Pentatomidae is a family of

insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (head, thorax and abdomen), three pairs ...

s belonging to the order Hemiptera, generally called shield bugs or stink bugs. Pentatomidae is the largest family in the superfamily

Pentatomoidea The Pentatomoidea are a superfamily of insects in the Heteroptera suborder of the Hemiptera order. As Hemiptera, they share a common arrangement of sucking mouthparts. The roughly 7000 species under Pentatomoidea are divided into 21 families ...

, and contains around 900 genera and over 4700 species.Robert G. Foottit, Peter H. Adler ''Insect Biodiversity: Science and Society'', John Wiley and Sons, 2009, As hemipterans, the pentatomids have piercing sucking mouthparts, and most are

phytophagous A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpar ...

, including several species which are severe pests on

agricultural Agriculture or farming is the practice of cultivating plants and livestock. Agriculture was the key development in the rise of sedentary human civilization, whereby farming of domesticated species created food surpluses that enabled peopl ...

crop A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydropo ...

s. However, some species, particularly in the subfamily

Asopinae Asopinae are a subfamily of stink bugs (family Pentatomidae). They are predatory stink bugs that are useful as biological control agents against pests, even against other Pentatomid species, which are all herbivorous. Genera # '' Afrius'' ...

, are predatory and may be considered beneficial.

Etymology

The name "Pentatomidae" is from the

Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...

''pente'' meaning "five" and ''tomos'' meaning "section", and refers to the five segments of their antennae. Pentatomids are generally called "shield bugs" in British

English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national id ...

, or "stink bugs" in

American English American English, sometimes called United States English or U.S. English, is the set of varieties of the English language native to the United States. English is the most widely spoken language in the United States and in most circumstances ...

. However, the term shield bugs is also applied broadly to include several related families (e.g.

Acanthosomatidae Acanthosomatidae is a family of Hemiptera, commonly named "shield bugs," for which Kumar in his 1979 world revision recognized 47 genera; now this number is 55 genera, with about 200 species, and is one of the least diversified families within Pe ...

Scutelleridae Scutelleridae is a family of true bugs. They are commonly known as jewel bugs or metallic shield bugs due to their often brilliant coloration. They are also known as shield-backed bugs due to the enlargement of the thoracic scutellum into a con ...

, and Cydnidae), or specifically only to refer to species in the family Acanthosomatidae. The term shield bug refers to the generalized body shape of adult bugs in these families which resembles a

heraldic shield In heraldry, an escutcheon () is a shield that forms the main or focal element in an achievement of arms. The word can be used in two related senses. In the first sense, an escutcheon is the shield upon which a coat of arms is displayed. In the ...

when viewed from above. The American name "stink bug" is specific to the Pentatomidae, and refers to their ability to release a pungent defensive spray when threatened, disturbed, or crushed. The composition of this spray may vary between species, and even by sex or age, but generally includes

aldehyde In organic chemistry, an aldehyde () is an organic compound containing a functional group with the structure . The functional group itself (without the "R" side chain) can be referred to as an aldehyde but can also be classified as a formyl group ...

s and

alkane In organic chemistry Organic chemistry is a subdiscipline within chemistry involving the science, scientific study of the structure, properties, and reactions of organic compounds and organic materials, i.e., matter in its various forms tha ...

s. Descriptions of the smells vary widely, and include oily, dusty, woody and earthy, and like

coriander Coriander (;

. In some species, the liquid contains cyanide compounds and a rancid almond scent, used to protect themselves and discourage predators. The term "stink bug" may also be a vernacular for unrelated insects such as pinacate beetles (in the genus ''

Eleodes ''Eleodes'' is a genus of darkling beetles, family Tenebrionidae. They are commonly known as pinacate beetles or desert stink beetles. They are endemic to western North America, with many species found in the Sonoran Desert. The name ''pinacate' ...

'').

Taxonomy

There are several subfamilies, of which the Aphylinae is often given family status, but is here retained as a subfamily, following Grazia ''et al.'' (2008). The subfamilies include: * Aphylinae Bergroth, 1906 – Australia # '' Aphylus'' Bergroth, 1906 # '' Neoaphylum'' Štys & Davidová-Vilímová, 2001 *

Spinola, 1850 ** ''

Troilus Troilus ( or ; grc, Τρωΐλος, Troïlos; la, Troilus) is a legendary character associated with the story of the Trojan War. The first surviving reference to him is in Homer's '' Iliad,'' composed in the late 8th century BCE. In Greek ...

