Nevada
Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
. However, these remains would not be excavated until Charles Camp and Samuel Welles of Berkeley led an expedition for the purpose in the mid 1950s. These fossils would take more than a decade to excavate, and the results of Camp's examination of the bones would not be published until a year after his
1975
It was also declared the ''International Women's Year'' by the United Nations and the European Architectural Heritage Year by the Council of Europe.
Events
January
* January 1 - Watergate scandal (United States): John N. Mitchell, H. R. ...
death. These giant ichthyosaurs were named '' Shonisaurus popularis'' and their final resting place is now known as Berlin-Ichthyosaur state park.
Other notable late 20th century advances in ichthyosaur research include the recognition of a new genus of ichthyosaur called '' Eurhinosaurus longirostris'' that had been misclassified as a species of ''Ichthyosaurus'' since 1854. In 1986
Christopher McGowan
Christopher is the English version of a Europe-wide name derived from the Greek name Χριστόφορος (''Christophoros'' or '' Christoforos''). The constituent parts are Χριστός (''Christós''), "Christ" or "Anointed", and φέρει� ...
would describe another, similar animal serendipitously discovered in England as ''
Excalibosaurus
''Excalibosaurus'' (meaning "Excalibur's lizard") is a monotypic genus of marine prehistoric reptiles ( ichthyosaurs) that lived during the Sinemurian stage (approximately 196.5 ± 2 Ma to 190.8 ± 1.0 Ma (million years ago)) of the Early Jura ...
sword
A sword is an edged, bladed weapon intended for manual cutting or thrusting. Its blade, longer than a knife or dagger, is attached to a hilt and can be straight or curved. A thrusting sword tends to have a straighter blade with a pointed t ...
21st century
The 21st (twenty-first) century is the current century in the ''Anno Domini'' era or Common Era, under the Gregorian calendar. It began on 1 January 2001 (2001, MMI) and will end on 31 December 2100 (Roman numerals, MMC).
Marking the beginnin ...
would see scholarly debate regarding the cause of the ichthyosaurs' extinction, especially regarding the potential role played by competition with the mosasaurs which had evolved around the time.
Edward Lhwyd
Edward Lhuyd FRS (; occasionally written Llwyd in line with modern Welsh orthography, 1660 – 30 June 1709) was a Welsh naturalist, botanist, linguist, geographer and antiquary. He is also named in a Latinate form as Eduardus Luidius.
Lif ...
Oxford
Oxford () is a city in England. It is the county town and only city of Oxfordshire. In 2020, its population was estimated at 151,584. It is north-west of London, south-east of Birmingham and north-east of Bristol. The city is home to the Un ...
's Ashmolean Museum. The book contained illustrations of unrecognized ichthyosaur vertebrae misidentified as belonging to fishes. 120 copies of the book were produced.
Joseph Anning
Mary Anning (21 May 1799 – 9 March 1847) was an English fossil collector, dealer, and palaeontologist who became known around the world for the discoveries she made in Jurassic marine fossil beds in the cliffs along the English Channel ...
discovered the first scientifically recognized ichthyosaur remains.1812
*
Mary Anning
Mary Anning (21 May 1799 – 9 March 1847) was an English fossil collector, dealer, and palaeontologist who became known around the world for the discoveries she made in Jurassic marine fossil beds in the cliffs along the English Channe ...
found the rest of the original ichthyosaur skeleton discovered by her brother Joseph.1814
* An anatomist with the
Royal College of Surgeons
The Royal College of Surgeons is an ancient college (a form of corporation) established in England to regulate the activity of surgeons. Derivative organisations survive in many present and former members of the Commonwealth. These organisations ar ...
named
Sir Everard Home
Sir Everard Home, 1st Baronet, FRS (6 May 1756, in Kingston upon Hull – 31 August 1832, in London) was a British surgeon.
Home was born in Kingston-upon-Hull and educated at Westminster School. He gained a scholarship to Trinity College, C ...
described the ichthyosaur discovered by the Annings. He thought it formed a "link between fishes and crocodiles".1818
* König described the new genus '' Ichthyosaurus''.
1819
* Everard Home changed his mind about the relationships of '' Ichthyosaurus''. Instead of linking fishes with crocodiles, he concluded that it linked salamanders and lizards. He tried to rename it Proteosaurus after the salamander genus ''
Proteus
In Greek mythology, Proteus (; Ancient Greek: Πρωτεύς, ''Prōteus'') is an early prophetic sea-god or god of rivers and oceanic bodies of water, one of several deities whom Homer calls the " Old Man of the Sea" ''(hálios gérôn)''. ...
'', but his original name had priority and is still considered official.
1820s
1822
* Conybeare described ''Ichthyosaurus communis'' and '' I. intermedius'' along with the species that would later come to be known as '' Leptonectes tenuirostris'' and '' Temnodontosaurus platyodon''.
1823
* Mary Anning or a member of her family discovered a complete ''Ichthyosaurus'' skeleton in
Lyme Regis
Lyme Regis is a town in west Dorset, England, west of Dorchester and east of Exeter. Sometimes dubbed the "Pearl of Dorset", it lies by the English Channel at the Dorset–Devon border. It has noted fossils in cliffs and beaches on the Heri ...
. The find aroused widespread curiosity about ancient life throughout Britain.1824
* The
Reverend George Young
The Reverend is an honorific style most often placed before the names of Christian clergy and ministers. There are sometimes differences in the way the style is used in different countries and church traditions. ''The Reverend'' is correctly ...
Yorkshire
Yorkshire ( ; abbreviated Yorks), formally known as the County of York, is a historic county in northern England and by far the largest in the United Kingdom. Because of its large area in comparison with other English counties, functions have ...
. He also described the specimen, noting that it shared traits with crocodiles, fishes, and dolphins. Young speculated that ichthyosaurs might still be discovered alive as more of the world's seas and oceans are explored.
1830s
1830
* Henry de la Beche illustrated a work titled "'' Duria Antiquior''", meaning "Ancient
Dorset
Dorset ( ; archaically: Dorsetshire , ) is a county in South West England on the English Channel coast. The ceremonial county comprises the unitary authority areas of Bournemouth, Christchurch and Poole and Dorset. Covering an area of ...
" for fossil hunter Mary Anning. This work, which prominently features plesiosaurs, has been regarded as the first attempt to accurately reconstruct the
Mesozoic
The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ...
Thomas W. Hawkins
Thomas W. Hawkins Jr. (born 10 January 1938 in Flushing, New York) is an American historian of mathematics.
Hawkins defended his Ph.D. thesis on ''"The Origins and Early Development of Lebesgue's Theory of Integration"'' at the University of ...
published a book titled ''
Memoirs of Ichthyosauri and Plesiosauri
A memoir (; , ) is any nonfiction narrative writing based in the author's personal memories. The assertions made in the work are thus understood to be factual. While memoir has historically been defined as a subcategory of biography or autobiog ...
''. He sold it in subscription format for L2.10, an exorbitantly high price for the period. The book included illustrations by landscape painter
John Samuelson Templeton
John is a common English name and surname:
* John (given name)
* John (surname)
John may also refer to:
New Testament
Works
* Gospel of John, a title often shortened to John
* First Epistle of John, often shortened to 1 John
* Second E ...
, although many of these inaccurately portray ichthyosaurs with straight tails and leaving the water to bask on rocks.
Suevoleviathan integer
''Suevoleviathan'' is an extinct genus of primitive ichthyosaur found in the Early Jurassic (Toarcian) of Holzmaden, Germany.
Taxonomy
The genus was named in 1998 by Michael Maisch for ''Leptopterygius disinteger'' and ''Ichthyosaurus integer ...
''.
1846
* Chaning Pearce discovered that a specimen of ''Ichthyosaurus communis'' contained the skeleton of another, tiny ''I. communis'' partially protruding from the larger individual's pelvis. After discussing the fossil with Sir Richard Owen, Pearce concluded that the specimen was a female killed and preserved in the act of giving birth.
1850s
1851
* Gideon Mantell described the new species '' Ichthyosaurus longirostris'' based on fossil found in Whitby, Yorkshire. Its upper jaw was so much longer than its lower jaw that it resembled a swordfish.
* Meyer described the new genus and species ''
Tholodus schmidi
''Tholodus'' is an extinct genus of basal (phylogenetics), basal ichthyopterygian known from the Middle Triassic (mid-late Anisian to late Ladinian stage) of Germany, northeastern Italy and possibly China. It was first named by Christian Erich He ...
''.
1852
* Quenstedt described the species that would later become known as ''
Contectopalatus atavus
''Contectopalatus'' was a primitive ichthyosaur, an extinct fish-like marine reptile from the Middle Triassic of Germany and China. It was originally named ''Ichthyosaurus atavus'' (Quenstedt, 1851/52), and later ''Mixosaurus atavus'' (Quensted ...
