The Sternorrhyncha suborder of the

Hemiptera Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from ...

contains the

aphid Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...

whiteflies Whiteflies are Hemipterans that typically feed on the undersides of plant leaves. They comprise the family Aleyrodidae, the only family in the superfamily Aleyrodoidea. More than 1550 species have been described. Description and taxonomy The A ...

, and

scale insect Scale insects are small insects of the Order (biology), order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient g ...

s, groups which were traditionally included in the now-obsolete order "Homoptera". The name "Sternorrhyncha" refers to the rearward position of the mouthparts relative to the head. Distributed worldwide, all members of this group are plant-feeders, many considered

pests PESTS was an anonymous American activist group formed in 1986 to critique racism, tokenism, and exclusion in the art world. PESTS produced newsletters, posters, and other print material highlighting examples of discrimination in gallery represent ...

feeding on major crops and

ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

s. Many exhibit modified

morphology Morphology, from the Greek and meaning "study of shape", may refer to: Disciplines *Morphology (archaeology), study of the shapes or forms of artifacts *Morphology (astronomy), study of the shape of astronomical objects such as nebulae, galaxies, ...

and/or life cycles, including phenomena such as flightless morphs,

parthenogenesis Parthenogenesis (; from the Greek + ) is a natural form of asexual reproduction in which the embryo develops directly from an egg without need for fertilization. In animals, parthenogenesis means the development of an embryo from an unfertiliz ...

sexual dimorphism Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

, and

eusociality Eusociality ( Greek 'good' and social) is the highest level of organization of sociality. It is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations wit ...

Phylogeny

The

phylogeny A phylogenetic tree or phylogeny is a graphical representation which shows the evolutionary history between a set of species or Taxon, taxa during a specific time.Felsenstein J. (2004). ''Inferring Phylogenies'' Sinauer Associates: Sunderland, M ...

of the extant Sternorrhyncha, inferred from analysis of small subunit (18S)

ribosomal RNA Ribosomal ribonucleic acid (rRNA) is a type of non-coding RNA which is the primary component of ribosomes, essential to all cells. rRNA is a ribozyme which carries out protein synthesis in ribosomes. Ribosomal RNA is transcribed from ribosomal ...

, is shown in the

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

. The evolutionary position of several fossil taxa are unclear. A suggested phylogeny is:

Groups

Well-known groups in the Sternorrhyncha include: *

s – (

Aphididae The Aphididae are a very large insect family in the aphid superfamily ( Aphidoidea), of the order Hemiptera. These insects suck the sap from plant leaves. Several thousand species are placed in this family, many of which are considered plant/cr ...

) * woolly and gall-making aphids (

Eriosomatinae Woolly aphids (subfamily: Eriosomatinae) are sap-sucking insects that produce a filamentous waxy white covering which resembles cotton or wool. The adults are winged and move to new locations where they lay egg masses. The nymphs often form ...

) * pine and spruce aphids (

Adelgidae The Adelgidae are a small family of the Hemiptera closely related to the aphids, and often included in the Aphidoidea with the Phylloxeridae or placed within the superfamily Phylloxeroidea as a sister of the Aphidoidea within the infraorder ...

) * phylloxerans ( Phylloxeridae, including the vine phylloxera) *

– (Aleyrodidae) * jumping plant lice ( Psyllidae and allied families) * Superfamily Coccoidea (

s) ** cottony cushion scales, giant coccids, and ground pearls (

Margarodidae The Margarodidae (illegitimately as Margodidae) or ground pearls are a family of scale insects within the superfamily Coccoidea. Members of the family include the Polish cochineal and Armenian cochineal (genus ''Porphyrophora'') and the original ...

) ** armoured scales (

Diaspididae Diaspididae is the largest family of scale insects with over 2650 described species in around 400 genera. As with all scale insects, the female produces a waxy protective scale beneath which it feeds on its host plant. Diaspidid scales are far mo ...

) **

cochineal The cochineal ( , ; ''Dactylopius coccus'') is a scale insect in the suborder Sternorrhyncha, from which the natural dye carmine is derived. A primarily sessility (motility), sessile parasitism, parasite native to tropical and subtropical Sout ...

insects (Dactylopiidae) ** lac scales (

Kerriidae ''Kerriidae'' is a family of Coccoidea, scale insects, commonly known as lac insects or lac scales. Some members of the genera ''Metatachardia'', ''Tachardiella'', ''Austrotacharidia'', ''Afrotachardina'', ''Tachardina'', and ''Kerria'' are raise ...

, Lacciferidae, Tachardinidae) ** soft scales ( Coccidae) ** pit scales ( Asterolecaniidae) **

mealybug Mealybugs are insects in the family Pseudococcidae, unarmored scale insects found in moist, warm habitats. Of the more than 2,000 described species, many are considered pests as they feed on plant juices of greenhouse plants, house plants and ...

s (Pseudococcidae) ** felted scales ( Eriococcidae)

References

External links

Suborder Sternorrhyncha—Plant-parasitic Hemipterans
at

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{{Authority control Insect suborders