'' Stål, 1868 *Cyrtocorinae Distant, 1880 # '' Ceratozygum'' Horváth, 1916 # '' Cyphothyrea'' Horváth, 1916 # '' Cyrtocoris'' White, 1842 # '' Pseudocyrtocoris'' Jensen-Haarup, 1926 *

Discocephalinae The Discocephalinae are a subfamily of shield bugs, erected by Fieber in 1860,Fieber FX (1860) ''Die europäischen Hemiptera. Halbflüger. (Rhynchota Heteroptera).'' 444 pp. Carl Gerold's Sohn (Wien). and found worldwide. Tribes and genera ''B ...

Fieber, 1860 ** '' Discocephala'' Laporte, 1832 ** '' Ochlerus'' Spinola, 1837 *Edessinae Fieber, 1860 # '' Anisoedessa'' Nunes & Fernandes, 2019 # '' Brachystethus'' Laporte, 1833 # '' Doesburgedessa'' Fernandes, 2010 # ''

Edessa (bug) ''Edessa'' is a large New World genus of stink bugs in the family Pentatomidae, containing over 250 described species. Several other genera used to be included until recently, such as ''Pygoda'', ''Ascra (genus), Ascra'', ''Paraedessa'', ''Do ...

'' Fabricius, 1803 # '' Grammedessa'' Correia & Fernandes, 2016 # '' Lopadusa'' Stål, 1860 # '' Mediocampus'' Thomas, 1994 # '' Neopharnus'' Van Duzee, 1910 # ''

Olbia (bug) Olbia (, ; sc, Terranoa; sdn, Tarranoa) is a city and commune of 60,346 inhabitants (May 2018) in the Italian insular province of Sassari in northeastern Sardinia, Italy, in the historical region of Gallura. Called in the Roman age, Civita ...

'' Stål, 1862 # '' Pantochlora'' Stål, 1870 # '' Paraedessa'' Silva & Fernandes, 2013 # '' Peromatus'' Amyot & Serville, 1843 # '' Pharnus'' Stål, 1867 # '' Plagaedessa'' Almeida & Fernandes, 2018 # '' Platistocoris'' Rider, 1998 # '' Praepharnus'' Barber & Bruner, 1932 * Pentatominae Leach, 1815 ** ''

Bathycoelia ''Bathycoelia''Amyot CJ-B, Audinet-Serville JG (1843) ''Histoire naturelle des insectes. Hémiptères. Librairie encyclopédique de Roret'', Paris. XXIII, 110. is a genus of shield bugs in the subfamily Pentatominae and the monotypic tribe Bathyc ...

'' Amyot & Serville, 1843 ** '' Pentatoma'' Olivier, 1789 *

Phyllocephalinae The Phyllocephalinae are a subfamily of shield bugs erected by Amyot and Serville in 1843. Tribes and genera ''BioLib'' lists the following genera in four tribes:Cressona'' Dallas, 1851 ** '' Megarrhamphus'' Bergroth, 1891 ** '' Phyllocephala'' Laporte, 1833 ** '' Tetroda'' Amyot & Serville, 1843 * Podopinae Amyot & Serville, 1843 ** '' Graphosoma'' Laporte de Castelnau, 1833 ** '' Podops'' Laporte de Castelnau, 1833 *Serbaninae (monotypic) # '' Serbana borneensis'' Distant, 1906 *Stirotarsinae (monotypic) # '' Stirotarsus'' Bergroth, 1911 *''

incertae sedis ' () or ''problematica'' is a term used for a taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertain ...

'' # '' Antillosciocoris'' Thomas, 2005 # ''

Asopus Asopus (; grc, Ἀ̄σωπός ''Āsōpos'') is the name of four different rivers in Greece and one in Turkey. In Greek mythology, it was also the name of the gods of those rivers. Zeus carried off Aegina, Asopus' daughter, and Sisyphus, who h ...

'' Burmeister, 1834 # '' Jostenicoris'' Arnold, 2011

Description

All pentatomids have 5-segmented antennae, and 3 tarsal segments on each foot. They generally have a large triangular scutellum in the center of the back. The body shape of adult pentatomids is generally " shieldlike," when viewed from above, but this varies between species, and is not true for the immature nymphal stages. The forewings of stink bugs are called hemelytra, with the basal half thickened while the apex is membranous. At rest, the wings are laid across the back of the insect, with the membranous wingtips overlapping. The hindwings are entirely membranous.

Economics

Several stink bugs and shield bugs are considered agricultural

pest Pest or The Pest may refer to: Science and medicine * Pest (organism), an animal or plant deemed to be detrimental to humans or human concerns ** Weed, a plant considered undesirable * Infectious disease, an illness resulting from an infection ** ...

s, because they can grow into large populations that feed on crops, damaging production, and they are resistant to many pesticides. They are a threat to cotton, corn, sorghum, soybeans, native and ornamental trees, shrubs, vines, weeds, and many cultivated crops. In

Mexico Mexico ( Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guate ...