Henry Coles
Henry Coles (born 11 March 1951) is a retired Australian rules footballer, who played for Collingwood and Melbourne in the Victorian Football League (VFL).
VFL career Collingwood
Coles was recruited from Kingsbury,cephalopods the ichthyosaur had eaten.
* Wagner described the species that would one day be known as ''
Aegirosaurus leptospondylus
''Aegirosaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaurs known from the late Jurassic and early Cretaceous of Europe. It was originally named as a species of ''Ichthyosaurus''.
Discovery and species
Originally describe ...
''.
1854
* Sir Richard Owen inaccurately reconstructed ''Ichthyosaurus'' as straight-tailed and able to bask on rocks for an outdoor exhibit at Sydenham.1858
* Quenstedt described the species that would later come to be known as '' Stenopterygius quadriscissus '' and ''
Stenopterygius triscissus
''Stenopterygius'' is an extinct genus of thunnosaur ichthyosaur known from Europe (England, France, Germany, Luxembourg and Switzerland). This genus of ichthyosaur was about long and weighed .Huene F. von 1939. Ein ganzes Ichthyosaurier-Skelett ...
''.
1860s
1860
* Sir Richard Owen tried to rename the Ichthyosauria the Ichthyopterygia.
* The first ichthyosaur fossils known from Australia were discovered in
Queensland
)
, nickname = Sunshine State
, image_map = Queensland in Australia.svg
, map_caption = Location of Queensland in Australia
, subdivision_type = Country
, subdivision_name = Australia
, established_title = Before federation
, established_ ...
and named ''
Ichthyosaurus australis
''Ichthyosaurus'' (derived from Greek ' () meaning 'fish' and ' () meaning 'lizard') is a genus of ichthyosaurs from the Early Jurassic (Hettangian - Pliensbachian), with possible Late Triassic record, from Europe ( Belgium, England, Ger ...
Oskar Fraas Oskar may refer to:
* oskar (gene), the Drosophila gene
* Oskar (given name), masculine given name
See also
* Oscar (disambiguation)
{{disambig ...
Cymbospondylus petrinus
''Cymbospondylus'' (a Greek word meaning "boat vertebrae") was a basal early ichthyosaur that lived between the early and middle Triassic period (249-237 million years ago). Previously, the genus was classified as a shastasaurid, but more rece ...
''.
1870s
1871
* Hulke described the species that would later become known as ''
Nannopterygius enthekiodon
''Nannopterygius'' (meaning "small wing/flipper" in Greek) is an extinct genus of ophthalmosaurid ichthyosaur that lived during the Middle Jurassic to the Early Cretaceous ( Callovian to Berriasian stages). Fossils are known from England, Kaza ...
''.
1874
* Seeley described the new genus and species '' Ophthalmosaurus icenicus''.
1876
* Blake described the species that would one day come to be known as ''
Temnodontosaurus crassimanus
''Temnodontosaurus'' (Greek for "cutting-tooth lizard"temno, meaning "to cut", odont meaning "tooth" and sauros meaning "lizard") is an extinct genus of ichthyosaur from the Early Jurassic period. They lived between 200 and 175 million years ...
''.
1879
* Marsh described the species that would one day come to be known as ''
Ophthalmosaurus natans
''Baptanodon'' is an ichthyosaur of the Late Jurassic period (160-156 million years ago), named for its supposed lack of teeth (although teeth of this genus have since been discovered). It had a graceful long dolphin-shaped body, and its jaws w ...
''.
1880s
1880
* Harry Govier Seeley published an argument in favor of live birth in ichthyosaurs. Seeley noted that many ichthyosaur specimens from Britain and
Germany
Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
contained miniature ichthyosaur skeletons inside them. While some researchers interpreted these remains as fossil stomach contents, Seeley observed that the smaller skeletons tend to be located too near the rear of the animal to be stomach contents. Instead, he interpreted the small skeletons as fetuses and the larger ones as pregnant females.1881
* Owen described the species ''
Ichthyosaurus breviceps
''Ichthyosaurus'' (derived from Greek ' () meaning 'fish' and ' () meaning 'lizard') is a genus of ichthyosaurs from the Early Jurassic (Hettangian - Pliensbachian), with possible Late Triassic record, from Europe (Belgium, England, Germany, Swi ...
''.
1886
* Bassani described the species that would later come to be known as '' Mixosaurus cornalianus''.
1887
* Georg Baur published a paper speculating on the origins of ichthyosaurs. He concluded that ichthyosaurs were descended from relatives of the Sphenodontidae, the family that includes the
modern tuatara
Tuatara (''Sphenodon punctatus'') are reptiles endemic to New Zealand. Despite their close resemblance to lizards, they are part of a distinct lineage, the order Rhynchocephalia. The name ''tuatara'' is derived from the Māori language and ...
Brachypterygius cantabridgiensis
''Brachypterygius'' (meaning ″short wing/paddle″ in Greek) is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur known from the Late Jurassic of England.McGowan, C. & Motani, R. ''Ichthyopterygia''. In Sues, H.-D. (ed.) Handbook ...
1895
* Merriam described the new genus and species ''
Shastasaurus pacificus
''Shastasaurus'' ("Mount Shasta lizard") is a very large extinct genus of ichthyosaur from the middle and late Triassic, and is the largest known marine reptile.Hilton, Richard P., ''Dinosaurs and Other Mesozoic Animals of California'', Universit ...
J. C. Merriam
John Campbell Merriam (October 20, 1869 – October 30, 1945) was an American paleontologist, educator, and conservationist. The first vertebrate paleontologist on the West Coast of the United States, he is best known for his taxonomy of ver ...
named the new species '' Shastasaurus alexandrae'' after Annie Alexander, the wealthy fossil enthusiast who financed much of his research and founded the
Museum of Paleontology
A museum ( ; plural museums or, rarely, musea) is a building or institution that cares for and displays a collection of artifacts and other objects of artistic, cultural, historical, or scientific importance. Many public museums make these ...
at Berkeley. He also described the species that would later come to be known as Californosaurus perrini.
1903
* J. C. Merriam described the new genus ''
Toretocnemus
''Toretocnemus'' is an extinct genus of ichthyosaur. Its remains have been found in California, United States, in Triassic layers of the Carnian Hosselkus Limestone.
History of research
The specimen that would eventually be cataloged as UCMP 81 ...
'' from
Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
California
California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the ...
. Along with the genus, he described the species ''
Toretocnemus californicus
''Toretocnemus'' is an extinct genus of ichthyosaur. Its remains have been found in California, United States, in Triassic layers of the Carnian Hosselkus Limestone.
History of research
The specimen that would eventually be cataloged as UCMP ...
'' and ''
Toretocnemus zitteli
''Toretocnemus'' is an extinct genus of ichthyosaur. Its remains have been found in California, United States, in Triassic layers of the Carnian Hosselkus Limestone.
History of research
The specimen that would eventually be cataloged as UCM ...
''.
1904
* Boulenger described the species that would later become known as '' Brachypterygius extremus''.
* Jaekel described the new genus ''
Stenopterygius
''Stenopterygius'' is an extinct genus of thunnosaur ichthyosaur known from Europe (England, France, Germany, Luxembourg and Switzerland). This genus of ichthyosaur was about long and weighed .Huene F. von 1939. Ein ganzes Ichthyosaurier-Skelett ...
''.
1906
* J. C. Merriam discovered the unusual remains of an animal with short jaws and button-shaped teeth in
Nevada
Nevada ( ; ) is a state in the Western region of the United States. It is bordered by Oregon to the northwest, Idaho to the northeast, California to the west, Arizona to the southeast, and Utah to the east. Nevada is the 7th-most extensive, ...
. He thought it was the first member of a new group of reptiles related to placodonts or rhynchosaurs but it would actually come to be recognized as ichthyosaurian and described as the genus ''
Omphalosaurus
''Omphalosaurus'' (from the Greek root "Button Lizard", for their button-like teeth) is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about ''O ...
''.1908
* Broili described the species that would later come to be known as '' Platypterygius platydactylus''.
1909
* Abel erected the genus '' Eurhinosaurus'' for the species ''Ichthyosaurus longirostris''.
1910s
1910
* Some putative ichthyosaur limb bones were found in
Spitsbergen
Spitsbergen (; formerly known as West Spitsbergen; Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipelago in northern N ...
. Carl Wiman would later refer these remains to the species ''
Omphalosaurus nevadanus
''Omphalosaurus'' (from the Greek root "Button Lizard", for their button-like teeth) is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about ''Omph ...
''. Wiman also described the new genus and species ''
Pessopteryx nisseri
''Pessopteryx'' is an extinct genus of ichthyosaur from the Early Triassic ( Olenekian) of Svalbard, Norway. The genus originally contained four species, ''P. nisseri'', ''P. arctica'', ''P. pinguis'', and ''P. minor'', which were named in 1910 b ...
