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''Suaeda fruticosa'' is a species of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of ...
in the
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
Amaranthaceae Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making i ...
(formerly placed in the family Chenopodiaceae). It is a small shrub, with very variable appearance over its wide range. It is a halophyte, and occurs in arid and semi-arid saltflats,
salt marsh A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is domin ...
es and similar
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s.


Description

''Suaeda fruticosa'' is a low shrub growing to a height of about . It is extremely variable throughout its wide range in height, growth habit, colouring, internode length, leaf shape, and the size and orientation of inflorescences and fruits. It is usually a rounded, much-branched bush but can be prostrate, climbing or straggling. It is densely-branched, the stems feeling very rough when the leaves are shed, pale green at first, becoming grey and fissured. The leaves are succulent, the smaller ones being long and narrow while the larger ones are elliptical. The flowers grow in clusters in the leaf axils. Some are bisexual, being drum-shaped and up to wide, with five succulent
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s fused to a third of their length. Others are entirely female, rather smaller with non-succulent tepals, fused for half their length, persistent and partly concealing the fruit. There are three stigmas. The perianth enlarges in the bisexual fruits but remains unchanged in the female fruits. Reproduction is mainly by seed, which are black and shining, slightly flattened, globular or drop-shaped. It has a chromosome number of 2n=36.


Taxonomy

The species has a complicated taxonomic history, and other species have been called ''S. fruticosa'' for a very long time. Two later homonyms of ''Suaeda fruticosa'' Forssk. ex J.F.Gmel. exist: *''Suaeda fruticosa'' (L.) Delile is a synonym of '' Suaeda vera'' Forssk. ex J.F.Gmel.. *''Suaeda fruticosa'' Hook. & Arn. is a synonym of ''
Suaeda nigra ''Suaeda nigra'', often still known by the former name ''Suaeda moquinii'', is a species of flowering plant in the amaranth family, known by the vernacular names bush seepweed or Mojave sea-blite. Taxonomy ''Suaeda nigra'' was first formally d ...
'' (Raf.) J.F.Macbr. In Europe, what was formerly called ''S. fruticosa'' is now known to be ''S. vera'', whereas in Africa what was formerly identified as ''S. fruticosa'' is now thought to be either ''S. vera'' or ''S. vermiculata''. The 'real' ''S. fruticosa'' Forssk. ex J.F.Gmel. occurs from the Arabian Peninsula and the Middle East eastwards to the Indian subcontinent, however, here it has in some cases been misidentified as ''S. vermiculata''. In Israel all three species may occur (or not), elsewhere ''S. fruticosa'' does not appear to occur in the same countries as the other two species. The name ''S. fruticosa'' has furthermore also been misapplied in North America to plants of the species ''S. nigra''.


Distribution

It occurs in the Arabian peninsula, Iran, Afghanistan and the Indian sub-continent.


Ecology

It is a common and widespread species growing on sometimes-flooded alluvial land, drier areas, coastal regions, salt flats and salt marshes on soils that are sandy and soils that have a lot of
clay Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4). Clays develop plasticity when wet, due to a molecular film of water surrounding the clay part ...
. This plant is common in the saltlands of the Indian sub-continent, and is one of the dominant plants in the ''Tamarix/Salvadora/Suaeda'' climax vegetation. Other associated plants include '' Zygophyllum simplex'', '' Cressa cretica'', ''
Caroxylon imbricatum ''Caroxylon imbricatum'', Synonym (taxonomy), synonym ''Salsola imbricata'', is a small species of shrub in the Family (biology), family Amaranthaceae. It grows in deserts and arid regions of north Africa, the Arabian Peninsula and southwestern As ...
'', '' Salsola stocksii'', '' Aeluropus lagopoides'' and ''
Sporobolus helvolus ''Sporobolus'' is a nearly cosmopolitan genus of plants in the grass family. The name ''Sporobolus'' means "seed-thrower", and is derived from Ancient Greek word (), meaning "seed", and the root of () "to throw", referring to the dispersion of ...
''. The few trees and shrubs growing in these saline habitats include '' Salvadora persica'', ''
Salvadora oleoides ''Salvadora oleoides'' is a small bushy evergreen tree found in India and Pakistan and southern Iran. The root and stem possess various antimicrobial agents and is traditionally used as toothbrush in Pakistan and India Habitat The vann is commo ...
'', ''
Tamarix dioica ''Tamarix dioica'' is a twiggy shrub or small tree that grows in saline habitats in western Asia. Common names include ghaz and khagal in Pakistan, lal jhau, urusia, ban jhau, nona-gach, urichiya in Bangladesh and nona jhau in the Sunderbans. D ...
'' and '' Capparis decidua''.


Uses

It is one of a number of plants high in sodium known as
barilla ''Barilla'' refers to several species of salt-tolerant ( halophyte) plants that, until the 19th century, were the primary source of soda ash and hence of sodium carbonate. The word "barilla" was also used directly to refer to the soda ash obtain ...
which were used to make
soda ash Sodium carbonate, , (also known as washing soda, soda ash and soda crystals) is the inorganic compound with the formula Na2CO3 and its various hydrates. All forms are white, odourless, water-soluble salts that yield moderately alkaline solutions ...
for use in the soap and glass industries. Large quantities were exported from India in the 18th and 19th centuries, and ''S. fruticosa'' and various chenopods, are still collected from the seasonal salt marshes in the
Rann of Kutch The Rann of Kutch (alternately spelled as Kuchchh) is a large area of salt marshes that span the border between India and Pakistan. It is located in Gujarat (primarily the Kutch district), India, and in Sindh, Pakistan. It is divided into t ...
for local use in the manufacture of soap and
baking soda Sodium bicarbonate ( IUPAC name: sodium hydrogencarbonate), commonly known as baking soda or bicarbonate of soda, is a chemical compound with the formula NaHCO3. It is a salt composed of a sodium cation ( Na+) and a bicarbonate anion ( HCO3� ...
. It also provides forage for
camel A camel (from: la, camelus and grc-gre, κάμηλος (''kamēlos'') from Hebrew or Phoenician: גָמָל ''gāmāl''.) is an even-toed ungulate in the genus ''Camelus'' that bears distinctive fatty deposits known as "humps" on its back. ...
s. The seeds could be a potential source of edible oils which are rich in unsaturated fatty acids. The plant is an obligate halophyte and can be used to reduce the salinity of soils.


References

{{Taxonbar, from=Q6430252 fruticosa Flora of Iran Flora of the Indian subcontinent