Thomas Richard Holtz Jr. (born September 13, 1965) is an American vertebrate
palaeontologist
Paleontology (), also spelled palaeontology or palæontology, is the scientific study of life that existed prior to, and sometimes including, the start of the Holocene epoch (roughly 11,700 years before present). It includes the study of fossi ...
, author, and principal lecturer at the
University of Maryland
The University of Maryland, College Park (University of Maryland, UMD, or simply Maryland) is a public university, public Land-grant university, land-grant research university in College Park, Maryland. Founded in 1856, UMD is the Flagship un ...
's Department of Geology. He has published extensively on the
phylogeny
A phylogenetic tree (also phylogeny or evolutionary tree Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, MA.) is a branching diagram or a tree showing the evolutionary relationships among various biological spe ...
,
morphology
Morphology, from the Greek and meaning "study of shape", may refer to:
Disciplines
*Morphology (archaeology), study of the shapes or forms of artifacts
*Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...
,
ecomorphology Ecomorphology or ecological morphology is the study of the relationship between the ecological role of an individual and its morphological adaptations. The term "morphological" here is in the anatomical context. Both the morphology and ecology exh ...
, and locomotion of terrestrial predators, especially on
tyrannosaurid
Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coelurosaurian theropod dinosaurs that comprises two subfamilies containing up to thirteen genera, including the eponymous ''Tyrannosaurus''. The exact number of genera ...
s and other
theropod
Theropoda (; ), whose members are known as theropods, is a dinosaur clade that is characterized by hollow bones and three toes and claws on each limb. Theropods are generally classed as a group of saurischian dinosaurs. They were ancestrally ...
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
s.
[Holtz, ''Thomas R. Holtz Jr.''.] He wrote the book ''
Dinosaurs
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
'' and is the author or co-author of the chapters "Saurischia", "Basal Tetanurae", and "Tyrannosauroidea" in the second edition of ''
The Dinosauria
''The Dinosauria'' is an extensive book on dinosaurs, compiled by David B. Weishampel, Peter Dodson, and Halszka Osmólska. It has been published in 2 editions, with the first edition published in 1990, consisting of material from 23 scientists.Be ...
''. He has also been consulted as a scientific advisor for the ''
Walking With Dinosaurs
''Walking with Dinosaurs'' is a 1999 six-part nature documentary television miniseries created by Tim Haines and produced by the BBC Science Unit the Discovery Channel and BBC Worldwide, in association with TV Asahi, ProSieben and France 3. ...
'' BBC series as well as the
Discovery
Discovery may refer to:
* Discovery (observation), observing or finding something unknown
* Discovery (fiction), a character's learning something unknown
* Discovery (law), a process in courts of law relating to evidence
Discovery, The Discover ...
special ''
When Dinosaurs Roamed America
''When Dinosaurs Roamed America'' (sometimes shortened to ''When Dinosaurs Roamed'' outside of the U.S.) is a two-hour American television program (produced in the style of a traditional nature documentary) that first aired on the Discovery Channe ...
'', and has appeared in numerous documentaries focused on prehistoric life, such as ''
Jurassic Fight Club'' on
History
History (derived ) is the systematic study and the documentation of the human activity. The time period of event before the History of writing#Inventions of writing, invention of writing systems is considered prehistory. "History" is an umbr ...
and ''
Monsters Resurrected
''Monsters Resurrected'' is an American documentary television series that premiered on September 13, 2009, on the Discovery Channel. The program reconstructs extinct animals of both Mesozoic and Cenozoic. It is also called ''Mega Beasts''.
F ...
'', ''
Dinosaur Revolution
''Dinosaur Revolution'' is a four-part American nature documentary produced by Creative Differences. It utilizes computer-generated imagery to portray dinosaurs and other animals from the Mesozoic era. The program was originally aired on the Disc ...
'' and ''
Clash of the Dinosaurs
''Clash of the Dinosaurs'' is a four-part television mini-series produced by Dangerous LTD for Discovery Channel. The show premiered on December 6, 2009, with the first two episodes scheduled back-to-back.
''Clash of the Dinosaurs'' was poorly r ...
'' on Discovery.
Holtz is also the director of the Science and Global Change Program within the
College Park Scholars
The University of Maryland, College Park (University of Maryland, UMD, or simply Maryland) is a public land-grant research university in College Park, Maryland. Founded in 1856, UMD is the flagship institution of the University System of Mary ...
living-learning community at the University of Maryland, College Park.
Early life
Holtz was born in California. He has stated that he wanted to be a dinosaur as a child, but upon learning that it is impossible to turn into a dinosaur he shifted his goals to study them. He attended
Johns Hopkins University
Johns Hopkins University (Johns Hopkins, Hopkins, or JHU) is a private research university in Baltimore, Maryland. Founded in 1876, Johns Hopkins is the oldest research university in the United States and in the western hemisphere. It consiste ...
