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Tornidae is a
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of very small and minute sea snails with an operculum,
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military * ...
gastropod
mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is es ...
s in the clade
Littorinimorpha Littorinimorpha is a large order of snails, gastropods, consisting primarily of sea snails ( marine species), but also including some freshwater snails ( aquatic species) and land snails ( terrestrial species).Bouchet P. & Rocroi J.-P. (Ed.); Fr ...
. This family used to be known as the Vitrinellidae.Bouchet, P.; Gofas, S. (2014). Tornidae Sacco, 1896 (1884). Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=124 on 2014-12-03 Iredale has shown that the family Adeorbidae Monterosato, 1884 should be called Tornidae (Iredale, Journal of Conchology, 14, 344) Family names cited with two dates (the second one in parentheses) are those ruled by Article 40(2) of
ICZN The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the Int ...
. "If ... a family-group name was replaced before 1961 because of the synonymy of the type genus, the replacement name is to be maintained if it is in prevailing usage. A name maintained by virtue of this Article retains its own author nd date, the first date citedbut takes the priority of the replaced name he date cited in parentheses, here alluding to Adeorbidae Monterosato, 1884


General characteristics

The shells of the snails in this family are minute, colorless and glassy. The paucispiral shell is umbilicated, auriform and depressed. The entire
aperture In optics, an aperture is a hole or an opening through which light travels. More specifically, the aperture and focal length of an optical system determine the cone angle of a bundle of rays that come to a focus in the image plane. An ...
is oblique. The shell has a simple
columella Lucius Junius Moderatus Columella (; Arabic: , 4 – ) was a prominent writer on agriculture in the Roman Empire. His ' in twelve volumes has been completely preserved and forms an important source on Roman agriculture, together with the ...
, and a rounded, sharp outer lip. The corneous operculum is paucispiral, and has an eccentric nucleus. The body of the animal differs from species in the family
Trochidae The Trochidae, common name top-snails or top-shells, are a family of various sized sea snails, marine gastropod molluscs in the subclass Vetigastropoda. This family is commonly known as the top-snails because in many species the shell resembles ...
by having no cirriform appendages of the foot.
George Washington Tryon George Washington Tryon Jr. (20 May 1838 – 5 February 1888) was an American malacologist who worked at the Academy of Natural Sciences in Philadelphia. Biography George Washington Tryon was the son of Edward K. Tryon and Adeline Savidt. ...
, Manual of Conchology vol. X, p.13; 1888-1889
The
radula The radula (, ; plural radulae or radulas) is an anatomical structure used by molluscs for feeding, sometimes compared to a tongue. It is a minutely toothed, chitinous ribbon, which is typically used for scraping or cutting food before the food ...
is taenioglossate. (Thiele, 1925)


Subfamilies

The following subfamilies were recognized in the 2005 taxonomy of Bouchet & Rocroi, but have now become alternate representations: * Circulinae Fretter & Graham, 1962 * Torninae Sacco, 1896 (1884) * Teinostomatinae Cossmann, 1917 (upgraded in 2021 to family level
Teinostomatidae Teinostomatidae is a family of very small and minute sea snails with an operculum, marine gastropod mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs ...
Cossmann, 1917) * Vitrinellinae Bush, 1897


