Spilomelinae is a very species-rich subfamily of the lepidopteran family

Crambidae The Crambidae are the grass moth family of lepidopterans. They are variable in appearance, the nominal subfamily Crambinae (grass moths) taking up closely folded postures on grass stems where they are inconspicuous, while other subfamilies includ ...

, the crambid snout

moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of ...

s. With 4,135 described species in 344 genera worldwide, it is the most speciose group among pyraloids.

Description

Imagines – the adult life stage – vary considerably in size: the forewing span ranges from 11.5 mm e.g. in '' Metasia'' to 50 mm in the robust-bodied '' Eporidia''. In resting position, the moths exhibit a characteristic triangular shape, with the wings usually folded over the abdomen, the forewings covering the hindwings. Some Spilomelinae diverge from this common resting pattern, like '' Maruca'' with widely spread wings, and '' Atomopteryx'' and '' Lineodes'' with narrow wings folded along the body. All Spilomelinae moths have well developed

compound eyes A compound eye is a visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidia, which are tiny independent photoreception units that consist of a cornea, lens, and photoreceptor cells which distin ...

, antennae and

mouthparts Mouthparts may refer to: * The parts of a mouth ** Arthropod mouthparts *** Insect mouthparts {{disambig ...

, although in the genera '' Niphopyralis'' and '' Siga'' the proboscis is lost. Synapomorphic characters of the subfamily comprise minute or obsolete maxillary palpi, ventrally projecting fornix tympani, and the female genitalia's ductus bursae with a weak

sclerotization Sclerotization is a biochemical process that produces the rigid shell of sclerotin that comprises an insect's chitinous exoskeleton. It is prominent in the thicker, armored parts of insects and arachnids, especially in the biting mouthparts and scl ...

or a granulose texture. The moths are furthermore characterized by an often bilobed praecinctorium, pointed spinula, and the absence of

chaetosemata This glossary of entomology describes terms used in the formal study of insect species by entomologists. A–C A synthetic chlorinated hydrocarbon insecticide, toxic to vertebrates. Though i ...

and of a retinacular hook. A gnathos or pseudognathos can be present or absent and is therefore of little

diagnostic Diagnosis is the identification of the nature and cause of a certain phenomenon. Diagnosis is used in many different disciplines, with variations in the use of logic, analytics, and experience, to determine " cause and effect". In systems engine ...

value, except for several genera of Agroterini, where the gnathos has a well-developed medial process.

Food plants

Food plant use is diverse within Spilomelinae, ranging from

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes except ...

s over

gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμν ...

s to a wide spectrum of

angiosperm Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of br ...

s. The larvae of '' Niphopyralis'' live in nests of weaver ants, where they feed on ant larvae. Steniini caterpillars are

detritivore Detritivores (also known as detrivores, detritophages, detritus feeders, or detritus eaters) are heterotrophs that obtain nutrients by consuming detritus (decomposing plant and animal parts as well as feces). There are many kinds of invertebrate ...

s. Many Spilomelinae tribes have a narrow food spectrum, with the larvae feeding on plants of only one or a few plant families, e.g. Lineodini on

Solanaceae The Solanaceae , or nightshades, are a family of flowering plants that ranges from annual and perennial herbs to vines, lianas, epiphytes, shrubs, and trees, and includes a number of agricultural crops, medicinal plants, spices, weeds, and o ...

, Hydririni primarily on

Sapindaceae The Sapindaceae are a family of flowering plants in the order Sapindales known as the soapberry family. It contains 138 genera and 1858 accepted species. Examples include horse chestnut, maples, ackee and lychee. The Sapindaceae occur in temp ...

and

Convolvulaceae Convolvulaceae (), commonly called the bindweeds or morning glories, is a family of about 60 genera and more than 1,650 species. These species are primarily herbaceous vines, but also include trees, shrubs and herbs. The tubers of several spec ...

, and Trichaeini on

Rubiaceae The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules ...

Economic impact

A number of Spilomelinae are considered "pest species", with their larvae feeding on a variety of economically important crops. Notable representatives are the genera of ''

Leucinodes ''Leucinodes'' is a genus of moths of the family Crambidae. It was first described by Achille Guenée in 1854. ''Leucinodes'' species have been documented as eggplant fruit borers. They are occasionally imported by accident from African and As ...

