Song Birds
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A songbird is a
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
belonging to the
suborder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
Passeri of the perching birds (
Passeriformes A passerine () is any bird of the order Passeriformes (; from Latin 'sparrow' and '-shaped') which includes more than half of all bird species. Sometimes known as perching birds, passerines generally have an anisodactyl arrangement of their ...
). Another name that is sometimes seen as the scientific or vernacular name is Oscines, from
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
''oscen'', "songbird". The Passeriformes contains 5,000 or so speciesEdwards, Scott V. and John Harshman. 2013. Passeriformes. Perching Birds, Passerine Birds. Version 06 February 2013 (under construction). http://tolweb.org/Passeriformes/15868/2013.02.06 in The Tree of Life Web Project, http://tolweb.org/ ccessed 2017/12/11 found all over the world, in which the vocal organ typically is developed in such a way as to produce a diverse and elaborate
bird song Bird vocalization includes both bird calls and bird songs. In non-technical use, bird songs (often simply ''birdsong'') are the bird sounds that are melodious to the human ear. In ornithology and birding, songs (relatively complex vocalization ...
. Songbirds form one of the two major lineages of extant perching birds (~4,000 species), the other being the
Tyranni The Tyranni (suboscines) are a suborder of passerine birds that includes more than 1,000 species, a large majority of which are South American. It is named after the type genus '' Tyrannus''. These have a different anatomy of the syrinx musculat ...
(~1,000 species), which are most diverse in the
Neotropics The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeog ...
and absent from many parts of the world. The Tyranni have a simpler
syrinx In classical Greek mythology, Syrinx () was an Arcadian nymph and a follower of Artemis, known for her chastity. Being pursued by Pan, she fled into the river Ladon, and at her own request was metamorphosed into a reed from which Pan then mad ...
musculature, and while their vocalizations are often just as complex and striking as those of songbirds, they are altogether more mechanical sounding. There is a third perching bird lineage, the
Acanthisitti The New Zealand wrens are a family (Acanthisittidae) of tiny passerines endemic to New Zealand. They were represented by seven Holocene species in four or five genera, although only two species in two genera survive today. They are understood to ...
of
New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...
, of which only two species remain alive today. Recent estimates indicate that songbirds originated 50 million years ago. The distribution of their basal lineages suggest that their origin and initial diversification occurred exclusively in the
Australian continent The continent of Australia, sometimes known in technical contexts as Sahul (), Australia-New Guinea, Australinea, or Meganesia to distinguish it from the country of Australia, is located within the Southern and Eastern hemispheres, near t ...
and only about 40 million years ago, oscines started to colonize
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
,
Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
, and eventually the
Americas The Americas, sometimes collectively called America, are a landmass comprising the totality of North America and South America.''Webster's New World College Dictionary'', 2010 by Wiley Publishing, Inc., Cleveland, Ohio. When viewed as a sing ...
.