'' as well as the species that would one day be known as ''
Isfjordosaurus minor
''Isfjordosaurus'' is an extinct genus of ichthyopterygian marine reptile that lived during the Early Triassic. Fossils have been found on the island of Spitsbergen, part of the Svalbard archipelago off the northern coast of Norway. It was formal ...
''.
* Andrews speculated that since many specimens of '' Ophthalmosaurus'' are toothless that it may have fed via suction feeding on soft foods like squid.
* Merriam described the new genus and species '' Phalarodon fraasi''.
1916
* Huene described the species now known as ''
Phalarodon major
''Phalarodon'' is an extinct genus of ichthyosaur. Its remains have been found in China, North America, and Spitsbergen. It measured between and weighed more than .
See also
* List of ichthyosaurs
* Timeline of ichthyosaur research
This time ...
1922
* German paleontologist Friedrich von Huene described the new genus '' Brachypterygius''. He also described the genera '' Nannopterygius'' and '' Platypterygius''.
1925
* von Huene described a new species of ''Shastasaurus''.1926
* von Huene erected the new genus '' Suevoleviathan'' for the species "''Leptopterygius''" ''disinteger''.
1927
* Huene described the species that would later come to be known as ''
Platypterygius hauthali
''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus o ...
Simeon Muller
Simeon () is a given name, from the Hebrew (Biblical ''Šimʿon'', Tiberian ''Šimʿôn''), usually transliterated as Shimon. In Greek it is written Συμεών, hence the Latinized spelling Symeon.
Meaning
The name is derived from Simeon, son ...
discovered the remains of 40 gigantic ichthyosaurs in Nevada.1929
* Wiman described the new genus and species '' Grippia longirostris''.
1930s
1931
* Huene described the new species '' Stenopterygius uniter'' as well as the species that would later become known as ''
Temnodontosaurus nuertingensis
''Temnodontosaurus'' (Greek for "cutting-tooth lizard"temno, meaning "to cut", odont meaning "tooth" and sauros meaning "lizard") is an extinct genus of ichthyosaur from the Early Jurassic period. They lived between 200 and 175 million years ...
''.
1934
* The most complete known ichthyosaur specimen from Australia, a '' Platypterygius'' was discovered in Queensland. Despite not being fully grown it was 18 feet in length.
* Kuhn described the genus ''
Californosaurus
''Californosaurus'' ('California lizard') is an extinct genus of ichthyosaur, an extinct marine reptile, from the Lower Hosselkus Limestone (Carnian, Late Triassic) of California.
Taxonomy
Merriam (1902) described it as a new species of ''Shas ...
''.
1939
* Nace described the genus that would one day be known as ''
Platypterygius americanus
''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus or ...
Platypterygius hercynicus
''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus ...
''.
1948
* Alfred Romer noted that Jurassic ichthyosaurs were so highly specialized for aquatic life that their anatomy exhibited no sign of descent from any known terrestrial reptile group.
1950s
1951
* Willy Ley, a German zoologist, observed that more than two hundred ichthyosaur specimens were being recovered from Holzmaden annually.1954
* Charles Camp and Samuel Welles of Berkeley lead the excavation of the gigantic ichthyosaur fossils discovered by Simeon Muller in the 20s. They estimated their body length to be 50 feet, which is roughly as large as a modern
humpback whale
The humpback whale (''Megaptera novaeangliae'') is a species of baleen whale. It is a rorqual (a member of the family Balaenopteridae) and is the only species in the genus ''Megaptera''. Adults range in length from and weigh up to . The hum ...
.1956
* Whitear reported brown pigmentation preserved in an ichthyosaur fossil.1957
* Camp's ichthyosaur excavation in Nevada ceased.
* Not far from Stowbridge in
Norfolk
Norfolk () is a ceremonial and non-metropolitan county in East Anglia in England. It borders Lincolnshire to the north-west, Cambridgeshire to the west and south-west, and Suffolk to the south. Its northern and eastern boundaries are the Nort ...
, workers serendipitously discovered a large partial ichthyosaur skeleton while digging a drainage channel. The specimen was later thought to belong to the genus ''Ophthalmosaurus''.
1960s
1963
* Camp resumed excavating the gigantic ichthyosaurs in Nevada.1965
* Colbert hypothesized that ichthyosaurs were descended from cotylosaurs.
* Camp's ichthyosaur excavation in Nevada ended, with 35–40 partial ichthyosaur skeletons unearthed.1968
* Pollard reported that '' Temnodontosaurus'' ate cephalopods.
1970s
1972
* McGowan argued that the narrow-finned longipinnate and the broad-finned latipinnate ichthyosaurs could be distinguished based on features of their skulls.
* Young and Dong described the new genera and species ''
Chaohusaurus geishanensis
''Chaohusaurus'' is an extinct genus of basal ichthyopterygian, depending on definition possibly ichthyosaur, from the Early Triassic of Chaohu and Yuanan, China.
Discovery
The type species ''Chaohusaurus geishanensis'' was named and descri ...
'' and '' Himalayasaurus tibetensis''.
1973
* McGowan described the cranial anatomy of ''Ichthyosaurus''. He observed that it had a very large
brain
The brain is an organ that serves as the center of the nervous system in all vertebrate and most invertebrate animals. It consists of nervous tissue and is typically located in the head ( cephalization), usually near organs for special ...
cerebellum
The cerebellum (Latin for "little brain") is a major feature of the hindbrain of all vertebrates. Although usually smaller than the cerebrum, in some animals such as the mormyrid fishes it may be as large as or even larger. In humans, the cere ...
as suggestive of highly coordinated bodily movements. He also noted that its large corpus striatum implied that it had a sophisticated repertoire of instinctive behaviors. He speculated that social and parental behavior may have been among them. By contrast, the lagena of Ichthyosaurus was small, suggesting that ichthyosaurs had weak sense of hearing.1974
* McGowan described the species that would later come to be known as ''
Temnodontosaurus eurycephalus
''Temnodontosaurus eurycephalus'' is an extinct species of marine reptile. It is thought to have been a nektonic carnivore. Its name comes from the Greek ''ευρύς'', for "wide" and ''κεφαλή'' for "head". It possibly belongs to a new ge ...
''.
1975
* Charles Camp died before he could finish his monograph on the Berlin-Ichthyosaur State Park ichthyosaurs.1976
* Joseph Gregory, a friend of Charles Camp, published some of Camp's research on the ichthyosaurs of Berlin-Ichthyosaur State Park.
* The new genus and species '' Shonisaurus popularis'' was described.
* McGowan observed that the eyes of the supposed ''Ophthalmosaurus'' discovered in Norfolk during 1950s were too small for the animal to be referred to that genus and reclassified it in a new one: ''
Grendelius
''Grendelius'' is a genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic (Kimmeridgian-Tithonian) of the UK and European Russia. It was a medium-sized ichthyosaur measuring about long.
Taxonomy
The type species, ''Grende ...
''. He also described the species that would later come to be known as ''
Brachypterygius mordax
''Grendelius'' is a genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic ( Kimmeridgian-Tithonian) of the UK and European Russia. It was a medium-sized ichthyosaur measuring about long.
Taxonomy
The type species, ''Grend ...
''.
1978
* Shikama, Kamei and Murata described the new genus and species '' Utatsusaurus hataii''.
1979
* McGowan observed that more ichthyosaur remains have been found in the
Early Jurassic
The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma  ...
strata of southern Germany than anywhere else in the world. He also speculated that in life most ichthyosaurs' biogeographic ranges would be as wide as those of modern
whales
Whales are a widely distributed and diverse group of fully aquatic placental marine mammals. As an informal and colloquial grouping, they correspond to large members of the infraorder Cetacea, i.e. all cetaceans apart from dolphins a ...
and dolphins. McGowan attributed the narrow ranges of most known ichthyosaur species to an artifact of the fossil record, since paleontologists can only recover fossils from accessible exposures of sedimentary rocks formed in settings conducive to fossilization, which may not correspond to the complete life range of a given species. He also rejected his own previous conclusion that latipinnate and longipinnate ichthyosaurs could be distinguished based on features of the skull and expressed doubt about the validity of the latipinnate-longipinnate dichotomy altogether.
1980s
1980
* Joseph Gregory published the rest of his late friend Charles Camp's research on the ichthyosaurs of Berlin-Ichthyosaur State Park. This publication contained a reconstruction of '' Shonisaurus'' portraying as a long-skulled long-finned animal with an unusually deep "pot belly".1983
*
Angela Kirton
Angela may refer to:
Places
* Angela, Montana
* Angela Lake, in Volusia County, Florida
* Lake Angela, in Lyon Township, Oakland County, Michigan
* Lake Angela, the reservoir impounded by the source dam of the South Yuba River
Fiction
* Ange ...
reported the presence of teeth in skulls attributable to ''Ophthalmosaurus'' from England. The toothlessness of many adult skulls suggest that either ''Ophthalmosaurus'' lost its teeth as it aged or the teeth were only loosely attached and prone to falling out after death.1984
* Wade observed that the longest digit in the flipper of ''Platypterygius'' was made up of a series of 30 bones.