, some species of stink bugs are called ''jumil'', ''chinche de monte'', ''xotlinilli'', or ''chumil'' (e.g. ''

Edessa mexicana Edessa (; grc, Ἔδεσσα, Édessa) was an ancient city ('' polis'') in Upper Mesopotamia, founded during the Hellenistic period by King Seleucus I Nicator (), founder of the Seleucid Empire. It later became capital of the Kingdom of Os ...

''). They are most often eaten in the states of

Morelos Morelos (), officially the Free and Sovereign State of Morelos ( es, Estado Libre y Soberano de Morelos), is one of the 32 states which comprise the Federal Entities of Mexico. It is divided into 36 municipalities and its capital city is Cu ...

and

Guerrero Guerrero is one of the 32 states that comprise the Administrative divisions of Mexico, 32 Federal Entities of Mexico. It is divided in Municipalities of Guerrero, 81 municipalities and its capital city is Chilpancingo and its largest city is Acap ...

. The flavor is sometimes said to resemble cinnamon, or sometimes a bitter medicinal flavor. ''Jumiles'' may be used for making sauces or as

taco A taco (, , ) is a traditional Mexican food consisting of a small hand-sized corn- or wheat-based tortilla topped with a filling. The tortilla is then folded around the filling and eaten by hand. A taco can be made with a variety of filli ...

filling. Since recent arrival in the U.S., populations of the

brown marmorated stink bug The brown marmorated stink bug (''Halyomorpha halys'') is an insect in the family Pentatomidae, native to China, Japan, Korea and other Asian regions. In September 1998 it was collected in Allentown, Pennsylvania, where it is believed to have b ...

have grown significantly. As of October 2014, brown marmorated stink bugs can be found in 41 out of 50 states within the U.S. In 2016 New Zealand's MPI put out an alert to prevent this invasive species from entering via imported cargo. Acoloba lanceolata01

European species

European species within this family include: * '' Acrosternum arabicum'' Wagner, 1959 * '' Acrosternum heegeri'' Fieber, 1861 * '' Acrosternum malickyi'' Josifov & Heiss, 1989 * '' Acrosternum millierei'' (Mulsant & Rey, 1866) * '' Acrosternum rubescens'' (Noualhier, 1893) * '' Aelia acuminata'' (Linnaeus, 1758) * '' Aelia albovittata'' Fieber, 1868 * '' Aelia angusta'' Stehlik, 1976 * '' Aelia cognata'' Fieber, 1868 * '' Aelia cribrosa'' Fieber, 1868 * '' Aelia furcula'' Fieber, 1868 * ''

Aelia germari Aelia may refer to: * ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 493 ...

'' Kuster, 1852 * ''

Aelia klugii Aelia may refer to: * ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 493 ...

'' Hahn, 1833 * ''

Aelia notata Aelia may refer to: * ''Aelia'' (bug), an insect genus in the tribe Aelini of the sub family Pentatominae Pentatominae is a subfamily of Pentatomidae, a family of shield bugs. This subfamily is the largest one within the Pentatomidae, having 493 ...

'' Rey, 1887 * '' Aelia rostrata'' Boheman, 1852 * '' Aelia sibirica'' Reuter, 1884 * '' Aelia virgata'' (Herrich-Schäffer, 1841) * '' Ancyrosoma leucogrammes'' (Gmelin, 1790) * ''

Andrallus spinidens ''Andrallus'' is a genus of predatory stink bugs Stink bug or stinkbug is a common name for several insects and may refer to: * Any of several bugs in the true bug (hemipteran) family Pentatomidae **''Halyomorpha halys'', or brown marmorated sti ...

'' (Fabricius, 1787) * ''

Antheminia absinthii ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Wagner, 1952) * '' Antheminia aliena'' (Reuter, 1891) * ''

Antheminia lunulata ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Goeze, 1778) * ''

Antheminia pusio ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Kolenati, 1846) * ''

Antheminia varicornis ''Antheminia'' is a genus of true bugs belonging to the family Pentatomidae. The genus was first described by Mulsant and Rey in 1866. The species of this genus are found in Europe, Northern America, Japan. Species These species belong to the ...

'' (Jakovlev, 1874) * '' Apodiphus amygdali'' (Germar, 1817) * '' Arma custos'' (Fabricius, 1794) * '' Arma insperata'' Horvath, 1899 * '' Asaroticus solskyi'' Jakovlev, 1873 * '' Bagrada abeillei'' Puton, 1881 * ''

Bagrada confusa ''Bagrada'' is a genus of stink bugs in the family Pentatomidae. Species * ''Bagrada abeillei'' Puton, 1881 * ''Bagrada elegans'' Puton, 1873 * ''Bagrada funerea'' Horvath, 1901 * ''Bagrada hilaris'' (Burmeister, 1835) (Bagrada bug) * ''Bagrada ...

