Steinernema
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''Steinernema'' is a genus of
nematode The nematodes ( or ; ; ), roundworms or eelworms constitute the phylum Nematoda. Species in the phylum inhabit a broad range of environments. Most species are free-living, feeding on microorganisms, but many are parasitic. Parasitic worms (h ...
s in the family of Steinernematidae. The genus ''Steinernema'' is named after the nematologist Gotthold Steiner. Nematodes of this genus have been shown to be effective as a biological control for agricultural pests of the Scarabaeidae family, such as '' Maladera formosae''.


Life cycle

Species form symbiotic relationships with '' Xenorhabdus'' and '' Photorhabdus'' bacteria. The free-living third stage juvenile, termed a dauer juvenile, enters its insect hosts through natural openings, such as the mouth, anus, and spiracles. Bacterial cells from the intestines are regurgitated into the insect. The insect hemolymph provides a rich medium for the bacterial cells which grow, releasing toxins and exoenzymes, causing the insect host to die from septicemia. The bacteria also produce other compounds to protect the insect from other microbes in the soil. The nematode moves out of its developmentally arrested third, nonfeeding stage, triggered by either bacterial or insect food signals. The nematodes feed on the bacteria and moult to the fourth stage, reaching adulthood within a few days. with separate male and female individuals. Nematode development continues for two to three generations. When insufficient nutrient remain adult development is suppressed. Developmentally arrested third stage juveniles accumulate and emerge into the soil, where they may survive for months until a new suitable host is found.


Species

The following species have been described:


Use in agriculture

Species of this genus can infect insects and are used as a biopesticide to infect agricultural pests. They can be used against a wide variety of species, including
weevils Weevils are beetles belonging to the Taxonomic rank, superfamily Curculionoidea, known for their elongated snouts. They are usually small – less than in length – and Herbivore, herbivorous. Approximately 97,000 species of weevils are known. ...
, cutworms, gnats and mole crickets.


References


Further reading

* * * {{Taxonbar, from=Q5304420 Rhabditida Rhabditida genera