Gorillas are primarily
herbivorous
A herbivore is an animal anatomically and physiologically evolved to feed on plants, especially upon vascular tissues such as foliage, fruits or seeds, as the main component of its diet. These more broadly also encompass animals that eat n ...
,
terrestrial
Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
great ape
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s that inhabit the tropical forests of
equatorial Africa
Equatorial Africa is an ambiguous term that sometimes is used to refer to the equatorial region of sub-Saharan Africa traversed by the Equator, more broadly to tropical Africa, or in a biological and geo-environmental sense to the intra-tropic ...
. The
genus
Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
''Gorilla'' is divided into two species: the
eastern gorilla
The eastern gorilla (''Gorilla beringei'') is a critically endangered species of the genus ''Gorilla'' and the largest living primate. At present, the species is subdivided into two subspecies. There are 6,800 eastern lowland gorillas or Grauer ...
and the
western gorilla
The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the species is found in a region of midwest Africa, geographically iso ...
, and either four or five
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
. The
DNA
Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...
of gorillas is highly similar to
that of humans, from 96 to 99% depending on what is included, and they are the next closest living relatives to
human
Humans (''Homo sapiens'') or modern humans are the most common and widespread species of primate, and the last surviving species of the genus ''Homo''. They are Hominidae, great apes characterized by their Prehistory of nakedness and clothing ...
s after the
chimpanzees
The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close relative the ...
.
Gorillas are the largest
living
Living or The Living may refer to:
Common meanings
*Life, a condition that distinguishes organisms from inorganic objects and dead organisms
** Living species, one that is not extinct
*Personal life, the course of an individual human's life
* ...
primate
Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...
s, reaching heights between , weights between , and arm spans up to , depending on species and sex. They tend to live in troops, with the leader being called a silverback. The eastern gorilla is distinguished from the western by darker fur colour and some other minor morphological differences. Gorillas tend to live 35–40 years in the wild.
Gorillas' natural habitats cover
tropical or subtropical forest in
Sub-Saharan Africa
Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...
. Although their range covers a small percentage of Sub-Saharan Africa, gorillas cover a wide range of elevations. The
mountain gorilla
The mountain gorilla (''Gorilla beringei beringei'') is one of the two subspecies of the eastern gorilla. It is listed as endangered by the IUCN .
There are two populations: One is found in the Virunga Mountains, Virunga volcanic mountains of C ...
inhabits the
Albertine Rift
The Albertine Rift is the western branch of the East African Rift, covering parts of Uganda, the Democratic Republic of the Congo (DRC), Rwanda, Burundi and Tanzania.
It extends from the northern end of Lake Albert to the southern end of Lake Tan ...
montane
cloud forest
A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest, is a generally tropical or subtropical, evergreen, Montane forest, montane, Tropical and subtropical moist broadleaf forests, moist forest characteri ...
s of the
Virunga Volcanoes
The Virunga Mountains (also known as Mufumbiro) are a chain of volcanoes in East Africa, in the area where Rwanda, the Democratic Republic of the Congo (DRC), and Uganda meet. The mountain range is a branch of the Albertine Rift Mountains, ...
, ranging in altitude from . Lowland gorillas live in dense forests and lowland swamps and marshes as low as
sea level
Mean sea level (MSL, often shortened to sea level) is an mean, average surface level of one or more among Earth's coastal Body of water, bodies of water from which heights such as elevation may be measured. The global MSL is a type of vertical ...
, with
western lowland gorilla
The western lowland gorilla (''Gorilla gorilla gorilla'') is one of two Critically Endangered subspecies of the western gorilla (''Gorilla gorilla'') that lives in Montane ecosystems#Montane forests, montane, Old-growth forest, primary and sec ...
s living in Central West African countries and
eastern lowland gorilla
The eastern lowland gorilla (''Gorilla beringei graueri'') or Grauer's gorilla is a Critically Endangered subspecies of eastern gorilla endemic to the mountainous forests of eastern Democratic Republic of the Congo. Important populations of th ...
s living in the
Democratic Republic of the Congo
The Democratic Republic of the Congo (DRC), also known as the DR Congo, Congo-Kinshasa, or simply the Congo (the last ambiguously also referring to the neighbouring Republic of the Congo), is a country in Central Africa. By land area, it is t ...
near its border with
Rwanda
Rwanda, officially the Republic of Rwanda, is a landlocked country in the Great Rift Valley of East Africa, where the African Great Lakes region and Southeast Africa converge. Located a few degrees south of the Equator, Rwanda is bordered by ...
.
There are thought to be around 316,000 western gorillas in the wild, and 5,000 eastern gorillas. Both species are classified as
Critically Endangered
An IUCN Red List critically endangered (CR or sometimes CE) species is one that has been categorized by the International Union for Conservation of Nature as facing an extremely high risk of extinction in the wild. As of December 2023, of t ...
by the IUCN; all subspecies are classified as Critically Endangered with the exception of the mountain gorilla, which is classified as
Endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
. There are many threats to their survival, such as poaching, habitat destruction, and disease, which threaten the survival of the species. However, conservation efforts have been successful in some areas where they live.