* Fossils of an unusual ichthyosaur whose upper jaw was much longer than its lower jaw were found.1985
* An adult ''Ichthyosaurus communis'' specimen was found with a tiny associated embryo of the same species in
Kilve
Kilve is a village in the Somerset West and Taunton district of Somerset, England, within the Quantock Hills Area of Outstanding Natural Beauty, the first AONB to be established, in 1957.
It lies on the A39 almost exactly equidistant from Bridg ...
,
Somerset
( en, All The People of Somerset)
, locator_map =
, coordinates =
, region = South West England
, established_date = Ancient
, established_by =
, preceded_by =
, origin =
, lord_lieutenant_office =Lord Lieutenant of Somerset
, lor ...
.
* A man named
Bernd Neubig
Bernd is a Low German short form of the given name Bernhard (English Bernard).
List of persons with given name Bernd
The following people share the name Bernd.
*Bernd Brückler (born 1981), Austrian hockey player
*Bernd Eichinger (1949–2011), G ...
discovered an ichthyosaur skeleton during construction of a railroad in Karlstadt, Germany. The specimen was the most complete ichthyosaur discovered in that region of the country.
* Gasparini reported the first scientifically documented ichthyosaur remains from Argentina. He referred the Argentine ichthyosaur to the genus ''Ophthalmosaurus''.
* Otschev and Efimov described the new species '' Platypterygius birjukovi''.
1986
* Christopher McGowan began leading a series of expeditions to Williston Lake,
British Columbia
British Columbia (commonly abbreviated as BC) is the westernmost Provinces and territories of Canada, province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include ...
, culminating in the discovery of the most complete known skeleton of ''Shastasaurus'', which also served as the type specimen of a new species that would later be described in 1994.
* German paleontologist Jurgen Reiss speculated that ichthyosaurs swam using their front flippers the way modern penguins do rather than propelling themselves with their tail flukes.
* McGowan named the ichthyosaur with an unusually long, protruding upper jaw from Somerset ''
Excalibosaurus
''Excalibosaurus'' (meaning "Excalibur's lizard") is a monotypic genus of marine prehistoric reptiles ( ichthyosaurs) that lived during the Sinemurian stage (approximately 196.5 ± 2 Ma to 190.8 ± 1.0 Ma (million years ago)) of the Early Jura ...
'', because it was found in the same general region where the legendary King Arthur obtained his sword.1987
* Mazin described the new species '' Omphalosaurus nettarhynchus''.
1988
* Massare and Callaway described a pregnant ''Mixosaurus'' specimen discovered in the
Alps
The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, ...
Italy
Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
. The specimen represented the oldest known evidence for live birth in ichthyosaurs.
* Three specimens attributable to a new Triassic ichthyosaur were discovered near Phattalung,
Thailand
Thailand ( ), historically known as Siam () and officially the Kingdom of Thailand, is a country in Southeast Asia, located at the centre of the Indochinese Peninsula, spanning , with a population of almost 70 million. The country is b ...
Dorset
Dorset ( ; archaically: Dorsetshire , ) is a county in South West England on the English Channel coast. The ceremonial county comprises the unitary authority areas of Bournemouth, Christchurch and Poole and Dorset. Covering an area of ...
, England, a new specimen of ''Grendelius'' was discovered. However, when Christoper McGowan studied this new specimen he realized that while distinct from ''Ophthalmosaurus'', ''Grendelius'' was apparently the same as the previously named genus ''Brachypterygius'' and the names were synonymized.
* McGowan observed that many '' Leptonectes'' specimens have somewhat protruding upper jaws.
* Remains of a Triassic ichthyosaur were discovered in
Sonora
Sonora (), officially Estado Libre y Soberano de Sonora ( en, Free and Sovereign State of Sonora), is one of the 31 states which, along with Mexico City, comprise the Federal Entities of Mexico. The state is divided into 72 municipalities; the ...
,
Mexico
Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...
.
* Callaway and Massare regarded the genus ''
Phalarodon
''Phalarodon'' is an extinct genus of ichthyosaur. Its remains have been found in China, North America, and Spitsbergen. It measured between and weighed more than .
See also
* List of ichthyosaurs
* Timeline of ichthyosaur research
This time ...
'' as a ''nomen dubium''.
* Sander described the new species ''
Cymbospondylus buchseri
''Cymbospondylus'' (a Greek word meaning "boat vertebrae") was a basal early ichthyosaur that lived between the early and middle Triassic period (249-237 million years ago). Previously, the genus was classified as a shastasaurid, but more r ...
''.
1990s
1990
* Bradley Kosch of Berlin-Ichthyosaur State Park criticized Camp's 1980 reconstruction of the park's famous ''Shonisaurus popularis'' as having too short a backbone and overly deep ribs, responsible for its supposedly deep pot belly. He noted that the illustration differed from both the text of Camp's published description and his own private field notes.
* Stephen Jay Gould published an article about the tail bend in ichthyosaurs.
* De Buffrenil and
Mazin
Mazin ( sr-cyr, Мазин) is a village in Croatia. It is connected by the D218 highway.
Population
According to the 2011 census, Mazin had 47 inhabitants.
1991 census
According to the 1991 census, settlement of Mazin had 362 inhabitants, ...
found a woven texture in the bones of ''Ichthyosaurus'', ''Omphalosaurus'', and ''Stenopterygius''. This texture is found only in the bones of quickly growing animals, suggesting that ichthyosaurs had high metabolic rates and may even have been
warm blooded
Warm-blooded is an informal term referring to animal species which can maintain a body temperature higher than their environment. In particular, homeothermy, homeothermic species maintain a stable body temperature by regulating metabolic proce ...
.
* Massare and Callaway observed that Triassic ichthyosaurs had more elongated body plans than their Jurassic successors.1991
* An archaeologist named Keary Walde discovered large fossil bones near the Sikanni Chief River. He reported his discovery to the Royal Tyrell Museum. The museum's curator,
Elizabeth Nicholls
Elizabeth (Betsy) Laura Nicholls (January 31, 1946 – October 18, 2004) was an American-Canadian paleontology, paleontologist who specialized in Triassic marine reptiles. She was a paleontologist at the Royal Tyrrell Museum in Alberta, Cana ...
, was also a respected marine reptile researcher, visited the site. She was astonished to realize that the bones discovered by Walde were the remains of an ichthyosaur roughly 75 feet long, one and a half times as big as the largest known ichthyosaur at the time.
* McGowan argued that ichthyosaur fossils with new-born remains protruding from the birth canal don't imply that the mother died giving birth. Instead, they could be expelled from an already dead mother by the pressure of gas-build up as her rotting carcass bloated. An example of this in modern marine life was documented where the carcasses of beached false killer whales were given a shallow burial. Several months later, the pregnant females' fetuses had been partly expelled. He also observed that roughly 35 ichthyosaur specimens a year were still being recovered from Holzmaden. He suggested that at least some ichthyosaurs may have been warm-blooded and swift swimming like modern tuna due to their similar body plans. He also noted that even if they weren't warm blooded their large body mass would help maintain stable body temperatures.
* Lingham-Soliar referred to the swimming style of advanced ichthyosaurs whose body remained stiff while large muscles powered the tail's swimming stroke as "axial oscillation".
* Mazin and others described the new genus and species '' Thaisaurus chonglakmanii''.
1992
* Field work on behalf of Alberta's Royal Tyrell Museum began in the Pink Mountains of British Columbia 60 miles north of where Christopher McGowan's field work was conducted. The Royal Tyrell researchers would discover Late Triassic ichthyosaur remains in the sediments of the Pardonet Formation.
* Judy Massare proposed an explanation for the high percentage of pregnant ''Stenopterygius'' at Holzmaden. She speculated that the area was used as a breeding ground, the way whales congregate to give birth in areas of shallow water today. The coordination of a large number of births at the same time and place would increase the young's chances of survival because there would be too many for the local predator population to eat. Massare suggested that another ichthyosaur genus, ''Leptopterygius'' (now known as ''Leptonectes''), was one such local predator that may have fed on vulnerable young ichthyosaurs.
* Nathalie Bardet dismissed claims of ichthyosaur fossils from rocks of more recent age than the Cenomanian. She also discussed possible causes for the extinction of the ichthyosaurs. She expressed doubt that ichthyosaurs were replaced by mosasaurs because they would not be in direct competition with each other. Instead she posited a connection to an extinction event that affected cephalopods at the boundary between the Cenomanian and Turonian ages. She proposed that the disappearance of these many cephalopod species may have deprived ichthyosaurs of their food source and caused their extinction.1993
* Charles Deeming and others published a paper on the 1985 ichthyosaur embryo discovered in Somerset. They observed that many pregnant ichthyosaur specimens contain fetuses oriented head-first toward the birth canal even though they were probably born tail-first as a precaution to prevent drowning. This suggests that there may have been complications during the pregnancy, like the fetus being too large to pass through the birth canal. If the decomposing fetus remained trapped in the mother, it would very likely kill her.