Description

The song in this clade is essentially territorial, because it communicates the identity and whereabouts of an individual to other birds, and also signals sexual intentions. Sexual selection among songbirds is highly based on mimetic vocalization. Female preference has shown in some populations to be based on the extent of a male's song repertoire. The larger a male's repertoire, the more females a male individual attracts. It is not to be confused with bird calls that are used for alarms and contact and are especially important in birds that feed or migrate in flocks. While almost all living birds give calls of some sort, well-developed songs are only given by a few lineages outside the songbirds. And still, not all songbirds proffer a call that is distinctly melodious. Songbirds do, however, possess a highly developed vocal organ, the
syrinx In classical Greek mythology, Syrinx () was an Arcadian nymph and a follower of Artemis, known for her chastity. Being pursued by Pan, she fled into the river Ladon, and at her own request was metamorphosed into a reed from which Pan then mad ...
, that enables their sonorous activity. This organ, also known as a song box, can be found where the windpipe meets diverging bronchial tubes which lead to the lungs. The organ is a solid, bony structure lined with a film of membranes which air passes through as the songbird calls. While the song boxes of songbirds vary in size and intricacy, this does not necessarily determine the songbird's ability to voice their song. Researchers believe this has more to do with the length of the windpipe. Other birds (especially non-passeriforms) sometimes have
songs A song is a musical composition performed by the human voice. The voice often carries the melody (a series of distinct and fixed pitches) using patterns of sound and silence. Songs have a structure, such as the common ABA form, and are usuall ...
to attract mates or hold territory, but these are usually simple and repetitive, lacking the variety of many oscine songs. The monotonous repetition of the
common cuckoo The cuckoo, common cuckoo, European cuckoo or Eurasian cuckoo (''Cuculus canorus'') is a member of the cuckoo order of birds, Cuculiformes, which includes the Geococcyx, roadrunners, the ani (bird), anis and the coucals. This species is a widesp ...
or little crake can be contrasted with the variety of a
nightingale The common nightingale, rufous nightingale or simply nightingale (''Luscinia megarhynchos''), is a small passerine bird which is best known for its powerful and beautiful song. It was formerly classed as a member of the thrush family Turdidae, ...
or marsh warbler. However, although many songbirds have songs that are pleasant to the human ear, this is not invariably the case. Many members of the
crow A crow is a bird of the genus ''Corvus'', or more broadly, a synonym for all of ''Corvus''. The word "crow" is used as part of the common name of many species. The related term "raven" is not linked scientifically to any certain trait but is rathe ...
family (
Corvidae Corvidae is a Cosmopolitan distribution, cosmopolitan Family (biology), family of Songbird, oscine passerine birds that contains the crows, ravens, Rook (bird), rooks, magpies, jackdaws, jays, treepies, choughs, and Nutcracker (bird), nutcrackers ...
) communicate with croaks or screeches, which sound harsh to humans. Even these, however, have a song of sorts, a softer twitter that is given between courting partners. And even though some
parrot Parrots (Psittaciformes), also known as psittacines (), are birds with a strong curved beak, upright stance, and clawed feet. They are classified in four families that contain roughly 410 species in 101 genus (biology), genera, found mostly in ...
s (which are not songbirds) can be taught to repeat human speech, vocal mimicry among birds is almost completely restricted to songbirds, some of which (such as the
lyrebird A lyrebird is either of two species of ground-dwelling Australian birds that compose the genus ''Menura'', and the family Menuridae. They are most notable for their impressive ability to mimic natural and artificial sounds from their environme ...
s or the aptly named
mockingbird Mockingbirds are a group of New World passerine birds from the family (biology), family Mimidae. They are best known for the habit of some species Mimicry, mimicking the songs of other birds and the sounds of insects and amphibians, often loudly ...
s) excel in imitating the sounds of other birds or even environmental noises. The birds from higher altitudes have evolved thicker downs (also known as jackets) to protect themselves from colder temperatures. Their feathers have outer and inner portions, with the lower down being fluffier and warmer to provide increased warmth.


Song repertoire and courtship

Sexual selection can be broken down into several different studies regarding different aspects of a bird's song. As a result, songs can vary even within a single species. Many believe that song repertoire and cognition have a direct relationship. However, a study published in 2013 has shown that cognitive abilities may not all be directly related to the song repertoire of a songbird. Specifically, spatial learning is said to have an inverse relationship with song repertoire. So for example, this would be an individual who does not migrate as far as others in the species but has a better song repertoire. This suggests an evolutionary trade-off between possible alleles. With natural selection choosing traits best fit for reproductive success, there could be a trade-off in either direction depending on which trait would produce a higher fitness at that time period. Song repertoire can be attributed to male songbirds as it is one of the main mechanisms of courtship. Song repertoires differ from male individual to male individual and species to species. Some species may typically have large repertoires while others may have significantly smaller ones.
Mate choice Mate choice is one of the primary mechanisms under which evolution can occur. It is characterized by a "selective response by animals to particular stimuli" which can be observed as behavior.Bateson, Paul Patrick Gordon. "Mate Choice." Mate Choi ...
in female songbirds is a significant realm of study as song abilities are continuously evolving. Males often sing to assert their dominance over other males in competition for a female, sometimes in lieu of a combative episode, and to arouse the female by announcing a readiness to mate. Though less frequent, females have also been known to sing occasionally a duet with a mate as an affirmation of their partnership. While some will sing their song from a familiar perch, other species common to grasslands will sing a familiar song each time they fly. Currently, there have been numerous studies involving songbird repertoires, unfortunately, there has not yet been a concrete evidence to confirm that every songbird species prefers larger repertoires. A conclusion can be made that it can vary between species on whether a larger repertoire is connected to better fitness. With this conclusion, it can be inferred that evolution via natural selection, or sexual selection, favors the ability to retain larger repertoires for these certain species as it leads to higher reproductive success. During times of courtship, it is said that male songbirds increase their repertoire by mimicking other species songs. The better the mimicking ability, retaining ability, and the quantity of other species mimicked has been proven to have a positive relationship with mating success. Female preferences cause the constant improvement of accuracy and presentation of the copied songs. Another theory known as the "song-sharing hypothesis" suggests that females prefer simpler, more homogenous songs that signal a male of familiar territory. As birdsong can be broken into regional dialects through this process of mimicry, the foreign song of a newcomer suggests the lack of territorial possession. This can be costly in the wake of territorial conflicts between disparate songbird populations and may compel a female to prefer a male spouting a familiar song of the area.