'' Horvath, 1936 * '' Bagrada elegans'' Puton, 1873 * '' Bagrada funerea'' Horvath, 1901 * '' Bagrada hilaris'' (Burmeister, 1835) * '' Bagrada stolida'' (Herrich-Schäffer, 1839) * '' Bagrada turcica'' Horvath, 1936 * ''

Brachynema cinctum ''Brachynema'' is the scientific name of two genera of organisms and may refer to: * ''Brachynema'' (bug), a genus of stinkbugs in the family Pentatomidae * ''Brachynema'' (plant), a genus of plants in the family Olacaceae {{Genus disambigu ...

'' (Fabricius, 1775) * ''

Brachynema germarii ''Brachynema'' is the scientific name of two genera of organisms and may refer to: * ''Brachynema'' (bug), a genus of stinkbugs in the family Pentatomidae * ''Brachynema'' (plant), a genus of plants in the family Olacaceae {{Genus disambigu ...

'' (Kolenati, 1846) * ''

Brachynema purpureomarginatum ''Brachynema'' is the scientific name of two genera of organisms and may refer to: * ''Brachynema'' (bug), a genus of stinkbugs in the family Pentatomidae * ''Brachynema'' (plant), a genus of plants in the family Olacaceae {{Genus disambigu ...

'' (Rambur, 1839) * '' Capnoda batesoni'' Jakovlev, 1889 * ''

Carpocoris coreanus ''Carpocoris'' is a genus of shield bug in the family Pentatomidae. The species of this genus are quite similar.Carpocoris fuscispinus ''Carpocoris fuscispinus'' is a species of shield bug in the family Pentatomidae. Subspecies * ''Carpocoris fuscispinus hahni'' (Flor, 1856) * ''Carpocoris fuscispinus incerta'' (Tamanini, 1959) * ''Carpocoris fuscispinus maculosa'' (Tamanini, ...

'' (Boheman, 1850) * ''

Carpocoris melanocerus ''Carpocoris'' is a genus of shield bug in the family Pentatomidae. The species of this genus are quite similar.Carpocoris pudicus'' (Poda, 1761) * '' Carpocoris purpureipennis'' (De Geer, 1773) * ''

Chlorochroa juniperina ''Chlorochroa juniperina'' is a European species of shield bug in the tribe Nezarini Nezarini is a tribe of stink bugs in the family Pentatomidae. Genera There are at least 21 described genera in the Nezarini including: * ''Acrosternum ...

'' (Linnaeus, 1758) * '' Chlorochroa pinicola'' (Mulsant & Rey, 1852) * ''

Chlorochroa reuteriana ''Chlorochroa''Stål C (1872) ''K. svenska VetenskAkad. Handl.'' 10, no. 4. is a genus of shield (stink) bugs in the family Pentatomidae, found in Europe and North America. There are over 20 described species in ''Chlorochroa''. Description A ...

'' (Kirkaldy, 1909) * '' Chroantha ornatula'' (Herrich-Schäffer, 1842) * ''

Codophila varia ''Codophila varia'' is a species of ''shield bug'' belonging to the subfamily Pentatominae in the family Pentatomidae. Distribution This species is present around most of Europe (Albania, Bosnia and Herzegovina, Bulgaria, Croatia, Cyprus, Franc ...

'' (Fabricius, 1787) * '' Crypsinus angustatus'' (Baerensprung, 1859) * '' Derula flavoguttata'' Mulsant & Rey, 1856 * ''

Dolycoris baccarum ''Dolycoris baccarum'', the sloe bug or hairy shieldbug, is a species of shield bug in the family Pentatomidae. Distribution and habitat This species is widespread in most of Europe and Central Asia. These shield bugs mainly inhabit hedgerows ...

'' (Linnaeus, 1758) * '' Dolycoris numidicus'' Horvath, 1908 * '' Dryadocoris apicalis'' (Herrich-Schäffer, 1842) * ''

Dybowskyia reticulata ''Dybowskyia'' is a genus of true bugs belonging to the family Pentatomidae and tribe Tarisini. The species of this genus are found in Europe and Japan. Species: * '' Dybowskyia reticulata'' (Dallas, 1851) References {{Taxonbar, from=Q40 ...

'' (Dallas, 1851) * '' Dyroderes umbraculatus'' (Fabricius, 1775) * '' Eudolycoris alluaudi'' (Noualhier, 1893) * ''

Eurydema cyanea ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * '' Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' ...