* Jennifer Hogler re-examined the tails of the early large ichthyosaurs '' Cymbospondylus'' and ''Shonisaurus'', finding the wedge-shaped vertebrae that form the tail-bend that composes the tail fluke. This find contradicted the widespread idea that these early ichthyosaurs had straight tails and therefore lacked well-developed flukes.
* Russell described the species that would later come to be known as ''
Arthropterygius chrisorum
''Arthropterygius'' is a widespread genus of ophthalmosaurid ichthyosaur which existed in Canada, Norway, Russia, and Argentina from the late Jurassic period and possibly to the earliest Cretaceous.
Description
''Arthropterygius'' appea ...
''.
* McGowan described the species that would later come to be known as '' Leptonectes solei''.
1994
* Axel Hungerbuhler bemoaned the taxonomic confusion caused by German geologist Friedrich Quenstedt's early research on the ichthyosaurs at Holzmaden. He attributed the bulk of this confusion to Quenstedt's unorthodox naming practices, which often exceeded the two-named binomials of standard biological nomenclature to consist of three or four names. Further, the fossils themselves were poorly organized and many of the type specimens he founded species on were unlabeled. Hungerbuhler named two new species of his own: ''Stenopterygius cuniceps'' and ''S. macrophasma''.
* Christopher McGowan named the new species of ''Shastasaurus'' discovered in British Columbia ''S. neoscapularis''. He also reviewed the species previously referred to the genus, finding many of them to be dubious, like ''S. altus'', ''S. careyi'', ''S. carinthiacus'', and ''S. osmonti''. However, two previously described species, the type, ''S. pacificus'', and the referred species ''S. alexandrae'' were found to be valid. He criticized Merriam for oversplitting ''Shastasaurus'' because most of the resulting names were useless and mislead the paleontological community into thinking that the genus was better understood than it really was.
* Fernandez described the new ichthyosaur genus '' Chacaisaurus'' from the
Vaca Muerta Formation
The Vaca Muerta Formation, commonly known as Vaca Muerta (Spanish for ''Dead Cow''), is a geologic formation of Late Jurassic to Early Cretaceous age, located in the Neuquén Basin in northern Patagonia, Argentina. It is well known as the host roc ...
of Argentina.
* Bardet and others reported ''Platypterygius'' fossils from late Cenomanian rocks in
Bavaria
Bavaria ( ; ), officially the Free State of Bavaria (german: Freistaat Bayern, link=no ), is a state in the south-east of Germany. With an area of , Bavaria is the largest German state by land area, comprising roughly a fifth of the total l ...
. These were the remains latest known ichthyosaurs.
* Fernández described the species that would later come to be known as '' Stenopterygius cayi''.
1995
* ''January:'' A man named Chris Moore discovered much of the front half of an ichthyosaur skeleton in the Belemnite Marls of
Seatown
Seatown is a coastal hamlet in Dorset, England, on the English Channel approximately west-southwest of Bridport. It lies within the civil parish of Chideock.
The coast at Seatown is part of the Jurassic Coast, a World Heritage Site stretching fo ...
, Dorset. The specimen was a juvenile of a new ''Leptopterygius'' species, named ''L. moorei'' after Moore. In life, the specimen was probably about 8 feet long.
* McGowan renamed the ichthyosaur genus ''Leptopterygius'' to ''Leptonectes''. He also synonymized ''Leptopterygius'' with ''Temnodontosaurus''. He also described the new genus and species '' Hudsonelpidia brevirostris''.
* Martill claimed that despite an abundance of ichthyosaur specimens with preserved soft tissue, there was no evidence that ichthyosaur skin was covered in scales.
* Tichy described the new species '' Omphalosaurus wolfi''.
* Nicholls and Brinkman described the new genus and species '' Parvinatator wapitiensis''.
1996
* Motani and others argued that sharks are the best modern analogues for ichthyosaurs because of their similar body plans.
* McGowan reported the serendipitous discovery of evidence for a new giant ichthyosaur species in a museum collection. He was studying the ichthyosaur fossils curated by the National Academy of Sciences in Philadelphia. One
Early Jurassic
The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma  ...
bone catalogued as a part of a shoulder (the coracoid) he recognized as actually being a gigantic skull bone (the
quadrate
Quadrate may refer to:
* Quadrate bone
* Quadrate (heraldry)
* Quadrate lobe of liver
* Quadrate tubercle
The quadrate tubercle is a small tubercle found upon the upper part of the femur. It serves as a point of insertion of the quadratus femori ...
). The misidentification was probably due to the quadrate's exceptionally large size as the coracoid is generally one of the larger bones in the ichthyosaur body. The implication of such a large quadrate being found was the existence a previously unknown ichthyosaur of similar or greater size to ''Shonisaurus'' itself. In life this animal may have been more than 50 feet long. He also described the new genus '' Leptonectes'' as well as the species that would later come to be known as '' Macgowania janiceps''.
* Motani, You, and McGowan observed that the primitive ichthyosaur '' Chensaurus'' had had a relatively large number of vertebrae. This suggests that it swam in an eel-like fashion. The researchers interpreted the course of ichthyosaur evolution as starting with an eel-like body plan, transitioning to a jack-like body plan, and reaching its culmination in a
tuna
A tuna is a saltwater fish that belongs to the tribe Thunnini, a subgrouping of the Scombridae ( mackerel) family. The Thunnini comprise 15 species across five genera, the sizes of which vary greatly, ranging from the bullet tuna (max le ...
-like body plan.
* Dal Sasso and Pinna described the new genus and species ''
Besanosaurus leptorhynchus
''Besanosaurus'' (meaning "Besano Italy.html" ;"title="ombardy, N. Italy">ombardy, N. Italylizard") is a genus of large ichthyosaur (a marine reptile, not a dinosaur) that lived during the middle Triassic period, approximately 235 million years ...
''.
1997
* Ryosuke Motani completed his doctoral dissertation: "Phylogeny of the Ichthyosauria with Special Reference to Triassic Forms".
* Callaway argued that it was inaccurate to refer to ichthyosaurs as ichthyopterygians.
* Callaway examined the mixosaurid fossils housed at the
natural history museum
A natural history museum or museum of natural history is a scientific institution with natural history collections that include current and historical records of animals, plants, fungi, ecosystems, geology, paleontology, climatology, and more ...
in Zurich. He regarded only two species of ''Mixosaurus'' to be valid; ''M. cornalianus'' and ''M. atavus''. He regarded ''M. maotaiensis'', ''M. natans'', and ''M. nordeskioeldii'' as junior synonyms of those species. He also regarded several species in the genus ''Phalarodon'' as synonymous with the two valid ''Mixosaurus'' species.
*
Martin Sander Martin may refer to:
Places
* Martin City (disambiguation)
* Martin County (disambiguation)
* Martin Township (disambiguation)
Antarctica
* Martin Peninsula, Marie Byrd Land
* Port Martin, Adelie Land
* Point Martin, South Orkney Islands
Aus ...
Bonn
The federal city of Bonn ( lat, Bonna) is a city on the banks of the Rhine in the German state of North Rhine-Westphalia, with a population of over 300,000. About south-southeast of Cologne, Bonn is in the southernmost part of the Rhine-Ru ...
named the ichthyosaur found in Karlstadt, Germany '' Shastasaurus neubigi'' in honor of its discoverer. The specimen originated in the Muschelkalk, which was deposited in a shallow sea. Because the waters were so shallow and large ichthyosaurs are so infrequently found in the Muschelkalk Sander speculated that the ichthyosaur entered the region where it would later fossilize by accident in a deviation from the normal range of the species.
* Fernández described the new genus and species ''
Caypullisaurus bonapartei
''Caypullisaurus'' is an extinct genus of platypterygiine ophthalmosaurid ichthyosaur from the Late Jurassic to the Early Cretaceous (Tithonian and Berriasian stages) of Argentina. Its holotype was collected from the Vaca Muerta Formation of Ce ...
''.
* Maisch and Matzke described the new genus and species '' Mikadocephalus gracilirostris''.
* Arkhangelsky described the species that would later come to be known as '' Ophthalmosaurus saveljeviensis''.
* Efimov described the species that would later come to be known as '' Platypterygius bedengensis''.
* Páramo described the new species ''
Platypterygius sachicarum
''Platypterygius'' is a historically paraphyletic genus of platypterygiine ichthyosaur from the Cretaceous period. It was historically used as a wastebasket taxon, and most species within ''Platypterygius'' likely are undiagnostic at the genus or ...
