Symmoriiformes is an extinct order of

cartilaginous fish Chondrichthyes (; ) is a class of jawed fish that contains the cartilaginous fish or chondrichthyans, which all have skeletons primarily composed of cartilage. They can be contrasted with the Osteichthyes or ''bony fish'', which have skeleto ...

. Originally named Symmoriida by Zangerl (1981), the name has since been corrected to Symmoriiformes to avoid confusion with a family. The symmoriiform fossils record begins during the late

Devonian The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian per ...

, and most had become extinct by the start of the

Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the s ...

, with the genus '' Dwykaselachus'' from the

Artinskian In the geologic timescale, the Artinskian is an age (geology), age or stage (stratigraphy), stage of the Permian. It is a subdivision of the Cisuralian Epoch (geology), Epoch or series (stratigraphy), Series. The Artinskian likely lasted between ...

Kungurian In the geologic timescale, the Kungurian is an age or stage of the Permian. It is the latest or upper of four subdivisions of the Cisuralian Epoch or Series. The Kungurian lasted between and million years ago (Ma). It was preceded by the Arti ...

of South Africa being the latest known uncontroversial occurrence. Teeth described from the

Valanginian In the geologic timescale, the Valanginian is an age or stage of the Early or Lower Cretaceous. It spans between 137.05 ± 0.2 Ma and 132.6 ± 0.2 Ma (million years ago). The Valanginian Stage succeeds the Berriasian Stage of the Lower Cretac ...

of France and Austria indicate that members of the family Falcatidae might have survived until the

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

; however, it has since been proposed that these teeth more likely belonged to neoselachian sharks.

Fossil distribution

Fossil evidence of Symmoriida have been found at Bear Gulch,

Fergus County, Montana Fergus County is a County (United States), county in the U.S. state of Montana. As of the 2020 United States census, 2020 census, the population was 11,446. Its county seat is Lewistown, Montana, Lewistown. The county was founded in 1885 and na ...

, Bethel Quarry, Pike County, Indiana, Kinshozan quarry,

Alaska Alaska ( ) is a non-contiguous U.S. state on the northwest extremity of North America. Part of the Western United States region, it is one of the two non-contiguous U.S. states, alongside Hawaii. Alaska is also considered to be the north ...

Gifu Prefecture is a Prefectures of Japan, prefecture of Japan located in the Chūbu region of Honshu. Gifu Prefecture has a population of 1,910,511 () and has a geographic area of . Gifu Prefecture borders Toyama Prefecture to the north; Ishikawa Prefecture ...

Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...

Bashkortostan Bashkortostan, officially the Republic of Bashkortostan, sometimes also called Bashkiria, is a republic of Russia between the Volga river and the Ural Mountains in Eastern Europe. The republic borders Perm Krai to the north, Sverdlovsk Oblast ...

Russian Federation Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders ...

and possibly also

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

Classification

Symmoriiformes were previously regarded as members of the Elasmobranchii, making them distant relatives of living sharks and rays. Zangerl (1981), who erected the order, considered them to be elasmobranchs, and this classification has been followed by some subsequent authors. The uncrushed braincase of '' Dwykaselachus'' indicates that symmoriiforms may be members of Holocephali, as much of the internal anatomy, including the otic labyrinth and brain space configuration are similar to those of living

chimaeras Chimaeras are Chondrichthyes, cartilaginous fish in the order (biology), order Chimaeriformes (), known informally as ghost sharks, rat fish (not to be confused with rattails), spookfish, or rabbit fish; the last two names are also applied to B ...

. Symmoriiformes have alternatively been considered early diverging

stem-group In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. ...

chondrichthyans.

Gallery

References

*http://palaeo.gly.bris.ac.uk/benton/vertclass.html {{Taxonbar, from=Q139057 Prehistoric cartilaginous fish orders