'' and '' Neoleucinodes'' with larvae feeding on

, '' Cnaphalocrocis'' and '' Marasmia'' damaging Poaceae like ''

Oryza ''Oryza'' is a genus of plants in the grass family. It includes the major food crop rice (species ''Oryza sativa'' and ''Oryza glaberrima''). Members of the genus grow as tall, wetland grasses, growing to tall; the genus includes both annual a ...

'', ''

Sorghum ''Sorghum'' () is a genus of about 25 species of flowering plants in the grass family ( Poaceae). Some of these species are grown as cereals for human consumption and some in pastures for animals. One species is grown for grain, while many ot ...

'' and '' Zea'', the legume pod borers of the genus '' Maruca'' on Fabaceae and

Amaranthaceae Amaranthaceae is a family of flowering plants commonly known as the amaranth family, in reference to its type genus '' Amaranthus''. It includes the former goosefoot family Chenopodiaceae and contains about 165 genera and 2,040 species, making i ...

, and '' Spoladea'', who feeds on a variety of different agriculturally important plant families. The box tree moth, ''

Cydalima perspectalis ''Cydalima perspectalis'' or the box tree moth is a species of moth of the family Crambidae, first described by Francis Walker, the English entomologist, in 1859. Native to Japan, China, Taiwan, Korea, far-east Russia and India, it has in ...

'', whose larvae feed on box trees, a prominent ornamental plant in many parks and gardens, has been accidentally introduced to Europe in the mid-2000s and to North America in 2018.

Systematics

Until the late 1990s, Spilomelinae were included in the subfamily

Pyraustinae Pyraustinae is a large subfamily of the lepidopteran family Crambidae, the crambid snout moths. It currently includes over 1,400 species; most of them tropical but some found in temperate regions including both North America and Europe. The P ...

tribe The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...

Spilomelini. In the past, Spilomelinae were believed to be

polyphyletic A polyphyletic group is an assemblage of organisms or other evolving elements that is of mixed evolutionary origin. The term is often applied to groups that share similar features known as homoplasies, which are explained as a result of conver ...

. However, a recent phylogenetic study by Mally et al. (2019), based on molecular genetic and morphological data, found the subfamily to be

monophyletic In cladistics for a group of organisms, monophyly is the condition of being a clade—that is, a group of taxa composed only of a common ancestor (or more precisely an ancestral population) and all of its lineal descendants. Monophyletic ...

. The study's authors furthermore proposed 13

tribes The term tribe is used in many different contexts to refer to a category of human social group. The predominant worldwide usage of the term in English is in the discipline of anthropology. This definition is contested, in part due to confl ...

within Spilomelinae: * Agroterini Acloque, 1897 * Asciodini Mally, Hayden, Neinhuis, Jordal & Nuss, 2019 * Herpetogrammatini Mally, Hayden, Neinhuis, Jordal & Nuss, 2019 * Hydririni Minet, 1982 * Hymeniini Swinhoe, 1900 * Lineodini Amsel, 1956 * Margaroniini Swinhoe & Cotes, 1889 * Nomophilini Kuznetzov & Stekolnikov, 1979 * Spilomelini Guenée, 1854 * Steniini Guenée, 1854 * Trichaeini Mally, Hayden, Neinhuis, Jordal & Nuss, 2019 *

Udeini Udeini is a Tribe (biology), tribe in the species-rich subfamily Spilomelinae in the Pyraloidea, pyraloid moth Family (biology), family Crambidae. The tribe was erected by Richard Mally, James E. Hayden, Christoph Neinhuis, Bjarte H. Jordal and M ...

Mally, Hayden, Neinhuis, Jordal & Nuss, 2019 *

Wurthiini Wurthiini is a tribe of the species-rich subfamily Spilomelinae in the pyraloid moth family Crambidae. Description Adult Wurthiini are small to medium-sized moths with wing spans of normally 20 to over 30 mm, whereas adults of ''Niphopy ...