''
1998
* Motani, Minoura, and Ando published a discussion of the ichthyosaur '' Utatsusaurus''. They noted its primitive position in the ichthyosaur family tree and the same number of vertebrae in the front part of its body as modern catsharks. They interpreted ''Utatsusaurus'' as a maneuverable shallow water predator that swam with eel-like undulations along most of its body length.
* Maisch named the family Leptonectidae for a clade including ''Eurhinosaurus'', ''Excalibosaurus'', and ''Leptonectes''. Distinguishing traits of the family include their large eyes, long flippers consisting of three or four main "digits", and their long over-biting snouts.
* Maisch and Matzke erected the genus ''
Wimanius
''Wimanius'' is a mixosaurian ichthyosaur, and the only genus in the family Wimaniidae. It existed during the Triassic Period in what is now Switzerland. It was described by Maisch and Matzke in 1998 based on fossils found in the Monte San Gior ...
''. The generic name honored Carl Wiman, who performed notable research on Triassic ichthyosaurs. They also named the species '' Wimanius odontopalatus''.
* Maisch and Matzke described the new genus ''
Contectopalatus
''Contectopalatus'' was a primitive ichthyosaur, an extinct fish-like marine reptile from the Middle Triassic of Germany and China. It was originally named ''Ichthyosaurus atavus'' (Quenstedt, 1851/52), and later ''Mixosaurus atavus'' (Quensted ...
''. It had a high crest running the midlength of its cranium for jaw muscle attachment giving it a powerful bite. It also would have been about 16 feet long in life, making it twice as long as any previously discovered mixosaurid.
* Maisch erected the genus '' Suevoleviathan'' to house the species ''Leptopterygius disinteger''.
* Darren Naish debunked the 1986 claim by Jurgen Riess that ichthyosaurs swam with their front flippers rather than their tail flukes. He concluded "if an animal has a propulsive surface on the end of its tail, it uses it."
* Efimov described the species that would later come to be known as '' Brachypterygius pseudoscythius''.
* Brinkmann described the species that would later come to be known as '' Mixosaurus kuhnschnyderi''.
1999
* Ryosuke Motani published a phylogeny of the ichthyosaurs. Motani regarded ''Thaisaurus'' as "incertae sedis" due to its pronounced similarity to '' Chaohusaurus'' and how little was known about its fossils. He also erected two new genera, ''
Macgowania
''Macgowania'' is an extinct genus of parvipelvian ichthyosaur known from British Columbia of Canada.
History of research
''Macgowania'' is known only from the holotype ROM 41992 ( RBCM EH 91.2.5), a partial skeleton which preserv ...
'' (named in honor of McGowan) and ''
Isfjordosaurus
''Isfjordosaurus'' is an extinct genus of ichthyopterygian marine reptile that lived during the Early Triassic. Fossils have been found on the island of Spitsbergen, part of the Svalbard archipelago off the northern coast of Norway. It was forma ...
'' (named after Isfjord, Spitsbergen).
* The expedition to the Sikanni Chief River, British Columbia led by Elizabeth Nicholls of the Royal Tyrell Museum excavated the animal's 18 foot long skull, which had to be split into pieces for removal. These pieces were so heavy a cargo helicopter was needed to transport them. The largest weighed 8,860 lbs.
* McGowan and Motani reported the results of their re-examination of the ''Shonisaurus'' specimens described by Camp from Berlin-Ichthyosaur state park. They concluded that of the three species Camp described from among the remains, only the type and most abundant species ''S. popularis'' was valid. The other two species he named, ''S. silberlingi'' and ''S. mulleri'', were merely junior synonyms of ''S. popularis''.
* Motani, Rothschild, and Wahl found that ''Temnodontosaurus'' had eyes up to 10 inches in diameter, the largest known of any animal.
* Fernandez described the new ichthyosaur genus '' Mollesaurus'' from the Los Molles Formation of Argentina. He also described the species ''Mollesaurus periallus''.
* Chun Li described the new ichthyosaur genus ''Qianichthyosaurus'' from Guizhou Province, China.
* Lingham-Soliar proposed a model for ichthyosaur extinction whereby the evolution of fishes capable of greater swimming speeds during the Cretaceous favored ambush predators like plesiosaurs and the newly evolved mosasaurs over the ichthyosaurs, who succumbed to the competition.
* McGowan and Milner described the new species ''Leptonectes moorei''.
* Efimov described the new genus and species ''Undorosaurus gorodischensis''. Efimov also described the species that would later come to be known as ''Ophthalmosaurus yasikovi''.
* Li described the new genus and species ''Qianichthyosaurus zhoui''.
''March 6th''
* An article about ichthyosaurs called "Dinodolphin" by Kate Douglas was published in New Scientist magazine.
''September''
* Rothschild, Motani, and Wahl presented an abstract to a meeting of the Society for Vertebrate Paleontology. There they reported evidence that some ''Ophthalmosaurus'' suffered from Decompression sickness, the bends.
21st century
2000s
2000 in paleontology, 2000
* Ryosuke Motani published an article on ichthyosaurs in Scientific American that documented the discovery of ''Utatsusaurus''.
* Swiss watch maker Rolex honored Elizabeth Nicholls as a Rolex Laureate and bestowed on her a $100,000 stipend, covering much of the expenses generated by her field work.
* Sander expressed doubt as to whether or not ''Omphalosaurus'' was really an ichthyosaur. He also cast doubt on the idea that the limb bones from Spitsbergen referred to the genus by Carl Wiman actually belonged to the same kind of animal as the jaws discovered by J. C. Merriam that served as its type specimen.
* Maisch and Matzke erected the new genus ''Callawayia'' for the species ''Shastasaurus neoscapularis''. They also erected the new genus ''Phantomosaurus''.
* Yin and others described the new genus and species ''Guizhouichthyosaurus tangae''. They also described the species that would later come to be known as ''Shastasaurus liangae''.
'' May''
* Nathalie Bardet and Marta Fernandez erected the new genus ''Aegirosaurus'' to house the species ''Ichthyosaurus leptospondylus''. This was first new ichthyosaur to be described from the Solnhofen lithographic limestone in over 50 years prior to its naming. The type specimen preserved extensive soft tissue traces. Bardet and Fernandez reported the presence of tiny scales covering the animal. They disagreed with Martill, who claimed in 1995 that no evidence of a scaley covering existed in ichthyosaurs.
2001 in paleontology, 2001
* Maisch and Matzke recognized the genus ''Phalarodon'' as diagnostic rather than dubious. They referred the species "''Mixsaurus''" ''major'' of the German Muschelkalk to that genus, and noted that this represented the first report of ''Phalarodon'' in that locality.
* Nicholls and Manabe erected the new genus ''Metashastasaurus'' for the species ''Shastasaurus neoscapularis''. However, since the genus ''Callawayia'' had already been named for this species by Maisch and Matzke it was never accepted as valid by the scientific community. Nicholls and Manabe also reported that since field work began in the Pardonet Formation of British Columbia's Pink Mountains, 65 ichthyosaur specimens had been recovered.
* Orndorff and others interpreted the burial site of the famous ichthyosaurs of Berlin-Ichthyosaur State Park as a deep ocean shelf rather than a shallow coastal area. They also proposed a new reinterpretation of the animal's deaths, suggesting that the deposit may represent a school of ''Shonisaurus'' that was paralyzed by neurotoxins in the fish or shellfish that they ate before sinking to their death. They compared their hypothesis to modern mass whale deaths off the coast of New England. However, this interpretation is considered doubtful as there is no evidence linking mass whale deaths with consumption of poisonous sealife.
* Dino Frey and others reported the presence of ichthyosaur vertebrae in the collection of the Faculty of Geoscience in Linares, Mexico. They expressed interest in returning to Mexico to study the remains further and search for new finds.
* Paleontologist Ben Kear collaborated with radiographer George Kourlis to perform a CT scan of ''Platypterigius''. They found that its inner ear bones were so thick that they could not transmit sound vibrations and concluded animal must have been deaf. The scan also exposed the sensory structures inside its nose that allowed it to smell as well as an unusual system of "channels and grooves". The researchers found embryonic remains inside a ''Platypteriguis'' from Hughenden, Queensland, as well as the remains of the belemnitida, belemnites, fish, and turtle hatchlings it ate.
* Arkhangelsky described the species that would later come to be known as ''Brachypterygius alekseevi''.
2002 in paleontology, 2002
* Li and You described the new species ''Cymbospondylus asiaticus''. The known remains of this species consisted of two skulls excavated from the Late Triassic Falang Formation of Guizhou Province, China. This was the first known example of the genus in Asia.
* Perkins speculated that the channels and grooves uncovered by Kear and Kourlis in the skull of Platypterigius via CT scan formed an electrosensory system similar to those possessed by modern sharks.