Genera

Genera in the family Tornidae include: * † '' Adeorbella'' Briart & Cornet, 1887 * '' Anticlimax'' Pilsbry & McGinty, 1946 * '' Aorotrema'' Schwengel & McGinty, 1942 * '' Canimarina'' Aguayo & Borro, 1946: synonym of ''Anticlimax'' Pilsbry & McGinty, 1946 * † '' Cantaurea'' Landau, da Silva & Heitz, 2016 * '' Caperella'' Laseron, 1958 * '' Circlotoma'' Laseron, 1958 * '' Circuitus'' Rubio & Rolán, 2017 * '' Circulter'' Laseron, 1958 * '' Cochliolepis'' Stimpson, 1958 * '' Cyclostremiscus'' Pilsbry & Olsson, 1945 * '' Discopsis'' de Folin, 1870 * ''
Discreliotia ''Pseudoliotia'' is a genus of sea snails, marine gastropod mollusks in the family Tornidae within the superfamily Truncatelloidea.Bouchet, P. (2013). Pseudoliotia Tate, 1898. In: MolluscaBase (2016). Accessed through: World Register of Marin ...
'' Laseron, 1958 * ''
Elachorbis ''Elachorbis'' is a genus of minute sea snails or micromolluscs, marine gastropod molluscs in the family Tornidae. Taxonomy The exact placement of this genus has long been contested. Tom Iredale established ''Elachorbis'' in 1914 within th ...
'' Iredale, 1914 * '' Episcynia'' Mörch, 1875 * ''
Esmeralda Esmeralda may refer to: Places * Esmeralda, Chile, a town in Chile * Esmeralda, Cuba * Esmeralda, Queensland, a locality in the Shire of Croydon, Australia * Esmeralda, Rio Grande do Sul, a municipality in Brazil * Esmeralda County, Nevada, USA ...
'' Pilsbry & Olsson, 1952 * '' Laciniorbis'' Martens, 1897 * '' Lophocochlias'' Pilsbry, 1921 * '' Lydiphnis'' Melvill, 1906 * † '' Megatyloma'' Cossmann, 1888 * '' Moeniatoma'' Laseron, 1958 * ''
Neusas ''Neusas'' is a genus of sea snails, marine gastropod mollusks in the family Tornidae Tornidae is a family of very small and minute sea snails with an operculum, marine gastropod mollusks in the clade Littorinimorpha. This family used to be k ...
'' Warén & Bouchet, 2001 * '' Ovini'' Simone, 2013 * '' Panastoma'' Pilsbry & Olsson, 1945 * ''
Parviturboides ''Parviturboides'' is a genus of gastropods belonging to the family Tornidae. The species of this genus are found in America. Species: *''Parviturboides avitus'' *''Parviturboides clausus'' *''Parviturboides copiosus'' *''Parviturboides d ...
'' Pilsbry & McGinty, 1949 * '' Peripitoma'' Laseron, 1958 * '' Pleuromalaxis'' Pilsbry & McGinty, 1945 * † '' Pterolabrella'' Maxwell, 1969 * '' Pygmaeorota'' Kuroda & Habe, 1954 * '' Scissilabra'' Bartsch, 1907 * '' Scrupus'' Finlay, 1927 * '' Sigaretornus'' Iredale, 1936 * ''
Solariorbis ''Solariorbis'' is a genus of gastropods belonging to the family Tornidae.MolluscaBase eds. (2022). MolluscaBase. Solariorbis Conrad, 1865. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aphia.php?p=taxdetai ...
'' Conrad, 1865 * '' Tholostoma'' Laseron, 1958 * '' Tornus'' Turton & Kingston, 1830 * ''
Tuberes ''Tuberes'' is a genus of sea snails, marine gastropod mollusks in the family Tornidae. MolluscaBase eds. (2022). MolluscaBase. Tuberes Rubio & Rolán, 2017. Accessed through: World Register of Marine Species at: https://www.marinespecies.org/aph ...
'' Rubio & Rolán, 2017 * ''
Uzumakiella ''Uzumakiella'' is a genus of sea snails, marine gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snai ...
'' Habe, 1958 * '' Vitridomus'' Pilsbry & Olsson, 1945 * '' Vitrinorbis'' Pilsbry & Olsson, 1952 * '' Woodringilla'' Pilsbry & Olsson, 1951 ;Genera brought into synonymy: * ''Adeorbis'' S.V. Wood, 1842: synonym of ''Tornus'' Turton & Kingston, 1830 * ''Climacia'' Dall, 1903: synonym of ''Anticlimax'' Pilsbry & McGinty, 1946 * ''Climacina'' Aguayo & Borro, 1946: synonym of ''Anticlimax'' Pilsbry & McGinty, 1946 * ''Liochrysta'' Laseron, 1958: synonym of ''Pseudoliotia'' Tate, 1898 * ''Lioprora'' Laseron, 1958: synonym of ''Canimarina'' Aguayo & Borro, 1946 * ''Miralabrum'' Pilsbry & Olsson, 1945: synonym of ''Cyclostremiscus'' Pilsbry & Olsson, 1945 * '' Ponocyclus'' Pilsbry, 1953: synonym of '' Cyclostremiscus'' Pilsbry & Olsson, 1945 * ''Pygmaerota'' Kuroda & Habe, 1952: synonym of ''Pygmaeorota'' Kuroda & Habe, 1954 * ''Soyorota'' Habe, 1961: synonym of ''Circulus'' Jeffreys, 1865 ;Alternate representations: * Subfamily Circulinae Fretter & Graham, 1962 * Subfamily Torninae Sacco, 1896 (1884) * Subfamily Vitrinellinae Bush, 1897


References

* Adam W. & Knudsen J. (1969). ''Quelques genres de mollusques prosobranches marins ou peu connus de l'Afrique occidentale''. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique 44(27): 1-69 * Backeljau, T. (1986). ''Lijst van de recente mariene mollusken van België
ist of the recent marine molluscs of Belgium Ist or IST may refer to: Information Science and Technology * Bachelor's or Master's degree in Information Science and Technology * Graduate School / Faculty of Information Science and Technology, Hokkaido University, Japan * Graduate School ...
'' Koninklijk Belgisch Instituut voor Natuurwetenschappen: Brussels, Belgium. 106 pp. *Vaught, K.C. (1989). ''A classification of the living Mollusca''. American Malacologists: Melbourne, FL (USA). . XII, 195 pp. * Rolán E. & Rubio F. 2002. ''The family Tornidae (Gastropoda, Rissooidea) in the East Atlantic.'' Resenas Malacologicas 13 (Suplemento), 98 pp. Sociedad Española de Malacología * Bouchet P. & Rocroi J.-P. (2005) ''Classification and nomenclator of gastropod families''. Malacologia 47(1-2): 1-397 * Rubio, F., Fernandez-Garcés, R. & Rolán, E., 2011. ''The family Tornidae (Gastropoda, Rissooidea) in the Caribbean and neighboring areas''. Iberus 29(2): 1-230 * * Powell A. W. B., ''New Zealand Mollusca'', William Collins Publishers Ltd, Auckland, New Zealand 1979
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{{Taxonbar, from=Q7826126 Truncatelloidea