Roepke, 1916

"Non-euspilomeline" tribes

Hydririni, Lineodini,

and

share several

plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, an ...

characters with the

sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

of Spilomelinae, the

. These plesiomorphies are: absence of longitudinal strips on the male abdominal segment 8; male genitalia with a straight to concave valva costa and a

phallus A phallus is a penis (especially when erect), an object that resembles a penis, or a mimetic image of an erect penis. In art history a figure with an erect penis is described as ithyphallic. Any object that symbolically—or, more precise ...

with an evenly sclerotized

apodeme An exoskeleton (from Greek ''éxō'' "outer" and ''skeletós'' "skeleton") is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton (endoskeleton) in for example, a human. In usage, some of the l ...

; female genitalia with a lanceolate “ediacaroid” signum in the corpus bursae and in several taxa with an appendix bursae attached to the corpus bursae. Due to these plesiomorphies, these four Spilomelinae tribes are referred to as "non-euspilomeline" tribes as opposed to the

"euspilomeline" group that represents a more derived group of Spilomelinae. The "non-euspilomeline" tribes form a paraphylum as they do not comprise the "euspilomelines".

"Euspilomeline" tribes

Pantographa limata – Basswood Leafroller Moth

Glyphodes stolalis (Crambidae Spilomelinae)

- 5268 – Psara obscuralis – Obscure Psara Moth (15439954193)

Duponchelia fovealis, Trawscoed, North Wales, July 2006 (19888933425)

The tribes Agroterini, Margaroniini, Spilomelini, Herpetogrammatini, Hymeniini, Asciodini, Trichaeini, Steniini and Nomophilini form the monophylum of "euspilomelines" (Greek ''eu-'' for "good" or "true"), all sharing a common ancestor. The

synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ...

of euspilomelines are: male abdominal tergite 8 with an emarginate anterior edge; male genitalia with partly sclerotized hair pencils on the anterior edge of vinculum-tegumen connection, with a convex valva costa, and the

without a coecum and the

of the phallus

reduced to a ventral, longitudinally sclerotized strip along the manica; female genitalia with a longitudinal membranous strip in the antrum, and the lack of a strongly sclerotised colliculum between antrum and ductus seminalis.

Genera currently unplaced in any Spilomelinae tribe

Based on the morphological

and characteristics of these tribes, Mally et al. (2019) assigned numerous Spilomelinae genera to these proposed tribes, so that about two thirds of the 339 Spilomelinae

genera Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial ...

are placed in these tribes, leaving 132 genera currently unplaced: *'' Aboetheta'' Turner, 1914 *'' Acicys'' Turner, 1911 *''

Aediodina ''Aediodina'' is a genus of moths of the family Crambidae. It contains only one species, ''Aediodina quaternalis'', which is found on Ambon Island and Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia, at t ...

'' Strand, 1919 *'' Agrammia'' Guenée, 1854 *'' Almonia'' Walker, 1866 *''

Ametrea ''Ametrea'' is a genus of moths of the family Crambidae. It contains only one species, ''Ametrea nebulalis'', which is found on the Sula Islands. References Spilomelinae Monotypic moth genera Moths of Indonesia Crambidae genera Taxa ...

'' Munroe, 1964 *'' Archernis'' Meyrick, 1886 (= ''Chrysommatodes'' Warren, 1896, ''Metoportha'' Meyrick, 1894, ''Protonoceras'' Warren, 1890) *'' Arxama'' Walker, 1866 *''

Atelocentra ''Atelocentra'' is a genus of moths of the family Crambidae. It contains only one species, ''Atelocentra chloraspis'', which is found in Australia, where it has been recorded from Queensland, New South Wales and Victoria. The wingspan is a ...

'' Meyrick, 1884 *''

Auchmophoba ''Auchmophoba'' is a genus of moths of the family Crambidae. Species *''Auchmophoba alternata'' (Warren, 1895) *''Auchmophoba costastrigalis'' (Hampson, 1896) Former species *''Auchmophoba tynnuta'' Turner, 1913 *''Auchmophoba sufetuloides'' Ham ...