* Peter Doyle gave an interview to ''New Scientist'' magazine discussing acid-etched belemnite shells, that he believed originated as ichthyosaur vomit.
* Ryosuke Motani compared the bodies and hydrodynamics of aquatic life with a "thunniform" body plan like dolphins, lamnid sharks, tunas, and the ichthyosaur ''Stenopterygius quadriscissus''. He modeled the motions of a thunniform body and its interaction with the water finding that, contrary to the conclusions of previous research, the tail fin of a thunniform animal was evolved to enable cruising at large body sizes rather than for "propuslive efficiency". He hypothesized that the similar body plans shared between ichthyosaurs and tunas suggest similar high swimming speeds and metabolic rates.
* Stuart Humphries and Graeme Ruxton published a study on the eyes of ''Ophthalmosaurus''. They calculated its eyes to be two and half to four times as light sensitive as the modern elephant seal. Since elephant seals can themselves dive thousands of feet deep, the researchers concluded that increased light sensitivity in deep water was probably not the only evolutionary pressure behind the evolution of large eyes in ''Ophthalmosaurus''. Its large eyes would have given it exceptionally clear vision as well, which would have been useful for tracking small prey and in possible social behavior.
* Thegarten Lingham-Soliar argued against Nathalie Bardet's attribution of ichthyosaur extinction to the loss of their preferred food sources in the Cenomanian-Turonian extinction event expanded on his own 1999 attribution of ichthyosaur extinction to biotic factors. During the Cretaceous the evolution of many fish groups trended towards faster swimming body types, making them harder to hunt for adult ichthyosaurs and harder to escape from for newborn ichthyosaurs. This ecological scene favored ambush predators like plesiosaurs and the newly evolved mosasaurs over the ichthyosaurs, who succumbed to the competition.
* Maisch and Lehmann described the new species ''Omphalosaurus peyeri''.
2003 in paleontology, 2003
* Elizabeth Nicholls, Chen Wei (ichthyologist), Chen Wei, and Makato Manabe published an extensive description of a new, complete specimen of ''Qianichthyosaurus'' from Guizhou. They observed that it was very similar to the genus ''Toretocnemus'' from California and concluded that similar ichthyosaur faunas spanned the Pacific Ocean during the Late Triassic.
* Maisch and Matzke described the new genus and species ''Quasianosteosaurus vikinghoegdai''.
2004 in paleontology, 2004
* Schmitz and others described the species that would later come to be known as ''Phalarodon callawayi''.
* Nicholls and Manabe described the species that would later come to be known as ''Shastasaurus sikkaniensis''.
2006 in paleontology, 2006
* Maxwell and Caldwell described the new genus and species ''Maiaspondylus lindoei''.
* Jiang and others described the species that would later come to be known as ''Barracudasauroides panxianensis''.
* Fröbisch, Sander and Rieppel described the new species ''Cymbospondylus nichollsi''.
2007 in paleontology, 2007
* Chen, Cheng, and Sander described the species that would later come to be known as ''Guizhouichthyosaurus wolongangensis''.
2008 in paleontology, 2008
* Maisch described the new genus and species ''Hauffiopteryx typicus''.
* Arkhangelsky and others described the new species ''Platypterygius ochevi''.
* Jiang and others described the new genus and species ''Xinminosaurus catactes''.
2010s
2010 in paleontology, 2010
* Maxwell described the new genus ''Arthropterygius''
* Druckenmiller and Maxwell described the new genus and species ''Athabascasaurus bitumineus''
* Maisch described the new genus ''Barracudasauroides'' and new species ''Omphalosaurus merriami''
* Chen and Cheng described the new species ''Mixosaurus xindianensis''
2011 in paleontology, 2011
* Fischer and others described the new genus and species ''Sveltonectes insolitus''
2012 in paleontology, 2012
* Fischer and others described the new genus and species ''Acamptonectes densus''
* Druckenmiller and others described the new genera and species ''Cryopterygius kristiansenae'' and ''Palvennia hoybergeti''
* Maxwell, Fernández, and Schoch described the new species ''Stenopterygius aaleniensis''
* Martin and others described the new species ''Temnodontosaurus azerguensis''
2013 in paleontology, 2013
* Chen and others described the new species ''Chaohusaurus, Chaohusaurus zhangjiawanensis''
* Cuthbertson, Russell and Anderson described the new genus and species ''Gulosaurus helmi''
* Fischer and others described the new genus and species ''Malawania anachronus''
* Fischer and others described the new genus and species ''Leninia stellans''.
* Fröbisch and others described the new genus and species ''Thalattoarchon saurophagis''
2014 in paleontology, 2014
* Roberts and others described the new genus and species ''Janusaurus lundi''
* Fischer and others described the new genus and species ''Sisteronia seeleyi''
* Arkhangelsky and Zverkov described the new species ''Undorosaurus, Undorosaurus trautscholdi''
* Fischer and others described the new genus and species ''Leninia stellans''.
* Anatomy, taxonomy and phylogenetic relationships of Cretaceous Ophthalmosauridae, ophthalmosaurids ''Simbirskiasaurus birjukovi'' and ''Pervushovisaurus bannovkensis'' are reevaluated by Fischer ''et al.'' (2014).
2015 in paleontology, 2015
* Motani and others described the new genus and species ''Cartorhynchus lenticarpus''
* Brusatte and others described the new genus and species ''Dearcmhara shawcrossi''
* Chen and others described the new genus and species ''Eretmorhipis carrolldongi''
* Lomax and Massare described the new species ''Ichthyosaurus anningae''
* Maxwell and others described the new genus and species ''Muiscasaurus catheti''
* A study of phylogenetic relationships of ichthyopterygians is published by Ji ''et al.'' (2015); the authors introduced a new name, Grippioidea, for the clade containing the last common ancestor of '' Utatsusaurus hataii'' and ''Grippia longirostris'', and all its descendants.
* An exceptionally large ichthyosaur Radius (bone), radius, possibly belonging to a member of Shastasauridae (which, if confirmed, would indicate that members of the family survived until
Early Jurassic
The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma  ...
) is described from the Hettangian Blue Lias Formation (south Wales, United Kingdom) by Martin ''et al.'' (2015).
2016 in paleontology, 2016
* Jiang and others described the new genus and species ''Sclerocormus parviceps''.
* Lomax described the new genus and species ''Wahlisaurus''.
* Lomax and Massare described the new species ''Ichthyosaurus larkini'' and ''Ichthyosaurus somersetensis, I. somersetensis''.
* Tyborowski described the new species ''Cryopterygius kielanae''.
* A study of taxonomic richness, disparity and evolutionary rates of ichthyosaurs throughout the Cretaceous period is published by Fischer ''et al.'' (2016).
* A restudy of ''" Platypterygius" campylodon'' is published by Fischer (2016), who transfers this species to the genus ''Pervushovisaurus''.
* A revision of the ichthyosaur material of the British Middle and Late Jurassic referable to '' Ophthalmosaurus icenicus'' is published by Moon & Kirton (2016).
2017 in paleontology, 2017
* Delsett and others described the new genus and species ''Keilhauia nui''.
* Paparella and others described the new genus and species ''Gengasaurus''.
* Lomax, Massare and Mistry described the new genus and species ''Protoichthyosaurus applebyi''.
* A study on the emergence date and changes of the evolutionary rate during the Ichthyosauromorpha, ichthyosauromorph evolution is published by Motani ''et al.'' (2017).
* A jaw fragment of a member of the genus ''
Omphalosaurus
''Omphalosaurus'' (from the Greek root "Button Lizard", for their button-like teeth) is an extinct genus of marine reptile from the Early Triassic to Middle Triassic, thought to be in the order of Ichthyosauria. Most of what is known about ''O ...
'' is described from the Middle Triassic (Anisian) Karchowice Formation (Poland) by Wintrich, Hagdorn & Sander (2017), representing the first record of ''Omphalosaurus'' from shallow marine carbonates and from the Muschelkalk facies.
* Description of three nearly complete and well-preserved skulls of '' Chaohusaurus chaoxianensis'', revealing new information on the skull anatomy of the species, is published by Zhou ''et al.'' (2017).
* A specimen of ''Ichthyosaurus somersetensis'' containing an embryo, representing the largest unequivocal specimen of a member of the genus ''Ichthyosaurus'', is described from the Lower Jurassic (lower Hettangian) Blue Lias Formation (United Kingdom) by Lomax & Sachs (2017).
* Plet ''et al.'' (2017) report the presence of Red blood cell, red and white blood cell-like structures as well as platelet-like structures, collagen and cholesterol in a vertebra of a member of the genus ''
Stenopterygius
''Stenopterygius'' is an extinct genus of thunnosaur ichthyosaur known from Europe (England, France, Germany, Luxembourg and Switzerland). This genus of ichthyosaur was about long and weighed .Huene F. von 1939. Ein ganzes Ichthyosaurier-Skelett ...