Taxonomy and systematics

Sibley and Alquist divided songbirds into two "
parvorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized ...
s",
Corvida The "Corvida" were one of two "parvorders" contained within the suborder Passeri, as proposed in the Sibley-Ahlquist taxonomy, the other being Passerida. Standard taxonomic practice would place them at the rank of infraorder. More recent res ...
and
Passerida Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorde ...
(standard
taxonomic 280px, Generalized scheme of taxonomy Taxonomy is a practice and science concerned with classification or categorization. Typically, there are two parts to it: the development of an underlying scheme of classes (a taxonomy) and the allocation ...
practice would rank these as
infraorder Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classific ...
s), distributed in Australo-Papua and
Eurasia Eurasia ( , ) is a continental area on Earth, comprising all of Europe and Asia. According to some geographers, Physical geography, physiographically, Eurasia is a single supercontinent. The concept of Europe and Asia as distinct continents d ...
respectively.Selvatti, A.P. ''et al''. (2015
A Paleogene origin for crown passerines and the diversification of the Oscines in the New World
''Molecular Phylogenetics and Evolution'', 88:1-15.
Subsequent molecular studies, however, show this treatment to be somewhat erroneous. Passerida is a highly diverse lineage, uniting over one-third of all bird species to include (in 2015) 3,885 species). These are divided into three major superfamilies (though not exactly corresponding to the Sibley-Ahlquist arrangement), in addition to some minor lineages. In contrast, Sibley & Alquist's "Corvida" is a phylogenetic grade and an artefact of the
phenetic In biology, phenetics (; ), also known as taximetrics, is an attempt to classify organisms based on overall similarity, usually with respect to Morphology (biology), morphology or other observable traits, regardless of their phylogeny or evoluti ...
methodology. The bulk of the "Corvida" make up the large clade Corvides (812 species as of 2015), which is a
sister group In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...
to the Passerida. The remaining 15 oscine families (343 species in 2015) form a series of basally branching sister groups to the Corvoid - Passerid clade. All of these groups, which form at least six successively branching basal clades, are found exclusively or predominantly in Australasia. Australian endemics are also prominent among basal lineages in both Corvoids and Passerids, suggesting that songbirds originated and diverged in Australia. Scrubbirds and lyrebirds, of which there are just two species of each, represent the oldest lineage of songbirds on Earth. The rufous scrubbird, ''Atrichornis rufescens'', is essentially confined to the Gondwana Rainforests of Australia World Heritage Area, occurring in both Queensland and New South Wales sections. It is now only found at elevations above . One of the earliest known fossil songbirds is '' Resoviaornis'' from the
Early Oligocene The Rupelian is, in the geologic timescale, the older of two age (geology), ages or the lower of two stage (stratigraphy), stages of the Oligocene epoch (geology), Epoch/series (stratigraphy), Series. It spans the time between . It is preceded b ...
of Poland.


