Szechuanosaurus
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''Szechuanosaurus'' (" Szechuan lizard") is an extinct
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of carnivorous
theropod Theropoda (; from ancient Greek , (''therion'') "wild beast"; , (''pous, podos'') "foot"">wiktionary:ποδός"> (''pous, podos'') "foot" is one of the three major groups (clades) of dinosaurs, alongside Ornithischia and Sauropodom ...
dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...
from the
Late Jurassic The Late Jurassic is the third Epoch (geology), epoch of the Jurassic Period, and it spans the geologic time scale, geologic time from 161.5 ± 1.0 to 143.1 ± 0.8 million years ago (Ma), which is preserved in Upper Jurassic stratum, strata.Owen ...
. Fossils referred to the genus have been found in
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
in the Oxfordian-?
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
(Peng ''et al''., 2005). Its
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is based on several undiagnostic teeth from the Kuangyuan Series. Additional possible specimens of ''Szechuanosaurus'' were also reported from the Kalaza Formation, also located in China. ''Szechuanosaurus'' is at times regarded as a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
'' due to the lack of diagnostic features in the fossilized teeth upon which the genus is based. Although the fossils are too fragmentary for confident identification, ''Szechuanosaurus'' is often interpreted as a medium-sized
allosaurid Allosauridae is an extinct family of medium to large bipedal, carnivorous allosauroid theropod dinosaurs from the Late Jurassic. Allosauridae is a fairly old taxonomic group, having been first named by the American paleontologist Othniel Charles ...
or perhaps a
metriacanthosaurid Metriacanthosauridae (Greek for "moderately-spined lizards") is an extinct family of allosauroid theropod dinosaurs that lived in Europe and Asia from the Middle Jurassic to the Early Cretaceous. The family is split into two subgroups: Metriacant ...
, capable of reaching lengths of around 7.3 meters.


Discovery and species

Three species have been assigned to this genus over the years. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
is ''Szechuanosaurus campi'', named by
Yang Zhongjian Yang Zhongjian, also Yang Chung-chien (; 1 June 1897 – 15 January 1979), courtesy name Keqiang (), also known as C.C. (Chung Chien) Young, was a Chinese paleontologist and zoologist. He was one of China's foremost vertebrate paleontologists. ...
("Chung Chien Young") in 1942 for four isolated tooth specimens: IVPP V235, two partial teeth; IVPP V236, a partial tooth; IVPP V238, several tooth fragments; and IVPP V239, a single tooth. The teeth form a
syntype In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part o ...
series and were not found together. Also a very fragmentary skeleton, specimen UCMP 32102, was referred to the genus by Yang.Young, C.C., 1942, "Fossil vertebrates from Kuangyuan, N. Szechuan, China", ''Bulletin of the Geological Society of China'', 22: 293-309 Some of the teeth indicate a large body-size. These fossils, although possibly
metriacanthosaurid Metriacanthosauridae (Greek for "moderately-spined lizards") is an extinct family of allosauroid theropod dinosaurs that lived in Europe and Asia from the Middle Jurassic to the Early Cretaceous. The family is split into two subgroups: Metriacant ...
, are now considered to be non-diagnostic, making ''S. campi'' a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''.Brusatte, S. L., Benson, R. B. J., and Xu, X. 2010. "The evolution of large-bodied theropod dinosaurs during the Mesozoic in Asia". ''
Journal of Iberian Geology ''Journal of Iberian Geology'' (formerly ''Cuadernos de Geología Ibérica'') is a triannual peer-reviewed scientific journal published by the Universidad Complutense de Madrid. The journal covers the field of geology and related earth science ...
'', 36: 275-296
The generic name refers to Szechuan. The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
honours the American paleontologist
Charles Lewis Camp Charles Lewis Camp (March 12, 1893 – August 14, 1975) was an American Palaeontology, palaeontologist and Zoology, zoologist, working from the University of California, Berkeley. He took part in excavations at the 'Placerias Quarry', in 1930 and ...
who had discovered UCMP 32102. A partial skeleton, CV 00214, was initially listed by Dong ''et al.'' (1978) in a faunal list as a new species of ''Szechuanosaurus'', ''Szechuanosaurus yandonensis''. At first, there was no description or illustration of it, making ''S. yandonensis'' at the time a ''
nomen nudum In Taxonomy (biology), taxonomy, a ''nomen nudum'' ('naked name'; plural ''nomina nuda'') is a designation which looks exactly like a scientific name of an organism, and may have originally been intended to be one, but it has not been published ...
''. Later,
Dong Zhiming Dong Zhiming (Chinese language, Chinese: 董枝明, Pinyin: ''Dǒng Zhimíng''; January 1937 – 20 October 2024) was a Chinese vertebrate paleontologist formerly employed at the Institute of Vertebrate Paleontology and Paleoanthropology (IVPP) ...
''et alii'' (1983) described it, and assigned it to ''Szechuanosaurus campi'' The affinities of this skeleton are uncertain, and it has only been briefly described. Holtz ''et al''. (2004) included it in their
phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
and found it to be the most basal
tetanuran Tetanurae (/ˌtɛtəˈnjuːriː/ or "stiff tails") is a clade that includes most theropod dinosaurs, including megalosauroids, allosauroids, and coelurosaurs (which includes tyrannosauroids, ornithomimosaurs, compsognathids and maniraptoran ...
. This individual was a medium-sized theropod, with an
ischium The ischium (; : is ...
(a pelvic bone) of ; for comparison, an ischium of '' Piatnitzkysaurus'' estimated to weigh is long. In 2000,
Daniel Chure Daniel commonly refers to: * Daniel (given name), a masculine given name and a surname * List of people named Daniel * List of people with surname Daniel * Daniel (biblical figure) * Book of Daniel, a biblical apocalypse, "an account of the activi ...
referred the specimen to " Szechuanoraptor dongi", itself an invalid ''nomen ex dissertatione''.Chure, D.J. (2000) ''A new species of'' Allosaurus ''from the Morrison Formation of Dinosaur National Monument (UT–CO) and a revision of the theropod family Allosauridae''. Unpublished Ph.D. dissertation, Columbia University, New York Carrano, Benson & Sampson (2012) synonymized it with ''
Yangchuanosaurus ''Yangchuanosaurus'' is an extinct genus of metriacanthosaurid theropod dinosaur that lived in China from the Middle Jurassic to Late Jurassic periods (Bathonian to Tithonian stages), and was similar (although slightly larger) in size and appea ...
shangyouensis'' from the same formation. The third species is ''Szechuanosaurus zigongensis'', named by Gao Yuhui in 1993 for an almost complete skeleton, specimen ZDM 9011.Gao, Y., 1993, "A new species of ''Szechuanosaurus'' from the Middle Jurassic of Dashanpu, Zigong, Sichuan", ''Vertebrata PalAsiatica'' 31(4): 308-314 It is an older species, from the
Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
, appearing to be distinct from the type species and therefore requires its own genus name. It was reassigned to ''
Yangchuanosaurus ''Yangchuanosaurus'' is an extinct genus of metriacanthosaurid theropod dinosaur that lived in China from the Middle Jurassic to Late Jurassic periods (Bathonian to Tithonian stages), and was similar (although slightly larger) in size and appea ...
'', as a ''Yangchuanosaurus zigongensis'', by Matthew Carrano, Roger Benson & Scott Sampson in 2012.


Notes


References


''Szechuanosaurus campi'' in The Theropod Database
{{Taxonbar, from=Q637535 Tetanurae Dinosaur genera Oxfordian dinosaurs Taxa named by Yang Zhongjian Fossil taxa described in 1942 Dinosaurs of China