'' (Fieber, 1864) * ''

Eurydema dominulus ''Eurydema dominulus'', also known by its common name painted bug, is a cabbage bug from the genus ''Eurydema''. The species was first described in 1911. Description ''Eurydema dominulus'' is a cabbage bug that is vividly orange and black. ...

'' (Scopoli, 1763) * ''

Eurydema eckerleini ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * '' Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * ''Eurydema herbacea'' ( ...

'' Josifov, 1961 * ''

Eurydema fieberi ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * '' Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' ...

'' Schummel, 1837 * '' Eurydema gebleri'' Kolenati, 1846 * ''

Eurydema herbacea ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * '' Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * '' Eurydema herbacea'' ...

'' (Herrich-Schäffer, 1833) * ''

Eurydema lundbaldi ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * ''Eurydema herbacea'' (Her ...

'' Lindberg, 1960 * '' Eurydema maracandica'' Oshanin, 1871 * '' Eurydema nana'' Fuente, 1971 * ''

Eurydema oleracea ''Eurydema oleracea'' is a species of shield bug in the family Pentatomidae and is commonly known as the rape bug, the crucifer shield bug, the cabbage bug or the brassica bug. Its specific name (zoology), specific name ''oleracea'' means "relat ...

'' (Linnaeus, 1758) * ''

Eurydema ornata ''Eurydema ornata'' is a species of shield bug in the family Pentatomidae. Description ''Eurydema ornata'' can reach a length of . The body has gray or bright red colors, with black markings. Sometimes it shows the red patches coloured white an ...

'' (Linnaeus, 1758) * ''

Eurydema rotundicollis ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * '' Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * ''Eurydema herbacea'' ( ...

'' (Dohrn, 1860) * '' Eurydema rugulosa'' (Dohrn, 1860) * ''

Eurydema sea ''Eurydema'' is a genus of shield bug in the family Pentatomidae. Species (following ) Subgenus ''Eurydema'' Laporte de Castelnau, 1833 * ''Eurydema eckerleini'' Josifov, 1961 * ''Eurydema gebleri'' Kolenati, 1846 * ''Eurydema herbacea'' (H ...

'' Pericart & De la Rosa 2004 * '' Eurydema spectabilis'' Horvath, 1882 * '' Eurydema ventralis'' Kolenati, 1846 * ''

Eysarcoris aeneus ''Eysarcoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. Species * ''Eysarcoris aeneus'' (Scopoli, 1763) * ''Eysarcoris confusus'' Fuente, 1971 * ''Eysarcoris distinctus'' (Schouteden) * ''Eysarcoris ...

'' (Scopoli, 1763) * ''

Eysarcoris ventralis ''Eysarcoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. Species * ''Eysarcoris aeneus'' (Scopoli, 1763) * ''Eysarcoris confusus'' Fuente, 1971 * ''Eysarcoris distinctus'' (Schouteden) * ''Eysarcoris ...

'' (Westwood, 1837) * ''

Eysarcoris venustissimus ''Stagonomus venustissimus'', common name woundwort shieldbug, is a species of shieldbug belonging to the family Pentatomidae, subfamily Pentatominae. Scientific name The species was first named by Fabricius as ''Cimex melanocephalus''. Howeve ...

'' (Schrank, 1776) * '' Graphosoma interruptum'' White, 1839 * ''

Graphosoma italicum ''Graphosoma italicum'' is a species of shield bug in the family Pentatomidae. It is also known as the Striped bug (or Italian striped bug) and Minstrel bug. Description ''G. italicum'' can reach a length of . The body is almost round, with a ...

'' (Müller, 1766) * '' Graphosoma lineatum'' (Linnaeus, 1758) * '' Graphosoma melanoxanthum'' Horvath, 1903 * '' Graphosoma semipunctatum'' (Fabricius, 1775) * ''

'' (Stål, 1855) * '' Holcogaster fibulata'' (Germar, 1831) * ''

Holcostethus albipes ''Holcostethus albipes'' is a shield bug belonging to the family Pentatomidae, subfamily Pentatominae. The species was first described by Johan Christian Fabricius in 1781. The head is trapezoidal and the lateral margin of pronotal callus is s ...

'' (Fabricius, 1781) * ''

Holcostethus evae ''Holcostethus'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. Species These 18 species belong to the genus ''Holcostethus''. * '' Holcostethus abbreviatus'' Uhler, 1872 * ''Holcostethus albipes ''Ho ...

'' Ribes, 1988 * '' Holcostethus sphacelatus'' (Fabricius, 1794) * ''

Jalla dumosa Jalla may refer to: *''Jalla! Jalla!'', a 2000 Swedish comedy film directed by Josef Fares * ''Jalla'' (bug), an insect genus in the subfamily Asopinae See also * * Yalla (disambiguation) *Jalla Jalaluhu ''Dhikr'' ( ar, ذِكْر}, , also s ...