Families

* Menuroidea ** Menuridae:
lyrebird A lyrebird is either of two species of ground-dwelling Australian birds that compose the genus ''Menura'', and the family Menuridae. They are most notable for their impressive ability to mimic natural and artificial sounds from their environme ...
s ** Atrichornithidae: scrub birds * Bowerbirds and Australian treecreepers **
Climacteridae There are seven species of Australasian treecreeper in the passerine bird family Climacteridae. They are medium-small, mostly brown birds with patterning on their underparts, and all are endemic to Australia-New Guinea. They resemble, but are n ...
: Australian treecreepers **
Ptilonorhynchidae Bowerbirds () make up the bird family (biology), family Ptilonorhynchidae. They are renowned for their unique courtship behaviour, where males build a structure and decorate it with sticks and brightly coloured objects in an attempt to attract a ...
:
bowerbird Bowerbirds () make up the bird family Ptilonorhynchidae. They are renowned for their unique courtship behaviour, where males build a structure and decorate it with sticks and brightly coloured objects in an attempt to attract a mate. The family ...
s * Meliphagoidea: honeyeaters and allies **
Maluridae The Australasian wrens are a family, Maluridae, of small, insectivorous passerine birds endemic to Australia and New Guinea. While commonly known as wrens, they are unrelated to the true wrens. The family comprises 32 species (including sixteen ...
:
fairy-wren The Australasian wrens are a family, Maluridae, of small, insectivorous passerine birds endemic to Australia and New Guinea. While commonly known as wrens, they are unrelated to the true wrens. The family comprises 32 species (including sixteen ...
s,
emu-wren The emu-wrens (''Stipiturus'') are a genus of passerine birds in the Australasian wren family, Maluridae. They are found only in Australia, where they inhabit scrubland, scrub, heathland and grassland. They are small birds, 12–19 cm long w ...
s and
grasswren Grasswrens (''Amytornis)'' are a genus of birds in the Australasian wren family, Maluridae. Taxonomy and systematics ''Amytornis'' is the only genus classified within the subfamily Amytornithinae, and form a separate clade to the related fair ...
s **
Meliphagidae The honeyeaters are a large and diverse family (biology), family, Meliphagidae, of small to medium-sized birds. The family includes the Epthianura, Australian chats, myzomelas, friarbirds, wattlebirds, Manorina, miners and melidectes. They are m ...
: true
honeyeater The honeyeaters are a large and diverse family, Meliphagidae, of small to medium-sized birds. The family includes the Australian chats, myzomelas, friarbirds, wattlebirds, miners and melidectes. They are most common in Australia and New Gui ...
s and chats ** Dasyornithidae: bristlebirds ** Pardalotidae: pardalotes **
Acanthizidae Acanthizidae—sometimes called Australian warblers—are a family of passerine birds which includes gerygones, thornbills '' Acanthiza'', and scrubwrens '' Sericornis''. The family Acanthizidae consists of small to medium passerine birds, with ...
: scrubwrens, thornbills, and gerygones * Australopapuan babblers ** Pomatostomidae: Australasian babblers * Logrunners ** Orthonychidae: logrunners * Other basal lineages ** Cnemophilidae: satinbirds '' Cnemophilus'' and '' Loboparadisea'' ** Melanocharitidae: berrypeckers and longbills **
Callaeidae Callaeidae (sometimes Callaeatidae) is a family of passerine birds endemic to New Zealand. It contains three genera, with five species in the family. One species, the huia, became extinct early in the 20th century, while the South Island kōk ...
: New Zealand wattlebirds kōkako, saddleback and † huia ** Notiomystidae: stitchbird * Corvides ** Paramythiidae: tit berrypeckers and crested berrypeckers ** Psophodidae:
whipbird Psophodidae is a family of passerine birds native to Australia and nearby areas. It has a complicated taxonomic history and different authors vary in which birds they include in the family. In the strictest sense, it includes only the five or six ...
s, jewel-babblers and quail-thrushes ** Platysteiridae: wattle-eyes and batis ** Malaconotidae: bush-shrikes ** Machaerirynchidae: boatbills **