, where he met his future wife. He then attended Yale, where he meet
John Ostrom
John Harold Ostrom (February 18, 1928 – July 16, 2005) was an American paleontologist who revolutionized modern understanding of dinosaurs in the 1960s.
As first proposed by Thomas Henry Huxley in the 1860s, Ostrom showed that dinosaurs were ...
who served as his academic advisor.
Theories
Holtz has come up with several new theories and hypotheses about the dinosaurs' classification. For example, he coined the terms
Maniraptoriformes
Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptorans. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of ''Ornithomimus'' and birds, ...
and
Arctometatarsus
An arctometatarsalian organism is one in which the proximal part of the middle metatarsal is pinched between the surrounding metatarsals. The trait appears to be highly homoplastic, common in certain sorts of dinosaurs accustomed to running (among ...
. He also proposed two classification systems for theropods. The first is the clade
Arctometatarsalia
Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and Nort ...
, made up of
tyrannosauroids
Tyrannosauroidea (meaning 'tyrant lizard forms') is a superfamily (or clade) of coelurosaurian theropod dinosaurs that includes the family Tyrannosauridae
Tyrannosauridae (or tyrannosaurids, meaning "tyrant lizards") is a family of coeluros ...
,
ornithomimosaurs
Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and North ...
, and
troodontids
Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...
, because all of these
coelurosaurs
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, tyra ...
had pinched middle
metatarsal
The metatarsal bones, or metatarsus, are a group of five long bones in the foot, located between the tarsal bones of the hind- and mid-foot and the phalanges of the toes. Lacking individual names, the metatarsal bones are numbered from the med ...
bones in their feet. In this proposed classification system, the tyrannosauroids were supposedly
basal
Basal or basilar is a term meaning ''base'', ''bottom'', or ''minimum''.
Science
* Basal (anatomy), an anatomical term of location for features associated with the base of an organism or structure
* Basal (medicine), a minimal level that is nec ...
to a clade known as
Bullatosauria
Maniraptoriformes is a clade of dinosaurs with pennaceous feathers and wings that contains ornithomimosaurs and maniraptorans. This group was named by Thomas Holtz, who defined it as "the most recent common ancestor of ''Ornithomimus'' and birds, ...
, which was made up of the
Troodontidae
Troodontidae is a clade of bird-like theropod dinosaurs. During most of the 20th century, troodontid fossils were few and incomplete and they have therefore been allied, at various times, with many dinosaurian lineages. More recent fossil disco ...
and the
Ornithomimosauria
Ornithomimosauria ("bird-mimic lizards") are theropod dinosaurs which bore a superficial resemblance to the modern-day ostrich. They were fast, omnivorous or herbivorous dinosaurs from the Cretaceous Period of Laurasia (now Asia, Europe and Nor ...
. These two groups were purported to form a
clade, because they both shared a common characteristic; which was a
skull capsule
The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, the ...
. However, later on, both of these classification systems were found to be
paraphyletic
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
, or "artificial", clades. For example, troodontids are now known to be deinonychosaurs, or "raptors", closely related to dromaeosaurids and birds. The discovery of basal tyrannosauroids, such as ''
Guanlong
''Guanlong'' (冠龍) is a genus of extinct proceratosaurid tyrannosauroid from the Late Jurassic of China. The taxon was first described in 2006 by Xu Xing ''et al.'', who found it to represent a new taxon related to ''Tyrannosaurus''. The ...
'', which lacked an arctometatarsus, also helped to disprove this theory. And, eventually, the "skull capsule" in troodontids and ornithomimosaurs was found to be an example of
convergent evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last com ...
, causing the clade Bullatosauria to be abandoned.

Holtz was also a key figure in the discovery that tyrannosauroids were not
carnosaur
Carnosauria is an extinct large group of predatory dinosaurs that lived during the Jurassic and Cretaceous periods. Starting from the 1990s, scientists have discovered some very large carnosaurs in the carcharodontosaurid family, such as ''Gig ...
s, as had been previously believed by most palaeontologists, but rather large
coelurosaur
Coelurosauria (; from Greek, meaning "hollow tailed lizards") is the clade containing all theropod dinosaurs more closely related to birds than to carnosaurs.
Coelurosauria is a subgroup of theropod dinosaurs that includes compsognathids, ty ...
s. One of the first scientists to theorize this, Holtz contributed greatly to the debunking of a
monophyletic
In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...
Carnosauria that contained Megalosauroidea, Allosauroidea and Tyrannosauroidea. However, fellow vertebrate paleontologist Oliver Rauhut is known to recover a monophyletic Carnosauria that contains both Megalosauroidea and Allosauroidea in many of his publications, although like most researcher he agrees that Tyrannosauroids are likely Coelurosaurians as based on the conclusions made by Holtz.
Selected publications
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Footnotes
References
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{{DEFAULTSORT:Holtz, Thomas R. Jr.
Living people
American paleontologists
University of Maryland, College Park faculty
1965 births