'' Turner, 1913 *'' Bacotoma'' Moore, 1885 (= ''Epactoctena'' Meyrick, 1937, ''Platamonia'' Lederer, 1863, ''Platamonina'' J. C. Shaffer & Munroe, 2007) *'' Cangetta'' Moore, 1886 *''

Carthade ''Carthade'' is a genus of moths of the family Crambidae. It contains only one species, ''Carthade caecalis'', which is found in Colombia Colombia (, ; ), officially the Republic of Colombia, is a country in South America with insular ...

'' Snellen, 1899 *'' Chabula'' Moore, 1886 *'' Chromodes'' Guenée, 1854 *''

Cissachroa ''Cissachroa'' is a monotypic moth genus of the family Crambidae described by Alfred Jefferis Turner in 1937. Its only species, ''Cissachroa callischema'', described in the same article, is found in Australia Australia, officially the C ...

'' Turner, 1937 *'' Coelorhyncidia'' Hampson, 1896 *'' Coptobasis'' Lederer, 1863 *'' Coremata'' Amsel, 1956 *''

Cotachena ''Cotachena'' is a genus of moths of the family Crambidae. Species *'' Cotachena aluensis'' Butler, 1887 *'' Cotachena alysoni'' Whalley, 1961 *'' Cotachena brunnealis'' Yamanaka, 2001 *'' Cotachena fuscimarginalis'' Hampson, 1916 *'' Cotachen ...

'' Moore, 1885 *'' Criophthona'' Meyrick, 1884 *'' Camptomastix'' Warren, 1892 (= ''Camptomastyx'' Hampson, 1896) *'' Cangetta'' Moore, 1886 (= ''Blechrophanes'' Turner, 1937) *''

'' Snellen, 1899 (= ''Chartade'' Neave, 1939) *''

Cavifrons ''Lineostriastiria biundulalis'' is a moth of the family Noctuidae. It is found in North America, where it has been recorded from Arizona, Oklahoma and Texas. The wingspan The wingspan (or just span) of a bird or an airplane is the distance ...

'' Zeller, 1872 *'' Ceratarcha'' Swinhoe, 1894 *'' Chabula'' Moore, 1886 *'' Chromodes'' Guenée, 1854 *''

'' Turner, 1937 *'' Coelorhyncidia'' Hampson, 1896 *'' Coptobasis'' Lederer, 1863 *'' Coremata'' Amsel, 1956 (= ''Culcita'' Amsel, 1957) *''

'' Moore, 1885 (= ''Mesothyris'' Warren, 1892, ''Syndicastis'' Meyrick, 1889) *'' Criophthona'' Meyrick, 1884 (= ''Conoprora'' Turner, 1913) *''

Daulia Daulis ( grc, Δαυλίς), at a later time Daulia (Δαυλία), and also Daulium or Daulion (Δαύλιον), was a town of ancient Phocis, near the frontiers of Boeotia, and on the road from Orchomenus and Chaeroneia to Delphi. Overview ...

'' Walker, 1859 (= ''Girtexta'' Swinhoe, 1890) *'' Deuterarcha'' Meyrick, 1884 *''

Deuterophysa ''Deuterophysa'' is a genus of moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approxima ...

'' Warren, 1889 *'' Dichocrocis'' Lederer, 1863 (= ''Zebrodes'' Warren, 1896) *'' Discothyris'' Warren, 1895 *'' Dracaenura'' Meyrick, 1886 *'' Duzulla'' Amsel, 1952 *'' Ebuleodes'' Warren, 1896 *'' Ectadiosoma'' Turner, 1937 *'' Elbursia'' Amsel, 1950 *'' Eudaimonisma'' T. P. Lucas, 1902 *'' Eulepte'' Hübner, 1825 (= ''Acrospila'' Lederer, 1863) *''

Eurybela ''Eurybela'' is a genus of moths of the family Crambidae. Species *'' Eurybela scotopis'' Turner, 1908 *'' Eurybela trophoessa'' (Turner, 1908) References Spilomelinae Crambidae genera Taxa named by Alfred Jefferis Turner {{Spilo ...

