Taiwanioideae
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''Taiwania'', with the single living species ''Taiwania cryptomerioides'', is a large
coniferous Conifers () are a group of conifer cone, cone-bearing Spermatophyte, seed plants, a subset of gymnosperms. Scientifically, they make up the phylum, division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a sin ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
in the cypress family
Cupressaceae Cupressaceae or the cypress family is a family of conifers. The family includes 27–30 genera (17 monotypic), which include the junipers and redwoods, with about 130–140 species in total. They are monoecious, subdioecious or (rarely) dioecio ...
.


Etymology

''Taiwania'' means 'from
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
', while ''cryptomerioides'' means 'resembling ''
Cryptomeria ''Cryptomeria'' (literally "hidden parts") is a monotypic genus of conifer in the cypress family Cupressaceae. It includes only one species, ''Cryptomeria japonica'' ( syn. ''Cupressus japonica'' L.f.). It is considered to be endemic to Japan, ...
''.Gledhill, David (2008). "The Names of Plants". Cambridge University Press. (hardback), (paperback). pp 127, 370


Taxonomy

The genus was formerly placed in the segregate family Taxodiaceae, it is now included in the
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unisp ...
subfamily Taiwanioideae of the family Cupressaceae. It is the second most basal member of the living Cupressaceae, with only '' Cunninghamia'' being more basal. Its lineage is thought to have diverged from the rest of Cupressaceae during the Middle
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
. The oldest fossil assignable to the genus is from the mid-
Cretaceous The Cretaceous ( ) is a geological period that lasted from about 143.1 to 66 mya (unit), million years ago (Mya). It is the third and final period of the Mesozoic Era (geology), Era, as well as the longest. At around 77.1 million years, it is the ...
(
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
-
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) of
Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...
. Other fossils of the genus are known from the Late Cretaceous of Asia, the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
of Asia and North America, and the
Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...
of Europe and Asia.


Range

It is native to eastern
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, growing in the
mountain A mountain is an elevated portion of the Earth's crust, generally with steep sides that show significant exposed bedrock. Although definitions vary, a mountain may differ from a plateau in having a limited summit area, and is usually higher t ...
s of central
Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...
, and locally in southwest
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
(Guizhou, Hubei, Sichuan, Yunnan,
Tibet Tibet (; ''Böd''; ), or Greater Tibet, is a region in the western part of East Asia, covering much of the Tibetan Plateau and spanning about . It is the homeland of the Tibetan people. Also resident on the plateau are other ethnic groups s ...
) and adjoining
Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
, and northern
Vietnam Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
. It is
endangered An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
by
illegal logging Illegal logging is the harvest, transportation, purchase, or sale of timber in violation of laws. The harvesting procedure itself may be illegal, including using corrupt means to gain access to forests; extraction without permission, or from a p ...
for its valuable
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
in many areas. It is very likely that the range was more extensive in the past before extensive felling for the wood. The populations in mainland Asia were treated as a distinct species ''Taiwania flousiana'' by some botanists, but the cited differences between these and the Taiwanese population are not consistent when a number of specimens from each area are compared.


Morphology

It is one of the largest tree species in Asia, reported to heights of up to tall and with a trunk up to diameter above buttressed base.Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are needle-like or awl-like and long on young trees up to about 100 years old, then gradually becoming more scale-like, long, on mature trees. The cones are small, long, with about 15–30 thin, fragile scales, each scale with two
seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...
s.


History

The genus is named after the country of Taiwan, from where it first became known to the botanical community in 1910. The wood is soft, but durable and attractively spicy scented, and was in very high demand in the past, particularly for
temple A temple (from the Latin ) is a place of worship, a building used for spiritual rituals and activities such as prayer and sacrifice. By convention, the specially built places of worship of some religions are commonly called "temples" in Engli ...
building and
coffin A coffin or casket is a funerary box used for viewing or keeping a corpse, for burial, entombment or cremation. Coffins are sometimes referred to as caskets, particularly in American English. A distinction is commonly drawn between "coffins" a ...
s. The rarity of the tree and its slow growth in
plantation Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...
s means legal supplies are now very scarce; the species has legal protection in China and Taiwan. '' Taiwania'' is also a journal that is published by
National Taiwan University National Taiwan University (NTU; zh, t=國立臺灣大學, poj=Kok-li̍p Tâi-oân Tāi-ha̍k, p=, s=) is a National university, national Public university, public research university in Taipei, Taiwan. Founded in 1928 during Taiwan under J ...
in Taiwan.


Extraordinary specimens

In 2022 a team of researchers measured a 79.1 meters (259.5 feet) ''Taiwania'' specimen in Shei-pa National Park. The tree was growing at an elevation of 2,000m. In 2023 a specimen was found measuring 84.1 meters in height.


See also

* Taiwania (supercomputer) * Taiwanica


References


External links


Conifers Around the World Special Report - ''"Taiwania Day" in Kyoto Prefectural Botanical Garden''
. {{Authority control Cupressaceae Monotypic conifer genera Trees of Myanmar Trees of China Trees of Taiwan Trees of Vietnam Vulnerable plants