'' (Linnaeus, 1758) * '' Leprosoma inconspicuum'' Baerensprung, 1859 * '' Leprosoma stali'' Douglas & Scott, 1868 * '' Leprosoma tuberculatum'' Jakovlev, 1874 * '' Macrorhaphis acuta'' Dallas, 1851 * ''

Mecidea lindbergi ''Mecidea'' is a genus of narrow stink bugs in the family Pentatomidae. There are about eight described species in ''Mecidea''. Species These eight species belong to the genus ''Mecidea'': * ''Mecidea indica'' Dallas, 1851 * '' Mecidea lindbe ...

'' Wagner, 1954 * ''

Mecidea pallidissima ''Mecidea'' is a genus of narrow stink bugs in the family Pentatomidae. There are about eight described species in ''Mecidea''. Species These eight species belong to the genus ''Mecidea'': * '' Mecidea indica'' Dallas, 1851 * '' Mecidea lindb ...

'' Jensen-Haarup, 1922 * '' Menaccarus arenicola'' (Scholz, 1847) * '' Menaccarus deserticola'' Jakovlev, 1900 * '' Menaccarus dohrnianus'' (Mulsant & Rey, 1866) * '' Menaccarus turolensis'' Fuente, 1971 * '' Mustha spinosula'' (Lefèbvre, 1831) * '' Neostrachia bisignata'' (Walker, 1867) * ''

Neottiglossa bifida ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * '' Neottiglossa bifida'' (A.Costa, 1847) * '' Neott ...

'' (A. Costa, 1847) * ''

Neottiglossa flavomarginata ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * '' Neotti ...

'' (Lucas, 1849) * '' Neottiglossa leporina'' (Herrich-Schäffer, 1830) * ''

Neottiglossa lineolata ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * '' Neotti ...

'' (Mulsant & Rey, 1852) * ''

Neottiglossa pusilla ''Neottiglossa'' is a genus of stink bugs in the family Pentatomidae. There are about 10 described species in ''Neottiglossa''. Species These 10 species belong to the genus ''Neottiglossa'': * ''Neottiglossa bifida'' (A.Costa, 1847) * '' Neotti ...

'' (Gmelin, 1790) * ''

Nezara viridula ''Nezara viridula'', commonly known as the southern green stink bug (USA), southern green shield bug (UK) or green vegetable bug (Australia and New Zealand), is a plant-feeding stink bug. Believed to have originated in Ethiopia, it can now be fo ...

'' (Linnaeus, 1758) * ''

Palomena formosa ''Palomena'' is a genus of shield bugs of the family Pentatomidae and tribe Nezarini; species are found in Europe and Asia. Species Species within this genus include: * '' Palomena angulata'' (Puton, 1871) * '' Palomena angulosa'' (Motschulsk ...

'' Vidal, 1940 * ''

Palomena prasina The green shield bug (''Palomena prasina'') is a European shield bug species in the family Pentatomidae. The name might equally apply to several other species in the tribe Nezarini, or if referred-to as a "green stink bug", it might more approp ...

'' (Linnaeus, 1761) * ''

Palomena viridissima ''Palomena viridissima'' is a European species of shield bug in the tribe Nezarini Nezarini is a tribe of stink bugs in the family Pentatomidae. Genera There are at least 21 described genera in the Nezarini including: * ''Acrosternum ...

'' (Poda, 1761) * ''

Pentatoma rufipes The forest bug or red-legged shieldbug (''Pentatoma rufipes'') is a species of shield bug in the family Pentatomidae, commonly found in most of Europe. It inhabits forests, woodlands, orchards, and gardens. Description ''P. rufipes'' can reach a ...

'' (Linnaeus, 1758) * ''

Peribalus congenitus ''Peribalus'' is a genus of shield bugs in the family Pentatomidae. Species *'' Peribalus congenitus'' Putshkov, 1965 *'' Peribalus inclusus'' (Dohrn, 1860) *'' Peribalus manifestus'' (Kiritshenko, 1952 *''Peribalus strictus ''Peribalus ...

'' Putshkov, 1965 * ''

Peribalus inclusus ''Peribalus'' is a genus of shield bugs in the family Pentatomidae. Species *''Peribalus congenitus'' Putshkov, 1965 *'' Peribalus inclusus'' (Dohrn, 1860) *''Peribalus manifestus'' (Kiritshenko, 1952 *''Peribalus strictus ''Peribalus st ...

