''Tikisuchus'' is an extinct
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
rauisuchid archosauromorph
Archosauromorpha ( Greek for "ruling lizard forms") is a clade of diapsid reptiles containing all reptiles more closely related to archosaurs (such as crocodilians and dinosaurs, including birds) rather than lepidosaurs (such as tuataras, li ...
. It is known from the
Late Triassic
The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch ...
Tiki Formation
The Tiki Formation is a Late Triassic (Carnian to Norian) geologic formation in Madhya Pradesh, northern India.[Shahdol District
Shahdol District is a district of Madhya Pradesh state in east central India. It old name was Matsya raj or Virat kingdom in the name of great king Maharaja Virata. With a total area of and a population of 10,66,063. Shahdol is an important dis ...]
of central
India
India, officially the Republic of India ( Hindi: ), is a country in South Asia. It is the seventh-largest country by area, the second-most populous country, and the most populous democracy in the world. Bounded by the Indian Ocean on the ...
and was the first rauisuchid to have been found in Asia. The
horizon
The horizon is the apparent line that separates the surface of a celestial body from its sky when viewed from the perspective of an observer on or near the surface of the relevant body. This line divides all viewing directions based on whether ...
from which remains have been found is
Carnian
The Carnian (less commonly, Karnian) is the lowermost stage of the Upper Triassic Series (or earliest age of the Late Triassic Epoch). It lasted from 237 to 227 million years ago (Ma). The Carnian is preceded by the Ladinian and is followe ...
in age. The
type species
In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...
is ''T. romeri'', named in honor of American paleontologist
Alfred Romer
Alfred Sherwood Romer (December 28, 1894 – November 5, 1973) was an American paleontologist and biologist and a specialist in vertebrate evolution.
Biography
Alfred Romer was born in White Plains, New York, the son of Harry Houston Romer an ...
. Romer was present at the Tiki locality during the excavation of the fossil, but died before the description of the genus in 1987. ''Tikisuchus'' is known only from one specimen, called
ISI
ISI or Isi may refer to:
Organizations
* Intercollegiate Studies Institute, a classical conservative organization focusing on college students
* Ice Skating Institute, a trade association for ice rinks
* Indian Standards Institute, former name of ...
R 305, which consists of the skull and some postcranial elements of a young individual.
Description
Compared to other rausuchids, the skull of ''Tikisuchus'' was very large. The skull's length is around 40% of the length of the presacral area between the head and the
sacrum
The sacrum (plural: ''sacra'' or ''sacrums''), in human anatomy, is a large, triangular bone at the base of the spine that forms by the fusing of the sacral vertebrae (S1S5) between ages 18 and 30.
The sacrum situates at the upper, back part ...
. The skull is deep, being wide at the back with a narrow rostrum. The teeth are large, recurved and serrated. Like other rauisuchids, it has rows of
osteoderm
Osteoderms are bony deposits forming scales, plates, or other structures based in the dermis. Osteoderms are found in many groups of extant and extinct reptiles and amphibians, including lizards, crocodilians, frogs, temnospondyls (extinc ...
s, or bony
scute
A scute or scutum (Latin: ''scutum''; plural: ''scuta'' "shield") is a bony external plate or scale overlaid with horn, as on the shell of a turtle, the skin of crocodilians, and the feet of birds. The term is also used to describe the anterior po ...
s, along its back. There are two rows of osteoderms. Each osteoderm is rectangular in shape and imbricates, or articulates tightly, with the ones around it. In other rauisuchids, the osteoderms are leaf-shaped rather than rectangular.
[
]
Paleobiology
Many other tetrapods were found in association with ''Tikisuchus'' representing a diverse Carnian paleofauna. Tetrapods from the Tiki site include ''Paleorhinus
''Paleorhinus'' (Greek: ''"Old Nose"'') is an extinct genus of widespread basal phytosaur known from the Late Triassic (late Carnian stage). The genus was named in 1904 based on the type species ''Paleorhinus bransoni'', which is known from Wyom ...
