Tetrabaena
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''Tetrabaena'' is a
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
genus of
green algae The green algae (: green alga) are a group of chlorophyll-containing autotrophic eukaryotes consisting of the phylum Prasinodermophyta and its unnamed sister group that contains the Chlorophyta and Charophyta/ Streptophyta. The land plants ...
containing the species ''Tetrabaena socialis''. It primarily occurs in freshwater, although it has been found in saltwater on at least one occasion. Although rare, it has a
cosmopolitan distribution In biogeography, a cosmopolitan distribution is the range of a taxon that extends across most or all of the surface of the Earth, in appropriate habitats; most cosmopolitan species are known to be highly adaptable to a range of climatic and en ...
and has even been found from
Antarctica Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean (also known as the Antarctic Ocean), it contains the geographic South Pole. ...
.


Description

''Tetrabaena'' are colonial organisms, consisting of four
cells Cell most often refers to: * Cell (biology), the functional basic unit of life * Cellphone, a phone connected to a cellular network * Clandestine cell, a penetration-resistant form of a secret or outlawed organization * Electrochemical cell, a d ...
. The four cells are arranged in a square, and attached to each other by the extensions of their cellular sheaths. Cells are ovoid, somewhat asymmetrical in shape. It a
motile Motility is the ability of an organism to move independently using metabolic energy. This biological concept encompasses movement at various levels, from whole organisms to cells and subcellular components. Motility is observed in animals, mi ...
flagellate A flagellate is a cell or organism with one or more whip-like appendages called flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryotes and the ...
, possessing two equal flagella per cell and two
contractile vacuole A contractile vacuole (CV) is a sub-cellular structure (organelle) involved in osmoregulation. It is found predominantly in protists, including unicellular algae. It was previously known as pulsatile or pulsating vacuole. Overview The contrac ...
s at the base of the flagella. Cells also contain a single cup-shaped
chloroplast A chloroplast () is a type of membrane-bound organelle, organelle known as a plastid that conducts photosynthesis mostly in plant cell, plant and algae, algal cells. Chloroplasts have a high concentration of chlorophyll pigments which captur ...
with a basal
pyrenoid Pyrenoids are sub-cellular phase-separated micro-compartments found in chloroplasts of many algae,Giordano, M., Beardall, J., & Raven, J. A. (2005). CO2 concentrating mechanisms in algae: mechanisms, environmental modulation, and evolution. ''An ...
. Although ''Tetrabaena'' possess an eyespot, it does not appear to function as a photoreceptor. ''Tetrabaena'' are capable of
sexual reproduction Sexual reproduction is a type of reproduction that involves a complex life cycle in which a gamete ( haploid reproductive cells, such as a sperm or egg cell) with a single set of chromosomes combines with another gamete to produce a zygote tha ...
, and are
isogamous Isogamy is a form of sexual reproduction that involves gametes of the same morphology (indistinguishable in shape and size), and is found in most unicellular eukaryotes. Because both gametes look alike, they generally cannot be classified as ma ...
.


Research significance

''Tetrabaena'' is a member of the order
Chlamydomonadales Chlamydomonadales, also known as Volvocales, are an order of flagellated or pseudociliated green algae, specifically of the Chlorophyceae.See the NCBIbr>webpage on Chlamydomonadales Data extracted from the Chlamydomonadales can form planar o ...
, a model lineage used in the study of the origins of
multicellular organism A multicellular organism is an organism that consists of more than one cell (biology), cell, unlike unicellular organisms. All species of animals, Embryophyte, land plants and most fungi are multicellular, as are many algae, whereas a few organism ...
s from single-celled ancestors. Within Chlamydomonadales, it is notable for being the colonial species with the smallest number of cells (4), placing it at the boundary between single- and multi-cellularity. As such, it has been characterized as the "simplest integrated multicellular organism." Genomic data and
molecular clock The molecular clock is a figurative term for a technique that uses the mutation rate of biomolecules to deduce the time in prehistory when two or more life forms diverged. The biomolecular data used for such calculations are usually nucleot ...
analysis suggest that ''Tetrabaena socialis'' may contain
cryptic species In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
.


References

Articles containing video clips Chlamydomonadales Monotypic algae genera Chlorophyta species {{Chlorophyceae-stub