'' Turner, 1908 *'' Eustenia'' Snellen, 1899 *''

Furcivena ''Furcivena'' is a genus of moths of the family Crambidae. Species *''Furcivena cyanoxantha'' Meyrick, 1933 *''Furcivena dialithalis'' Hampson, 1917 *''Furcivena euclidialis'' (Hampson, 1906) *''Furcivena rhodoneurialis'' Hampson, 1898 *''Furcive ...

'' Hampson, 1896 *'' Gadessa'' Moore, 1885 *''

Gethosyne ''Gethosyne'' is a genus of moths of the family Crambidae. It contains only one species, ''Gethosyne aequivocalis'', which is found in Meghalaya, in Northeast India , native_name_lang = mni , settlement_type = , image_ ...

'' Warren, 1896 *'' Glauconoe'' Warren, 1892 *'' Goniorhynchus'' Hampson, 1896 *'' Hemopsis'' Kirti & Rose, 1987 *'' Heterudea'' Dognin, 1905 *'' Hyalea'' Guenée, 1854 *'' Indogrammodes'' Kirti & Rose, 1989 *'' Ischnurges'' Lederer, 1863 (= ''Nesolocha'' Meyrick, 1886) *'' Legrandellus'' J. C. Shaffer & Munroe, 2007 *'' Lepidoneura'' Hampson, 1896 *''

Leucinodella Agroterini is a tribe of the species-rich subfamily Spilomelinae in the pyraloid moth family Crambidae. The tribe was erected by Alexandre Noël Charles Acloque in 1897. Genera Agroterini currently contains 30 genera, altogether comprising ...

'' Strand, 1918 *'' Leucochromodes'' Amsel, 1956 *'' Leucophotis'' Butler, 1886 *''

Lipararchis ''Lipararchis'' is a genus of moths of the family Crambidae. Species *''Lipararchis aspilus'' (Turner, 1915), described from Australia in the genus ''Calamochrous'' *''Lipararchis hyacinthopa'' Meyrick, 1934, the type species, described from Vuni ...

'' Meyrick, 1934 *'' Luma'' Walker, 1863 (= Pelena Moore, 1886, Pelina Hampson, 1897, ''Petena'' Neave, 1940) *'' Mabra'' Moore, 1885 (type species: ''Asopia eryxalis'' Walker, 1859, by original designation) *'' Macaretaera'' Meyrick, 1886 (= ''Trichoptychodes'' Swinhoe, 1894, ''Trigonophylla'' Turner, 1937) *'' Macrobela'' Turner, 1939 *'' Malaciotis'' Meyrick, 1934 *'' Malickyella'' Mey & Speidel, 2010 *'' Meekiaria'' Munroe, 1974 *'' Merodictya'' Warren, 1896 *'' Mesocondyla'' Lederer, 1863 *'' Metallarcha'' Meyrick, 1884 (= ''Panopsia'' Turner, 1913) *'' Metoeca'' Warren, 1896 *'' Metraeopsis'' Dognin, 1905 *''

Microgeshna ''Microgeshna'' is a genus of moths of the family Crambidae. It contains only one species, ''Microgeshna laportei'', which is found on the Seychelles, where it has been recorded from Aldabra Aldabra is the world's second-largest coral ato ...

'' J. C. Shaffer & Munroe, 2007 *''

Microphysetica ''Microphysetica'' is a genus of moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approxim ...

'' Hampson, 1917 (= ''Falx'' Amsel, 1956, ''Falcimorpha'' Amsel, 1957) *'' Mimudea'' Warren, 1892 *'' Mukia'' Amsel, 1954 *'' Myriostephes'' Meyrick, 1884 *'' Myrmidonistis'' Meyrick, 1887 *''

Nacoleia Nakoleia ( el, Νακώλεια and Νακόλεια) also known as Nakolaion (Νακώλαιον), Latinized as Nacolia or Nacolea, was an ancient and medieval city in Phrygia. It corresponds to present-day Seyitgazi, Eskişehir Province in t ...