'' from Toarcian Posidonia Shale (
Germany
Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
).
* An articulated skeleton of an Ophthalmosauridae, ophthalmosaurid ichthyosaur is described from the Upper Jurassic (Kimmeridgian) Katrol Formation (India) by Prasad ''et al.'' (2017).
2018 in paleontology, 2018
* A study aiming to identify sexual dimorphism, taxonomic variation and individual variation among the specimens of '' Chaohusaurus chaoxianensis'' is published by Motani ''et al.'' (2018).
* A survey of the form and distribution of pathological structures in the skeletons of ichthyosaurs is published by Pardo-Pérez ''et al.'' (2018).
* A study on the microanatomy of vertebral centra of ichthyosaurs, aiming to establish whether there is any variation between the primitive and the most derived forms, is published by Houssaye, Nakajima & Sander (2018).
* Humerus, Humeri of ''Pessopteryx nisseri'' and vertebrae referred to the genus '' Cymbospondylus'' are described from the Lower Triassic Vikinghøgda Formation (
Spitsbergen
Spitsbergen (; formerly known as West Spitsbergen; Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipelago in northern N ...
, Norway) by Engelschiøn ''et al.'' (2018).
* A large, isolated jaw fragment of a giant ichthyosaur is described from the Upper Triassic (Rhaetian) Westbury Mudstone Formation (United Kingdom) by Lomax ''et al.'' (2018), who also reinterpret some putative dinosaur limb bone shafts from the Upper Triassic of Aust Cliff as more likely to be ichthyosaur fossils.
* Ichthyosaur fossils, including an incomplete skeleton of a member of the genus '' Leptonectes'', are described from the Lower Jurassic (Pliensbachian) of Asturias (Spain) by Fernández, Piñuela & García-Ramos (2018).
* Remains of ichthyosaur embryos, still situated within a fragment of the rib-cage of the parent animal, are described from the Lower Jurassic (Toarcian) Whitby Mudstone Formation (United Kingdom) by Boyd & Lomax (2018).
* Ichthyosaur remains from the Lower Cretaceous Agrio Formation (Neuquén Basin, Argentina) are described by Lazo ''et al.'' (2018).
* New specimen of ''
Phalarodon
''Phalarodon'' is an extinct genus of ichthyosaur. Its remains have been found in China, North America, and Spitsbergen. It measured between and weighed more than .
See also
* List of ichthyosaurs
* Timeline of ichthyosaur research
This time ...
fraasi'' is described from the Middle Triassic Botneheia Formation (Svalbard, Norway) by Økland ''et al.'' (2018).
* Redescription of the relocated holotype of '' Suevoleviathan integer'' is published by Maxwell (2018), who considers the species ''Suevoleviathan disinteger'' to be a Synonym (taxonomy), junior synonym of ''S. integer''.
* A study on specimens of '' Temnodontosaurus'' from the
Early Jurassic
The Early Jurassic Epoch (in chronostratigraphy corresponding to the Lower Jurassic Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic-Jurassic extinction event, 201.3 Ma  ...
of southern
Germany
Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
, aiming to document the types of pathologies observed in the skeletons of specimens assigned to this genus, is published by Pardo-Pérez ''et al.'' (2018).
* Four isolated partial skulls from the Lower Jurassic of the United Kingdom, previously identified as '' Ichthyosaurus communis'', are assigned to the species ''Protoichthyosaurus prostaxalis'' and ''P. applebyi'' by Lomax & Judith Massare, Massare (2018), providing new information on the anatomy of these taxa.
* A reassessment of ''Ichthyosaurus communis'' and ''I. intermedius'' is published by Massare & Lomax (2018), who consider the latter species to be a Synonym (taxonomy), junior synonym of the former.
* A study on the cellular and molecular composition of integumental tissues in an exceptionally preserved specimen of ''
Stenopterygius
''Stenopterygius'' is an extinct genus of thunnosaur ichthyosaur known from Europe (England, France, Germany, Luxembourg and Switzerland). This genus of ichthyosaur was about long and weighed .Huene F. von 1939. Ein ganzes Ichthyosaurier-Skelett ...
'' is published by Lindgren ''et al.'' (2018).
* New specimen of ''Palvennia hoybergeti'', providing new information on the anatomy of this species, is described from the Upper Jurassic Slottsmøya Member of the Agardhfjellet Formation (
Spitsbergen
Spitsbergen (; formerly known as West Spitsbergen; Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipelago in northern N ...
, Norway) by Delsett ''et al.'' (2018).
* A revision of British ichthyosaur taxa of the Late Jurassic is published by Moon & Kirton (2018).
2019 in paleontology, 2019
* Two new specimens of ''Eretmorhipis carrolldongi'', revealing superficial Convergent evolution, convergence with the modern platypus, are described from the Lower Triassic Jialingjiang Formation (China) by Cheng ''et al.'' (2019).
* A study on the phylogenetic relationships of ichthyosaurs will be published by Moon (2019).
* A study on the evolution of ichthyosaur body forms and on its impact on the energy demands of ichthyosaur swimming is published by Gutarra ''et al.'' (2019).
* A study on the flexibility and function of ichthyosaur tails, as indicated by comparisons with shark tails, is published by Crofts, Shehata & Flammang (2019).
* A study on the effects of methodology, missing data and exceptional preservation of fossil specimens in lagerstätten on known morphological diversity of fossil animals, as indicated by fossil record of ichthyosaurs, is published by Flannery Sutherland ''et al.'' (2019).
* Second specimen of ''Wahlisaurus massarae'' is reported from a quarry in
Somerset
( en, All The People of Somerset)
, locator_map =
, coordinates =
, region = South West England
, established_date = Ancient
, established_by =
, preceded_by =
, origin =
, lord_lieutenant_office =Lord Lieutenant of Somerset
, lor ...
(United Kingdom), from the base of the Blue Lias Formation (Triassic–Jurassic boundary) by Lomax, Evans & Carpenter (2019), extending known geographic and stratigraphic range of the species.
* Partial skeleton of a large ichthyosaur from the Lower Jurassic (Sinemurian) of Warwickshire, England is described by Lomax, Porro & Larkin (2019), who assign this specimen to the species ''Protoichthyosaurus prostaxalis''.
* A neonate specimen of '' Ichthyosaurus communis'' is described by Lomax ''et al.'' (2019).
* A study on the variation of the hindfin Morphology (biology), morphology in the specimens of ''Ichthyosaurus'' and on its taxonomic utility is published by Massare & Lomax (2019).
* A study on the bone microstructure of the skeleton of a specimen of ''
Stenopterygius
''Stenopterygius'' is an extinct genus of thunnosaur ichthyosaur known from Europe (England, France, Germany, Luxembourg and Switzerland). This genus of ichthyosaur was about long and weighed .Huene F. von 1939. Ein ganzes Ichthyosaurier-Skelett ...
quadriscissus'' from the Lower Jurassic Posidonia Shale (
Germany
Germany, officially the Federal Republic of Germany (FRG),, is a country in Central Europe. It is the most populous member state of the European Union. Germany lies between the Baltic and North Sea to the north and the Alps to the sou ...
) is published by Anderson ''et al.'' (2019).
*A study on ontogentic variation in the braincase of ''
Stenopterygius
''Stenopterygius'' is an extinct genus of thunnosaur ichthyosaur known from Europe (England, France, Germany, Luxembourg and Switzerland). This genus of ichthyosaur was about long and weighed .Huene F. von 1939. Ein ganzes Ichthyosaurier-Skelett ...
'' is published by Miedema and Maxwell
* A study on the anatomy of an Ophthalmosauridae, ophthalmosaurid rostrum fragment from the Upper Jurassic (Oxfordian (stage), Oxfordian) in the Morawica quarry in the Świętokrzyskie Mountains (Poland), and on its implications for reconstructing the internal Morphology (biology), morphology of the ophthalmosaurid cranial region and inferring the functional adaptations and palaeoecology of these reptiles, will be published by Tyborowski, Skrzycki & Dec (2019).
* A revision of the type series of all three species of ''Undorosaurus'' is published by Zverkov & Efimov (2019).
* A study on the taxonomy and phylogeny of ichthyosaurs belonging to the genus ''Arthropterygius'' is published by Zverkov & Prilepskaya (2019).
* New fossil remains of '' Platypterygius sachicarum'' (a new skull and associated postcranial remains of upper Barremian age) are described from Villa de Leyva, Colombia by Maxwell ''et al.'' (2019), representing the first documented postcranial remains of this species.
* Campos, Fernández and Herrera described the new species ''Arthropterygius thalassonotus''
* Huang and others described the new species ''Chaohusaurus brevifemoralis''
2020 in paleontology, 2020
* Barrientos Lara and others described the new genus and species ''Acuetzpalin carranzai''.
See also
* History of paleontology
** Timeline of paleontology
* Timeline of plesiosaur research
* Timeline of mosasaur research