Thalassemys
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''Thalassemys'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
thalassochelydia Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including '' Eurysternum'', ''Plesiochelys'' and '' Thalassemys'' to the exclusion of '' Pel ...
n
turtle Turtles are reptiles of the order (biology), order Testudines, characterized by a special turtle shell, shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Crypt ...
from the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
of western and central
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. While the genus was originally named by Rütimeyer in 1859 for a large
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
and other associated fragments from the late
Kimmeridgian In the geologic timescale, the Kimmeridgian is an age in the Late Jurassic Epoch and a stage in the Upper Jurassic Series. It spans the time between 154.8 ±0.8 Ma and 149.2 ±0.7 Ma (million years ago). The Kimmeridgian follows the Oxfordian ...
of the Reuchenette Formation of
Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...
, although the taxon was not validly named until
1873 Events January * January 1 ** Japan adopts the Gregorian calendar. ** The California Penal Code goes into effect. * January 17 – American Indian Wars: Modoc War: First Battle of the Stronghold – Modoc Indians defeat the Unit ...
when Rütimeyer designated the type species ''T. hugii''. Rütimeyer also named ''T. gresslyi'' from the Reunchenette Formation in the same paper as ''T. hugii'', but it cannot be differentiated from the type material of ''T. hugii'' and is therefore a junior synonym. A large assemblage of shell and postcranial material from the Reunchenette was named as a species of '' Eurysternum'', ''E. ignoratum'', by Bräm in
1965 Events January–February * January 14 – The First Minister of Northern Ireland and the Taoiseach of the Republic of Ireland meet for the first time in 43 years. * January 20 ** Lyndon B. Johnson is Second inauguration of Lynd ...
. While originally distinguished based on the presence of fontanelles on the
plastron The turtle shell is a shield for the ventral and dorsal parts of turtles (the Order (biology), order Testudines), completely enclosing all the turtle's vital organs and in some cases even the head. It is constructed of modified bony elements such ...
, the feature was later identified on ''T. hugii'' and ''E. ignoratum'' was designated a junior synonym. Additional material from the
Kimmeridge Clay The Kimmeridge Clay is a sedimentary rock, sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation (geology), form ...
of the
United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...
has also been referred to ''T. hugii''. The second valid species of ''Thalassemys'' is ''T. marina'', named in
1903 Events January * January 1 – Edward VII is proclaimed Emperor of India. * January 10 – The Aceh Sultanate was fully annexed by the Dutch forces, deposing the last sultan, marking the end of the Aceh War that have lasted for al ...
by Fraas for a partial carapace and half a plastron from the late Kimmeridgian deposits of Schnaitheim,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. It was suggested to be a species of the related taxon '' Eurysternum'' by Maisch in
2001 The year's most prominent event was the September 11 attacks against the United States by al-Qaeda, which Casualties of the September 11 attacks, killed 2,977 people and instigated the global war on terror. The United States led a Participan ...
, who referred additional material from Schnaitheim to the species. In
2003 2003 was designated by the United Nations as the International Year of Fresh water, Freshwater. In 2003, a Multi-National Force – Iraq, United States-led coalition 2003 invasion of Iraq, invaded Iraq, starting the Iraq War. Demographic ...
, it was suggested that ''T. marina'' was a synonym of the related turtle '' Palaeomedusa'', which was based on an unknown but restored region of the original carapace. ''Thalassemys heusseri'' was validly named in
1924 Events January * January 12 – Gopinath Saha shoots Ernest Day, whom he has mistaken for Sir Charles Tegart, the police commissioner of Calcutta, and is arrested soon after. * January 20–January 30, 30 – Kuomintang in Ch ...
by Oertel for a partial carapace from the Kimmeridgian or
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
of Holzen, Germany, a different spelling from the invalid name "T. heuseri" he used previously in
1815 Events January * January 2 – Lord Byron marries Anna Isabella Milbanke in Seaham, county of Durham, England. * January 3 – Austria, Britain, and Bourbon-restored France form a secret defensive alliance treaty against Pr ...
. As the type specimen was lost, the species cannot be distinguished from others of the genus, and is therefore a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''. In addition to ''E. ignoratum'', Bräm also named the species ''Thalassemys moseri'' in 1965 for a partial carapace from the Reunchenette Formation. An additional shell and also a skull were referred to the taxon in
1980 Events January * January 4 – U.S. President Jimmy Carter proclaims a United States grain embargo against the Soviet Union, grain embargo against the USSR with the support of the European Commission. * January 6 – Global Positioning Sys ...
by Rieppel, who suggested that the species should be included in ''
Plesiochelys ''Plesiochelys'' ("near turtle") is a genus of late Jurassic European and Asian turtle. The type species is ''Plesiochelys etalloni''. Fossil records have discovered ''Plesiochelys bigleri'' and ''Plesiochelys etalloni'' from the Kimmeridge Clay ...
'' rather than ''Thalassemys''. This skull was given the name '' Jurassichelon'' in
2015 2015 was designated by the United Nations as: * International Year of Light * International Year of Soil __TOC__ Events January * January 1 – Lithuania officially adopts the euro as its currency, replacing the litas, and becomes ...
, leaving the species ''T. moseri'', based on other material, in an uncertain position. Further assessment of the material by Anquetin and colleagues in
2017 2017 was designated as the International Year of Sustainable Tourism for Development by the United Nations General Assembly. Events January * January 1 – Istanbul nightclub shooting: A gunman dressed as Santa Claus opens fire at the ...
concluded that ''T. moseri'' was distinct from the species ''J. oleronensis'' named for the skull, but was representative of the same genus. ''Thalassemys bruntrutana'' was described in 2015 by Püntener and colleagues for an almost complete carapace and plastron from a lower member of the Reunchenette Formation than other species, with referred material also from the British
Kimmeridge Clay The Kimmeridge Clay is a sedimentary rock, sedimentary deposit of fossiliferous marine clay which is of Late Jurassic to lowermost Cretaceous age and occurs in southern and eastern England and in the North Sea. This rock formation (geology), form ...
of the
Isle of Purbeck The Isle of Purbeck is a peninsula in Dorset, England. It is bordered by water on three sides: the English Channel to the south and east, where steep cliffs fall to the sea; and by the marshy lands of the River Frome, Dorset, River Frome and Poo ...
. While the genus ''Thalassemys'' was placed in the family Thalassemydidae, it represents the only definitive member of the family due to the uncertain relationships of other members of
Thalassochelydia Thalassochelydia is a clade of extinct marine turtles from the Late Jurassic and earliest Cretaceous of Europe and South America. The group is defined as including '' Eurysternum'', ''Plesiochelys'' and '' Thalassemys'' to the exclusion of '' Pel ...
, primarily due to the lack of cranial material in ''Thalassemys''.


References

* {{Taxonbar, from=Q7709531 Thalassochelydia Prehistoric turtle genera Kimmeridgian genera Late Jurassic turtles Late Jurassic reptiles of Europe Jurassic England Fossils of England Jurassic Germany Fossils of Germany Jurassic Switzerland Fossils of Switzerland Fossil taxa described in 1873