Vangidae The family Vangidae (from ''vanga'', Malagasy language, Malagasy for the hook-billed vanga, ''Vanga curvirostris'') comprises a group of often shrike-like medium-sized birds distributed from Asia to Africa, including the vangas of Madagascar to ...
: vangas, woodshrikes, and helmetshrikes ** Pityriasidae: Bornean bristlehead **
Artamidae Artamidae is a family of passerine birds found in Australia, the Indo-Pacific region, and Southern Asia. It includes 24 extant species in six genera and three subfamilies: Peltopsinae (with one genus, '' Peltops''), Artaminae (with one genus con ...
:
butcherbird Butcherbirds are songbirds closely related to the Australian magpie. Most are found in the genus ''Cracticus'', but the black butcherbird is placed in the monotypic genus ''Melloria''. They are native to Australasia. Taxonomy Together with thr ...
s, currawongs and
Australian magpie The Australian magpie (''Gymnorhina tibicen'') is a black and white passerine bird native to Australia and southern New Guinea, and introduced to New Zealand, and the Fijian island of Taveuni. Although once considered to be three separate ...
(formerly in Cracticidae) ** Rhagologidae: mottled whistlers ** Aegithinidae:
iora The ioras are a small family, Aegithinidae, of four passerine bird species found in south and southeast Asia. The family is composed of a single genus, ''Aegithina''. They were formerly grouped with the leafbirds and fairy-bluebirds, in the fam ...
s ** Campephagidae:
cuckooshrike The cuckooshrikes and allies in the family (biology), family Campephagidae are small to medium-sized passerine bird species found in the subtropical and tropical Africa, Asia and Australasia. The 93 species are divided into 11 genus, genera. The ...
s and trillers ** Mohouidae: whiteheads ** Neosittidae:
sittella The sittellas are a family, Neosittidae, of small passerine birds found only in Australasia. They resemble nuthatches, but whilst they were considered to be in that family for many years they are now afforded their own family. They do not migrat ...
s ** Eulacestomidae: ploughbills ** Oreoicidae: Australo-Papuan bellbirds **
Pachycephalidae The Pachycephalidae are a family of bird species that includes the whistlers, shrikethrushes, and three of the pitohuis, and is part of the ancient Australo-Papuan radiation of songbirds. The family includes 69 species that are separated into fi ...
:
whistlers Whistler may refer to: * Someone who whistles Places Canada * Whistler, British Columbia, a resort town ** Whistler railway station ** Whistler Secondary School * Whistler Blackcomb, a ski resort in British Columbia * Whistler Mountain, Bri ...
, shrike-thrushes,
pitohui The pitohuis are bird species endemic to New Guinea. The onomatopoeic name is thought to be derived from that used by New Guineans from nearby Dorey (Manokwari), but it is also used as the name of a genus '' Pitohui'' which was established by th ...
s and allies ** Laniidae:
shrike Shrikes () are passerine birds of the family Laniidae. The family is composed of 34 species in two genera. The family name, and that of the larger genus, '' Lanius'', is derived from the Latin word for "butcher", and some shrikes are also known ...
s **
Vireonidae The vireos make up a family (biology), family, Vireonidae, of small to medium-sized passerine birds found in the New World (Canada to Argentina, including Bermuda and the West Indies) and Southeast Asia. The family contains 62 species and is d ...
: vireos ** Oriolidae: orioles, figbirds and † piopio (formerly Turnagridae) ** Dicruridae: drongos **
Rhipiduridae The family Rhipiduridae are small insectivorous birds of Australasia, Southeast Asia and the Indian subcontinent that includes the fantails and silktails. Taxonomy and systematics The family contains 55 species which are divided into four genera ...
:
fantail Fantails are small insectivorous songbirds of the genus ''Rhipidura'' in the family Rhipiduridae, native to Australasia, Southeast Asia and the Indian subcontinent. Most of the species are about long, specialist aerial feeders, and named as "f ...
s **