History and etymology
The word ''gorilla'' comes from the history of
Hanno the Navigator
Hanno the Navigator (sometimes "Hannon"; , ; ) was a Ancient Carthage, Carthaginian explorer (sometimes identified as a king) who lived during the 5th century BC, fifth century BC, known for his Navy, naval expedition along the coast of West A ...
( 500 BC), a
Carthaginian The term Carthaginian ( ) usually refers to the civilisation of ancient Carthage.
It may also refer to:
* Punic people, the Semitic-speaking people of Carthage
* Punic language
The Punic language, also called Phoenicio-Punic or Carthaginian, i ...
explorer on an expedition to the west
Africa
Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...
n coast to the area that later became
Sierra Leone
Sierra Leone, officially the Republic of Sierra Leone, is a country on the southwest coast of West Africa. It is bordered to the southeast by Liberia and by Guinea to the north. Sierra Leone's land area is . It has a tropical climate and envi ...
. Members of the expedition encountered "savage people, the greater part of whom were women, whose bodies were hairy, and whom our interpreters called Gorillae". It is unknown whether what the explorers encountered were what we now call gorillas, another species of ape or monkeys, or humans.
[ Skins of gorillai women, brought back by Hanno, are reputed to have been kept at Carthage until Rome destroyed the city 350 years later at the end of the ]Punic Wars
The Punic Wars were a series of wars fought between the Roman Republic and the Ancient Carthage, Carthaginian Empire during the period 264 to 146BC. Three such wars took place, involving a total of forty-three years of warfare on both land and ...
, 146 BC.
In 1625 Andrew Battel mentioned the existence of the animal, under the name Pongo:
A century and a half after Battel's story was published, one writer called Radermacher wrote that "the large species, described by Buffon and other authors as of the size of a man, is held by many to be a Chimera
Chimera, Chimaera, or Chimaira (Greek for " she-goat") originally referred to:
* Chimera (mythology), a fire-breathing monster of ancient Lycia said to combine parts from multiple animals
* Mount Chimaera, a fire-spewing region of Lycia or Cilicia ...
."
The American physician and missionary Thomas Staughton Savage and naturalist Jeffries Wyman first described the western gorilla
The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the species is found in a region of midwest Africa, geographically iso ...
in 1847 from specimens obtained in Liberia
Liberia, officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to Guinea–Liberia border, its north, Ivory Coast to Ivory Coast–Lib ...
. They called it ''Troglodytes gorilla'', using the then-current name of the chimpanzee genus. The species name was derived , as described by Hanno.
Evolution and classification
The closest relatives of gorillas are the other two Homininae
Homininae (the hominines) is a subfamily of the family Hominidae (hominids). (The Homininae——encompass humans, and are also called "African hominids" or "African apes".) This subfamily includes two tribes, Hominini and Gorillini, both having ...
genera, chimpanzees and humans, all of them having diverged from a common ancestor about 7 million years ago. Human gene sequences differ only 1.6% on average from the sequences of corresponding gorilla genes, but there is further difference in how many copies each gene has.
Until recently, gorillas were considered to be a single species, with three subspecies: the western lowland gorilla, the eastern lowland gorilla and the mountain gorilla. There is now agreement that there are two species, each with two subspecies. More recently, a third subspecies has been claimed to exist in one of the species. The separate species and subspecies developed from a single type of gorilla during the Ice Age
An ice age is a long period of reduction in the temperature of Earth's surface and atmosphere, resulting in the presence or expansion of continental and polar ice sheets and alpine glaciers. Earth's climate alternates between ice ages, and g ...
, when their forest habitats shrank and became isolated from each other.[ ] Primatologists
Primatology is the scientific study of non-human primates. It is a diverse discipline at the boundary between mammalogy and anthropology, and researchers can be found in academic departments of anatomy, anthropology, biology, medicine, psycholo ...
continue to explore the relationships between various gorilla populations. The species and subspecies listed here are the ones upon which most scientists agree.
The proposed third subspecies of ''Gorilla beringei'', which has not yet received a trinomen
In biology, trinomial nomenclature is the system of names for taxa below the rank of species. These names have three parts. The usage is different in zoology and botany.
In zoology
In zoological nomenclature, a trinomen (), trinominal name, or ...
, is the Bwindi population of the mountain gorilla, sometimes called the Bwindi gorilla.
Some variations that distinguish the classifications of gorilla include varying density, size, hair colour, length, culture, and facial widths. Population genetics of the lowland gorillas suggest that the western and eastern lowland populations diverged around 261 thousand years ago.