'' Walker, 1859 (= ''Aplomastix'' Warren, 1890, ''Orthocona'' Warren, 1896, ''Semioceros'' Meyrick, 1884) *'' Nankogobinda'' Rose & Kirti, 1986 *'' Nausinoe'' Hübner, 1825 (= ''Lepyrodes'' Guenée, 1854, ''Phalangiodes'' Guenée, 1854) *'' Nausinoella'' J. C. Shaffer & Munroe, 2007 *'' Neostege'' Hampson, 1910 *'' Nevrina'' Guenée, 1854 *'' Nistra'' Walker, 1859 *'' Notesia'' Yamanaka, 1992 *'' Ommatobotys'' J. C. Shaffer & Munroe, 2007 *''

Orphnophanes ''Orphnophanes'' is a genus of moths of the family Crambidae described by Julius Lederer (entomologist), Julius Lederer in 1863. Species *''Orphnophanes ankarampotsyalis'' Marion & Viette, 1956 *''Orphnophanes eucerusalis'' (Walker, 1859) *''Orph ...

'' Lederer, 1863 (= ''Syntomodora'' Meyrick, 1894) *'' Orthospila'' Warren, 1890 *''

Osiriaca ''Osiriaca'' is a monotypic moth genus of the family Crambidae described by Francis Walker in 1866. Its only species, ''Osiriaca ptousalis'', described by the same author in 1859, is known from Australia Australia, officially the Comm ...

'' Walker, 1866 (= ''Myriotis'' Meyrick, 1885) *'' Otiophora'' Turner, 1908 *'' Paramecyna'' Amsel, 1961 *'' Pectinobotys'' Munroe, 1959 *'' Pelinopsis'' Dognin, 1905 *'' Physematia'' Lederer, 1863 *''

Piletocera ''Piletocera'' is a genus of moths of the family Crambidae. The genus was first described by Julius Lederer in 1863. Species *'' Piletocera aegimiusalis'' (Walker, 1859) *'' Piletocera aequalis'' Walker, 1866 *'' Piletocera agathanalis'' Sch ...

'' Lederer, 1863 (= ''Alutefa'' Swinhoe, 1900, ''Danaga'' Moore, 1885, ''Diplotyla'' Meyrick, 1886, ''Ellogima'' Turner, 1913, ''Erebangela'' Meyrick, 1886, ''Graphicopoda'' Butler, 1886, ''Hormatholepis'' Butler, 1886, ''Ptilaeola'' Meyrick, 1886, ''Rinecera'' Butler, 1884, ''Sematosopha'' Meyrick, 1937, ''Strepsimela'' Meyrick, 1886) *''

Plateopsis ''Plateopsis'' is a monotypic moth genus of the family Crambidae described by William Warren in 1896. Its only species, ''Plateopsis vespertilio'', described in the same article, is found in Meghalaya Meghalaya (, or , meaning "abode of cl ...

'' Warren, 1896 *'' Platygraphis'' Dyar, 1918 *'' Pleonectoides'' Hampson, 1891 *'' Polythlipta'' Lederer, 1863 *''

Praeacrospila ''Praeacrospila'' is a genus of moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approxim ...

'' Amsel, 1956 *'' Praephostria'' Amsel, 1956 *'' Pramadea'' Moore, 1888 *''

Preneopogon ''Preneopogon'' is a genus of moth Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximat ...

'' Warren, 1896 *''

Prionopaltis ''Prionopaltis'' is a genus of moths of the family Crambidae described by William Warren in 1892. Species *'' Prionopaltis consocia'' Warren, 1892 *'' Prionopaltis sericea'' Warren, 1892 *'' Prionopaltis subdentalis'' Swinhoe, 1894 Referenc ...

'' Warren, 1892 *'' Prorodes'' Swinhoe, 1894 (= ''Idiostrophe'' Warren, 1896) *'' Proternia'' Meyrick, 1884 *'' Protonoceras'' Warren, 1890 *'' Pycnarmon'' Lederer, 1863 (= ''Aripana'' Moore, 1886, ''Entephria'' Lederer, 1863, ''Eutrichotis'' Swinhoe, 1900, ''Pyralocymatophora'' Strand, 1918, ''Satanastra'' Meyrick, 1890) *'' Pylartes'' Walker, 1863 *'' Pyradena'' Munroe, 1958 *'' Ravanoa'' Moore, 1885 *'' Rehimena'' Walker, 1866 *'' Rhimphaliodes'' Hampson, 1893 (= ''Rhimphaleodes'' Hampson, 1896) *''

Rhynchetria ''Rhynchetria'' is a genus of moths of the family Crambidae. It contains only one species, ''Rhynchetria damasales'', which is found on Java. References Natural History Museum Lepidoptera genus database Pyraustinae Crambidae genera Mono ...