Monarchidae The monarchs or monarch flycatchers (family Monarchidae) comprise a family of over 100 passerine birds which includes shrikebills, paradise flycatchers, and Grallina, magpie-larks. Monarchids are small insectivore, insectivorous songbirds with l ...
:
monarch A monarch () is a head of stateWebster's II New College Dictionary. "Monarch". Houghton Mifflin. Boston. 2001. p. 707. Life tenure, for life or until abdication, and therefore the head of state of a monarchy. A monarch may exercise the highest ...
s and allies ** Platylophidae: jayshrikes **
Corvidae Corvidae is a Cosmopolitan distribution, cosmopolitan Family (biology), family of Songbird, oscine passerine birds that contains the crows, ravens, Rook (bird), rooks, magpies, jackdaws, jays, treepies, choughs, and Nutcracker (bird), nutcrackers ...
:
crow A crow is a bird of the genus ''Corvus'', or more broadly, a synonym for all of ''Corvus''. The word "crow" is used as part of the common name of many species. The related term "raven" is not linked scientifically to any certain trait but is rathe ...
s,
magpie Magpies are birds of various species of the family Corvidae. Like other members of their family, they are widely considered to be intelligent creatures. The Eurasian magpie, for instance, is thought to rank among the world's most intelligent c ...
s, and jays **
Corcoracidae Corcoracidae is a family of passerine birds known as the Australian mudnesters. The family has sometimes been called Struthideidae in the past; however, despite Struthideidae being an older name than Corcoracidae, the latter name takes precedence ...
:
white-winged chough The white-winged chough (''Corcorax melanorhamphos'') is one of only two surviving species of the Australian mud-nest builders family, Corcoracidae, and is the Monotypic taxon, only member of the genus ''Corcorax''. It is native to southern and e ...
and
apostlebird The apostlebird (''Struthidea cinerea''), also known as the grey jumper, lousy jack or happy family, is a quick-moving, gray or black bird about 33 cm (13 in) long. It is a native to Australia where it roams woodlands, eating insects an ...
** Melampittidae: melampittas ** Ifritidae: ifritabirds ** Paradisaeidae: birds of paradise *
Passerida Passerida is, under the Sibley-Ahlquist taxonomy, one of two parvorders contained within the suborder Passeri (standard taxonomic practice would place them at the rank of infraorder). While more recent research suggests that its sister parvorde ...
** Petroicidae: Australasian robins ** Picathartidae:
rockfowl The picathartes, rockfowl, or bald crows are a small genus of two passerine bird species forming the family Picathartidae found in the rain-forests of tropical west and central Africa. They have unfeathered heads, and feed on insects and inverteb ...
** Chaetopidae: rockjumpers ** Eupetidae: rail-babbler **
Bombycillidae The waxwings are three species of passerine birds classified in the genus ''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestripe ...
:
waxwing The waxwings are three species of passerine birds classified in the genus ''Bombycilla''. They are pinkish-brown and pale grey with distinctive smooth plumage in which many body feathers are not individually visible, a black and white eyestripe, ...
s and allies ** Ptiliogonatidae: silky-flycatchers **
Hypocoliidae The grey hypocolius or simply hypocolius (''Hypocolius ampelinus'') is a small passerine bird species. It is the sole member of the genus ''Hypocolius'' and it is placed in a family of its own, the Hypocoliidae. This slender and long tailed bird ...
: hypocolius ** Dulidae: palmchat ** †
Mohoidae Mohoidae, also known as the Hawaiian honeyeaters, was a Family (biology), family of Hawaiian species of now Holocene extinction event, recently Extinction, extinct, Nectarivore, nectarivorous songbirds in the genera ''Moho (genus), Moho'' (ōō) ...
: some Hawaiian honeyeaters, '' Moho'' and '' Chaetoptila'' genera, not closely related to
Meliphagidae The honeyeaters are a large and diverse family (biology), family, Meliphagidae, of small to medium-sized birds. The family includes the Epthianura, Australian chats, myzomelas, friarbirds, wattlebirds, Manorina, miners and melidectes. They are m ...