'' (Dohrn, 1860) * ''

Peribalus strictus ''Peribalus strictus'', common name Vernal Shieldbug, is a species of shield bugs in the family Pentatomidae. Subspecies *''Peribalus strictus capitatus'' Jakovlev, 1889 *''Peribalus strictus strictus'' (Fabricius, 1803) *''Peribalus strict ...

'' (Fabricius, 1803) * '' Perillus bioculatus'' (Fabricius, 1775) * '' Picromerus bidens'' (Linnaeus, 1758) * ''

Picromerus brachypterus ''Picromerus'' is a genus of shield bugs in the family Pentatomidae. Genera * '' Picromerus bidens'' (Linnaeus, 1758) * '' Picromerus brachypterus'' Ahmad & Önder, 1990 * ''Picromerus conformis'' (Herrich-Schäffer, 1841) * ''Picromerus el ...

'' Ahmad & Onder, 1990 * ''

Picromerus conformis ''Picromerus'' is a genus of shield bugs in the family Pentatomidae. Genera * '' Picromerus bidens'' (Linnaeus, 1758) * '' Picromerus brachypterus'' Ahmad & Önder, 1990 * '' Picromerus conformis'' (Herrich-Schäffer, 1841) * ''Picromerus e ...

'' (Herrich-Schäffer, 1841) * '' Picromerus nigridens'' (Fabricius, 1803) * '' Piezodorus lituratus'' (Fabricius, 1794) * ''

Piezodorus punctipes ''Piezodorus'' is a genus of Pentatomidae, a family of shield bugs. Species *''Piezodorus guildinii'' Westwood *'' Piezodorus lituratus'' (Fabricius, 1794) – gorse shield bug *'' Piezodorus punctipes'' Puton, 1889 *''Piezodorus rubrofasciat ...

'' Puton, 1889 * '' Piezodorus teretipes'' (Stål, 1865) * ''

Pinthaeus sanguinipes ''Pinthaeus sanguinipes'' is a species of stink bug ( family Pentatomidae). Description ''Pinthaeus sanguinipes'' can reach a length of . These stink bugs overwinter as adults, females lay their eggs in May and the adults are present from Augus ...

'' (Fabricius, 1781) * '' Podops annulicornis'' Jakovlev, 1877 * '' Podops calligerus'' Horvath, 1887 * '' Podops curvidens'' Costa, 1843 * ''

Podops dilatatus ''Podops''Laporte de Castelnau FL (1833) Mémoire de cinquante espèces nouvelles ou peu connues d'Insectes. ''Annales de la Société entomologique de France'' 1: 386–415. is a genus of Palaearctic bugs, in the family Pentatomidae; it is the ...

'' Puton, 1873 * '' Podops inunctus'' (Fabricius, 1775) * '' Podops rectidens'' Horvath, 1883 * '' Putonia torrida'' Stål, 1872 * '' Rhacognathus punctatus'' (Linnaeus, 1758) * '' Rhaphigaster nebulosa'' (Poda, 1761) * '' Rubiconia intermedia'' (Wolff, 1811) * '' Schyzops aegyptiaca'' (Lefèbvre, 1831) * ''

Sciocoris angularis ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Puton, 1889 * '' Sciocoris angusticollis'' Puton, 1895 * ''

Sciocoris conspurcatus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Klug, 1845 * ''

Sciocoris convexiusculus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Puton, 1874 * '' Sciocoris cursitans'' (Fabricius, 1794) * '' Sciocoris deltocephalus'' Fieber, 1861 * ''

Sciocoris distinctus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Fieber, 1851 * ''

Sciocoris galiberti ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Ribaut, 1926 * '' Sciocoris helferi'' Fieber, 1851 * '' Sciocoris hoberlandti'' Wagner, 1954 * ''

Sciocoris homalonotus ''Sciocoris homalonotus'' is a species of shield bug belonging to the family Pentatomidae, subfamily Pentatominae. It is approximately 6–7 mm in size. The species was first described by Franz Xaver Fieber in 1851. The apex of the cho ...

'' Fieber, 1851 * '' Sciocoris luteolus'' Fieber, 1861 * ''

Sciocoris macrocephalus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Fieber, 1851 * '' Sciocoris maculatus'' Fieber, 1851 * ''

Sciocoris microphthalmus ''Sciocoris microphthalmus'' is a species of stink bug in the family Pentatomidae. It is found in Europe and Northern Asia (excluding China), North America, and Southern Asia. References External links * Sciocorini Articles created by ...

