Steinheim Skull
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The Steinheim skull is a
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...
ized
skull The skull, or cranium, is typically a bony enclosure around the brain of a vertebrate. In some fish, and amphibians, the skull is of cartilage. The skull is at the head end of the vertebrate. In the human, the skull comprises two prominent ...
of a ''
Homo neanderthalensis Neanderthals ( ; ''Homo neanderthalensis'' or sometimes ''H. sapiens neanderthalensis'') are an extinct group of archaic humans who inhabited Europe and Western and Central Asia during the Middle to Late Pleistocene. Neanderthal extinctio ...
'' or ''
Homo heidelbergensis ''Homo heidelbergensis'' is a species of archaic human from the Middle Pleistocene of Europe and Africa, as well as potentially Asia depending on the taxonomic convention used. The species-level classification of ''Homo'' during the Middle Pleis ...
'' found on 24 July 1933 near Steinheim an der Murr,
Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...
. It is estimated to be between 250,000 and 350,000 years old. The skull is slightly flattened and has a cranial capacity between 950 and 1280 cc. Sometimes referred to as ''Homo steinheimensis'' in older literature, the original fossil is housed in the State Museum of Natural History in Stuttgart, Germany. Some believe that the Steinheim skull may have belonged to an adult female due to its gracile nature.


Classification

The "primitive man of Steinheim" is a single find. The designation "Steinheim skull" can be seen as a reference to the location of the fossil, but in no way identifies with a certain taxon. The skull shows characteristics of both ''H. heidelbergensis'' and Neanderthals. It is therefore classified by most paleoanthropologists to ''H. heidelbergensis'' and is believed to be a transitional form of ''H. heidelbergensis'' to Neanderthals. This has sometimes been referred to as "pre-Neanderthal.” Reinhard Ziegler: ''4 Millionen Jahre Mensch.'' Spektrum der Wissenschaft, Mai 1999, Seite 130 ff. The inner ear of the fossil has a feature in which Neanderthals and ''H. sapiens'' differ. The location of the semicircular canals of the inner ear in the temporal bone of the skull base is similar to the situation in Neanderthals, while the semicircular canals of the older ''H. erectus,'' as with ''H. sapiens'', are closer. Until the late 1980s, the fossil was sometimes referred to as ''H. sapiens steinheimensis''. During this time Neanderthals were also referred to as ''H. sapiens neanderthalensis''. Today, however, many paleoanthropologists classify Neanderthals and modern humans as separate species, with ''H. heidelbergensis'' their possible last common ancestor (LCA). Therefore that two distinct species are to be considered: ''H. neanderthalensis'' and ''H. sapiens.'' A 2016 study by Chris Stringer found ''
Homo heidelbergensis ''Homo heidelbergensis'' is a species of archaic human from the Middle Pleistocene of Europe and Africa, as well as potentially Asia depending on the taxonomic convention used. The species-level classification of ''Homo'' during the Middle Pleis ...
'' to have not been the common ancestor of Neanderthals and humans, being instead placed entirely on the Neanderthal side of the tree along with
Denisovan The Denisovans or Denisova hominins ( ) are an extinct species or subspecies of archaic human that ranged across Asia during the Lower and Middle Paleolithic, and lived, based on current evidence, from 285 thousand to 25 thousand years ago. D ...
s (close relatives of the Neanderthals) and the Steinheim skull, which was classified as an early Neanderthal fossil. Taking this into account, the LCA of modern humans and Neanderthals needs to be dated even further back, e. g. to ''Homo erectus'', see Neanderthal §Evolution. Sequencing of
genome A genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding genes, other functional regions of the genome such as ...
( aDNA) and
proteome A proteome is the entire set of proteins that is, or can be, expressed by a genome, cell, tissue, or organism at a certain time. It is the set of expressed proteins in a given type of cell or organism, at a given time, under defined conditions. P ...
( ancient protein) of further human fossils has to be done in order to shed light on the relation of ''H. heidelbergensis'' and the Steinheim skull to the pre-neanderthal "Neandersovans", the proposed LCA of Neanderthals and Denisovans. A 2021 study by Chris Stringer found Steinheim sits as a sole sister to a group containing '' H. antecessor and H. longi'', plus ''H.'' ''sapiens.'' Together they are considered as sister to the Neanderthals.


Discovery

Prior to discovery of the Steinheim skull in the gravel pit, many objects such as bones of elephants, rhinos and wild horses had been unearthed. Therefore, the archaeologists excavating the site were already sensitized to possible skeletal remains in the quarry. Fritz Berckhemer traveled on the same day of the skull's discovery and reviewed the still hidden skull in the wall. The next day, together with Max Bock, he began the careful excavation. It was clear, on the basis of the shape and dimensions of the skull, that it was not a monkey, as had initially been suspected. It turned out to be a human skull from the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
. The skull was roughly cleaned, hardened and plastered so it would arrive safe and sound in the Museum of Natural History. The find in Steinheim turned up no other artifacts. There were no artifacts from the people as well as no other skeletal remains found. There were also no tools such as stone tools or bone implements discovered at the site. It can be assumed, however, that the hominin would have been capable of creating tools and using them for different purposes. For example, evidence of this can be found around the same time frame of “ Swanscombe man", where they found some fist wedges from the Acheulean tool culture.


Brain tumor

The investigations completed at the Eberhard Karls University of Tuebingen in 2003 by Alfred Czarnetzki, Carsten M. Pusch and Erwin Schwaderer, showed that the owner of the skull suffered from a meningioma, which is an arachnoid tumor. Meningiomas are a diverse set of tumors that arise from the
meninges In anatomy, the meninges (; meninx ; ) are the three membranes that envelop the brain and spinal cord. In mammals, the meninges are the dura mater, the arachnoid mater, and the pia mater. Cerebrospinal fluid is located in the subarachnoid spac ...
, which is the membranous layers surrounding the central nervous system. The slow-growing tumor had a size of 51 mm × 43 mm × 25 mm and a volume of . It is believed that this tumor may have caused headaches. It is also possible that no neurological deficiencies were suffered, due to the slow growing nature of meningiomas. Whether the tumor should be considered the cause of death cannot be determined from the remains. Having the rest of the skeleton would be necessary for further investigation into a possible cause of death. Meningiomas are very rare with roughly 2 out of 100,000 causing symptoms, so finding evidence of one in such ancient remains is a very exciting discovery.idw-online vom 11. August 2003: ''Tübinger Forscher finden erstmals Schädeltumor bei frühen Menschen.''
Abbildung des Schädels (Memento vom 16. Oktober 2008 im ''Internet Archive'')
It is the earliest evidence of a meningioma tumor on record.


See also

* List of fossil sites ''(with link directory)'' * List of hominina (hominid) fossils ''(with images)''


References


External links

* {{portal bar, Evolutionary biology, Paleontology Homo heidelbergensis fossils