** Hylocitreidae: hylocitrea ** Stenostiridae: fairy-flycatcher and allies **
Paridae The tits, chickadees, and titmice constitute the Paridae, a family of small passerine birds which occur mainly in the Northern Hemisphere and Africa. Many were formerly classified in the genus ''Parus''. Eurasian and African members of this f ...
: tits,
chickadee The chickadees are a group of North American birds in the family Paridae included in the genus '' Poecile''. Species found in North America are referred to as chickadees; species found elsewhere in the world are called tits. They are small-si ...
s, and
titmouse ''Baeolophus'' is a genus of birds in the family Paridae, commonly called tits. Its members are known as titmouses or titmice. All the species are native to North America. In the past, most authorities retained ''Baeolophus'' as a subgenus w ...
** Remizidae: penduline-tits ** Nicatoridae: nicators ** Panuridae: bearded reedling **
Alaudidae Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
:
lark Larks are passerine birds of the family Alaudidae. Larks have a cosmopolitan distribution with the largest number of species occurring in Africa. Only a single species, the horned lark, occurs in North America, and only Horsfield's bush lark occ ...
s **
Pycnonotidae The bulbuls are members of a family (biology), family, Pycnonotidae, of medium-sized perching bird, passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and in ...
:
bulbul The bulbuls are members of a family, Pycnonotidae, of medium-sized passerine songbirds, which also includes greenbuls, brownbuls, leafloves, and bristlebills. The family is distributed across most of Africa and into the Middle East, tropic ...
s ** Hirundinidae: swallows and Martin (bird), martins ** Pnoepygidae: Pnoepyga, wren-babblers ** Macrosphenidae: crombecs and African warblers ** Cettiidae: bush-warblers and allies ** Scotocercidae: streaked scrub-warbler ** Erythrocercidae: yellow flycatchers ** Aegithalidae: long-tailed tits ** Phylloscopidae: leaf-warblers and allies. Recently split from Sylviidae. ** Acrocephalidae: reed warblers and allies ** Locustellidae: Locustellidae, grassbirds and allies ** Donacobiidae: black-capped donacobius ** Bernieridae: Malagasy warblers ** Cisticolidae: cisticolas and allies ** Timaliidae: babblers ** Pellorneidae: ground babblers ** Leiothrichidae: laughingthrushes and allies ** Sylviidae: Old World warblers ** White-eye, Zosteropidae: white-eyes ** Arcanatoridae: dapple-throat and allies ** Promeropidae: sugarbirds ** Irenidae: fairy-bluebirds ** Regulidae: kinglets ** Elachuridae: elachuras ** Hyliotidae: hyliotas ** Troglodytidae: wrens ** Polioptilidae: gnatcatchers ** Sittidae: nuthatches ** Tichodromidae: wallcreeper ** Certhiidae: treecreepers ** Mimidae:
mockingbird Mockingbirds are a group of New World passerine birds from the family (biology), family Mimidae. They are best known for the habit of some species Mimicry, mimicking the songs of other birds and the sounds of insects and amphibians, often loudly ...
s and thrashers ** Sturnidae: starlings ** Buphagidae: oxpeckers ** Turdidae: Thrush (bird), thrushes and allies ** Muscicapidae: Old World flycatchers and Chat (bird), chats ** Cinclidae: dippers ** Chloropseidae: leafbirds ** Dicaeidae: flowerpeckers ** Nectariniidae: sunbirds ** Old World sparrow, Passeridae: true Old World sparrow, sparrows ** Ploceidae: weavers and widowbirds ** Estrildid finch, Estrildidae: estrildid finches (waxbills, munias, etc.) ** Viduidae: indigo birds and Viduidae, whydahs ** Peucedramidae: olive warbler ** Prunellidae: accentor ** Motacillidae: wagtails and pipits ** Urocynchramidae: Przevalski's finch ** Fringillidae: true finches and Hawaiian honeycreepers (formerly Drepanididae) ** Parulidae: New World warblers, for example the black-throated blue warblers and allies ** Icteridae: American icterid, blackbirds, New World orioles, grackles and cowbirds. ** Coerebidae: bananaquit ** Emberizidae: Emberiza, buntings **Passerellidae: New World sparrows ** Thraupidae: tanagers, true honeycreepers and allies ** Calcariidae: snow buntings and longspurs ** Cardinalidae: cardinals and allies


See also

* Song system


References


External links


Oscines
Tree of Life web project article July 31, 2006. {{Authority control Passeri, Songbirds, Extant Eocene first appearances