'' Flor, 1860 * ''

Sciocoris modestus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Horvath, 1903 * '' Sciocoris ochraceus'' Fieber, 1861 * '' Sciocoris orientalis'' Linnavuori, 1960 * '' Sciocoris pallens'' Klug, 1845 * ''

Sciocoris pentheri ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Wagner, 1953 * ''

Sciocoris pictus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' Wagner, 1959 * '' Sciocoris sideritidis'' Wollaston, 1858 * '' Sciocoris sulcatus'' Fieber, 1851 * ''

Sciocoris umbrinus ''Sciocoris'' is a genus of shield bugs belonging to the family Pentatomidae, subfamily Pentatominae. The genus was erected by Carl Fredrik Fallén in 1829. Subgenera and species Subgenus ''Aposciocoris'' Wagner, 1965 * '' Sciocoris cerrutii'' ...

'' (Wolff, 1804) * '' Sciocoriscanariensis'' Lindberg, 1953 * ''

Scotinophara sicula ''Scotinophara'' is a genus of Pentatomidae, shield bugs in the Tribe (biology), tribe Podopini and sometimes called "black bugs".Scotinophara subalpina'' (Bergroth, 1893) * '' Stagonomus amoenus'' (Brullé, 1832) * '' Stagonomus bipunctatus'' (Linnaeus, 1758) * '' Stagonomus devius'' Seidenstucker, 1965 * '' Stagonomus grenieri'' (Signoret, 1865) * ''

Staria lunata ''Staria lunata'' is a species of ''shield bug'' belonging to the family Pentatomidae. It is the only species of the genus. Distribution and habitat This species is present in most of Europe ( Albania, Austria, Bosnia and Herzegovina, Bulg ...

'' (Hahn, 1835) * ''

Stenozygum coloratum ''Stenozygum'' is a genus of stink bugs in the family Pentatomidae. It is widely distributed in Australian-Melanesian, Oriental, Palearctic and Ethiopian regions. Description Species in the genus ''Stenozygum'' are associated with plants like ' ...

'' (Klug, 1845) * '' Sternodontus binodulus'' Jakovlev, 1893 * '' Sternodontus obtusus'' Mulsant & Rey, 1856 * '' Tarisa dimidiatipes'' Puton, 1874 * '' Tarisa elevata'' Reuter, 1901 * '' Tarisa flavescens'' Amyot & Serville, 1843 * '' Tarisa pallescens'' Jakovlev, 1871 * '' Tarisa salsolae'' Kerzhner, 1964 * '' Tarisa subspinosa'' (Germar, 1839) * '' Tholagmus flavolineatus'' (Fabricius, 1798) * '' Tholagmus strigatus'' (Herrich-Schäffer, 1835) * '' Trochiscocoris hemipterus'' (Jakovlev, 1879) * '' Trochiscocoris rotundatus'' Horvath, 1895 * '' Troilus luridus'' (Fabricius, 1775) * '' Ventocoris achivus'' (Horvath, 1889) * '' Ventocoris falcatus'' (Cyrillus, 1791) * '' Ventocoris fischeri'' (Herrich-Schäffer, 1851) * '' Ventocoris halophilum'' (Jakovlev, 1874) * '' Ventocoris modestus'' (Jakovlev, 1880) * '' Ventocoris philalyssum'' (Kiritshenko, 1916) * '' Ventocoris ramburi'' (Horvath, 1908) * '' Ventocoris rusticus'' (Fabricius, 1781) * '' Ventocoris trigonus'' (Krynicki, 1871) * '' Vilpianus galii'' (Wolff, 1802) * '' Zicrona caerulea'' (Linnaeus, 1758) Eurydema oleracea.jpg, Image:Chinche hedionda depredadora (Euthyrhynchus floridanus).jpg, '' Euthyrhynchus floridanus'', nymph Image:Troilus luridus 20020828.jpg, '' Troilus luridus'' Image:Pentatomidae spec.jpg, '' Raphigaster nebulosa'' Image:Eurydema ornatum en Tenerife.jpg, '' Eurydema ornatum'' File:Perillus nymph.jpg, '' Perillus'', nymph

References

External links

When Twenty-Six Thousand Stinkbugs Invade Your Home
by

Kathryn Schulz Kathryn Schulz is an American journalist and author. She is a staff writer at ''The New Yorker''. In 2016, she won the Pulitzer Prize for Feature Writing for her article on the risk of a major earthquake and tsunami in the Pacific Northwest. B ...

. ''The New Yorker,'' March 12, 2018. Ed Yong called it a "pure delight."
I.A.D. Robertson: ''The Pentatomoidea (Hemiptera: Heteroptera) of Sub-Saharan Africa : a database''
Malindi, 2009

– photos and information

* I. A. D. Robertson: ''The Pentatomoidea (Hemiptera: Heteroptera) of Sub-Saharan Africa : a database''. Malindi, 2009
Online version in Repository Naturalis Leiden
{{Taxonbar, from=Q652142 Heteroptera families Shield bugs Taxa named by William Elford Leach la:Cimex foetidus