'' Klunder van Gijen, 1913 *'' Sagariphora'' Meyrick, 1894 *'' Sameodesma'' Hampson, 1918 *'' Sedenia'' Guenée, 1854 *'' Sericophylla'' Turner, 1937 (= ''Sericophora'' Turner, 1937) *'' Stenorista'' Dognin, 1905 *'' Syllepte'' Hübner, 1823 (= ''Arthriobasis'' Warren, 1896, ''Haitufa'' Swinhoe, 1900, ''Haliotigris'' Warren, 1896, ''Neomabra'' Dognin, 1905, ''Nothosalbia'' Swinhoe, 1900, ''Polycorys'' Warren, 1896, ''Subhedylepta'' Strand, 1918, ''Sylepta'' Hübner, 1825, ''Syllepta'' Hübner, 1826, ''Troctoceras'' Dognin, 1905) *'' Syngropia'' Hampson, 1912 *'' Syntrita'' Dognin, 1905 *'' Tabidia'' Snellen, 1880 *'' Tanaophysopsis'' Munroe, 1964 *''

Tasenia ''Tasenia'' is a genus of moths of the family Crambidae. It contains only one species, ''Tasenia nigromaculalis'', which is found on Java. References Natural History Museum Lepidoptera genus database Pyraustinae Crambidae genera Taxa na ...

'' Snellen, 1901 *'' Torqueola'' Swinhoe, 1906 *'' Trichoceraea'' Sauber in G. Semper, 1902 *''

Trigonobela ''Trigonobela'' is a genus of moths of the family Crambidae described by Alfred Jefferis Turner in 1915. Species *''Trigonobela nebridopepla'' Turner, 1915 *''Trigonobela perfenestrata'' (Butler, 1882) References

Spilomelinae Crambidae ...

'' Turner, 1915 *'' Trithyris'' Lederer, 1863 *'' Troctoceras'' Dognin, 1905 *'' Tylostega'' Meyrick, 1894 *''

Voliba ''Voliba'' is a genus of moths of the family Crambidae described by Walker in 1866. Species *''Voliba asphyctopa'' Turner, 1908 *''Voliba gigantea'' Hampson, 1912 *''Voliba leptomorpha'' Turner, 1908 *''Voliba psammoessa'' Turner, 1908 *''Voliba ...

'' Walker, 1866 (= ''Gabrisa'' Walker, 1866, ''Stereocopa'' Meyrick, 1885) *'' Xanthomelaena'' Hampson, 1896 *'' Zagiridia'' Hampson, 1897

Genera excluded from Spilomelinae

'' Aporocosmus'' Butler, 1886 was transferred to Odontiinae, '' Orthoraphis'' Hampson, 1896 to

Lathrotelinae Lathrotelinae is a subfamily of the pyraloid family Crambidae described by John Frederick Gates Clarke in 1971. It currently comprises 45 species in six genera. Description Characteristic features of the Lathrotelinae are the undulating wing o ...

, '' Hydropionea'' Hampson, 1917 and ''

Plantegumia ''Plantegumia'' is a genus of moths of the family Crambidae. The three described species are distributed in Central and South America. The genus was formerly treated in Spilomelinae, but is now placed in Glaphyriinae Glaphyriinae is a subf ...

'' Amsel, 1956 to

Glaphyriinae Glaphyriinae is a subfamily of the lepidopteran family Crambidae. It was described by William Trowbridge Merrifield Forbes in 1923. The subfamily currently comprises 509 species in 75 genera. The larvae of Glaphyriinae predominantly feed on p ...

, and '' Prooedema'' Hampson, 1896 to

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{{Taxonbar, from=Q2310651 Moth subfamilies Taxa named by Achille Guenée