'', a phytosaur
Phytosaurs (Φυτόσαυροι in greek) are an extinct group of large, mostly semiaquatic Late Triassic archosauriform reptiles. Phytosaurs belong to the order Phytosauria. Phytosauria and Phytosauridae are often considered to be equivalent gr ...
, ''Metoposaurus
''Metoposaurus'' meaning "front lizard" is an extinct genus of stereospondyl temnospondyl amphibian, known from the Late Triassic of Germany, Italy, Poland, and Portugal. https://www.app.pan.pl/archive/published/app47/app47-535.pdf This mostly ...
'', a temnospondyl
Temnospondyli (from Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order of small to giant tetrapods—often considered primitive amphibians—that flourished worldwide during the Carb ...
, and ''Paradapedon
''Hyperodapedon'' is a genus of rhynchosaurs (beaked, archosaur-like reptiles) from the Late Triassic period ( Carnian stage). Fossils of the genus have been found in Africa, Asia, Europe and North and South America. Its first discovery and na ...
'', a rhynchosaur
Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaur ...
. The Tiki fauna is similar to that of the German
German(s) may refer to:
* Germany (of or related to)
**Germania (historical use)
* Germans, citizens of Germany, people of German ancestry, or native speakers of the German language
** For citizens of Germany, see also German nationality law
**Ger ...
Keuper
The Keuper is a lithostratigraphic unit (a sequence of rock strata) in the subsurface of large parts of west and central Europe. The Keuper consists of dolomite, shales or claystones and evaporites that were deposited during the Middle and Late ...
.[
]Theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s were also present in the Tiki Formation. Both ''Tikisuchus'' and the theropods were large terrestrial predators, and, having been found at the same locality, likely came in close contact with one another. The similar lifestyles of the two carnivores may have resulted in competition for the same food sources. Possible prey would have included rhynchosaur
Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaur ...
s, trilophosaurs, dicynodont
Dicynodontia is an extinct clade of anomodonts, an extinct type of non-mammalian therapsid. Dicynodonts were herbivorous animals with a pair of tusks, hence their name, which means 'two dog tooth'. Members of the group possessed a horny, typica ...
s, and aetosaur
Aetosaurs () are heavily armored reptiles belonging to the extinct order Aetosauria (; from Greek, (aetos, "eagle") and (, "lizard")). They were medium- to large-sized omnivorous or herbivorous pseudosuchians, part of the branch of archosaurs ...
s. The authors of the original description of ''Tikisuchus'', Sankar Chatterjee
Sankar Chatterjee (born May 28, 1943) is a paleontologist, and is the Paul W. Horn Professor of Geosciences at Texas Tech University and Curator of Paleontology at the Museum of Texas Tech University. He earned his Ph. D. from the University of ...
and Pranab Majumdar, suggested that competition between ''Tikisuchus'' and theropods was low because of abundant food resources and stabilized ecological interactions. Chatterjee and Majumdar thought that there was an "ecological balance" during the Carnian stage on the basis of little change in the paleofauna of the time.[ They considered the ]paleoclimate
Paleoclimatology ( British spelling, palaeoclimatology) is the study of climates for which direct measurements were not taken. As instrumental records only span a tiny part of Earth's history, the reconstruction of ancient climate is important t ...
to have been warm with seasonal wet and dry seasons conducive to the growth of tropical forests. At the end of the Carnian, however, the authors claimed that many prey resources became extinct and the forest environment was replaced by a more open environment. The limited resources would have heightened competition between theropods and rauisuchids like ''Tikisuchus''. Chatterjee and Majumdar considered theropods to be agile pursuit predators while rauisuchids were considered slow ambush predators. Therefore, they suggested that theropods, which were more suited to living in an open environment, outcompeted rauisuchids at the end of the Triassic to become the dominant large land carnivores by the beginning of the Jurassic.[ However, more recent studies suggest that dinosaurs gained dominance only after the Triassic-Jurassic extinction event in a case of opportunism with no other large archosaurs such as rauisuchids to compete with.]
References
{{Taxonbar, from=Q3281023
Rauisuchids
Prehistoric reptiles of Asia
Fossil taxa described in 1987
Prehistoric pseudosuchian genera