Characteristics
Wild male gorillas weigh , while adult females weigh . Adult males are tall, with an arm span that stretches from . Female gorillas are shorter at , with smaller arm spans. Colin Groves
Colin Peter Groves (24 June 1942 – 30 November 2017) was a British-Australian biologist and anthropologist. Groves was professor of biological anthropology at the Australian National University in Canberra, Australia.
Education
Born in Englan ...
(1970) calculated the average weight of 42 wild adult male gorillas at 144 kg, while Smith and Jungers (1997) found the average weight of 19 wild adult male gorillas to be 169 kg. Adult male gorillas are known as silverbacks due to the characteristic silver hair on their backs reaching to the hips. The tallest wild gorilla recorded was a silverback with an arm span of , a chest of , and a weight of , shot in Alimbongo, northern Kivu
Kivu is the name for a large region in the eastern Democratic Republic of the Congo that borders Lake Kivu. It was a ''Région'' (read 'province') of the country under the rule of Mobutu Sese Seko from 1966 to 1988. As an official ''Région'' ...
in May 1938. The heaviest wild gorilla recorded was a silverback shot in Ambam
Ambam is a town and commune in the Vallée-du-Ntem department, South Province, Cameroon on the border with Equatorial Guinea and Gabon. This frontier town is located approximately 220 km from Yaoundé and as of 2005 had a population of 41 ...
, Cameroon
Cameroon, officially the Republic of Cameroon, is a country in Central Africa. It shares boundaries with Nigeria to the west and north, Chad to the northeast, the Central African Republic to the east, and Equatorial Guinea, Gabon, and the R ...
, which weighed . The tallest gorilla in captivity was Gust, a western lowland gorilla that was captured as a baby in Belgian Congo
The Belgian Congo (, ; ) was a Belgian colonial empire, Belgian colony in Central Africa from 1908 until independence in 1960 and became the Republic of the Congo (Léopoldville). The former colony adopted its present name, the Democratic Repu ...
and spent his life at Antwerp Zoo
Antwerp Zoo () is a zoo in the centre of Antwerp, Belgium, located next to the Antwerpen-Centraal railway station. It is the oldest animal park in the country, and one of the oldest in the world, established on 21 July 1843.
History
Since its ...
. He was tall. Males in captivity can be overweight
Being overweight is having more body fat than is optimally healthy. Being overweight is especially common where food supplies are plentiful and lifestyles are sedentary.
, excess weight reached epidemic proportions globally, with more than ...
and reach weights up to .
The eastern gorilla is more darkly coloured than the western gorilla, with the mountain gorilla being the darkest of all. The mountain gorilla also has the thickest hair. The western lowland gorilla can be brown or greyish with a reddish forehead. In addition, gorillas that live in lowland forest are more slender and agile than the more bulky mountain gorillas. The eastern gorilla also has a longer face and broader chest than the western gorilla. Like humans, gorillas have individual fingerprints.
Their eye colour is dark brown, framed by a black ring around the iris. Gorilla facial structure is described as mandibular prognathism
Prognathism is a positional relationship of the Human mandible, mandible or maxilla to the skeletal base where either of the jaws protrudes beyond a predetermined imaginary line in the coronal plane of the skull.
In the case of ''mandibular'' ...
, that is, the mandible
In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla).
The jawbone i ...
protrudes farther out than the maxilla
In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
. Adult males also have a prominent sagittal crest
A sagittal crest is a ridge of bone running lengthwise along the midline of the top of the skull (at the sagittal suture) of many mammalian and reptilian skulls, among others. The presence of this ridge of bone indicates that there are excepti ...
.
Gorillas move around by knuckle-walking
Knuckle-walking is a form of quadrupedal walking in which the forelimbs hold the fingers in a partially flexed posture that allows body weight to press down on the ground through the knuckles. Gorillas and chimpanzees use this style of locomoti ...
, although they sometimes walk upright for short distances, typically while carrying food or in defensive situations. A 2018 study investigating the hand posture of 77 mountain gorillas at Bwindi Impenetrable National Park
The Bwindi Impenetrable National Park is a national park in southwestern Uganda. It is part of the Bwindi Impenetrable Forest and is situated along the Democratic Republic of the Congo border next to the Virunga National Park and on the edge of t ...
(8% of the population) found that knuckle walking was done only 60% of the time, and they also supported their weight on their fists, the backs of their hands/feet, and on their palms/soles (with the digits flexed). Such a range of hand postures was previously thought to have been used by only orangutans. Studies of gorilla handedness
In human biology, handedness is an individual's preferential use of one hand, known as the dominant hand, due to and causing it to be stronger, faster or more Fine motor skill, dextrous. The other hand, comparatively often the weaker, less dext ...
have yielded varying results, with some arguing for no preference for either hand, and others right-hand dominance for the general population.
Studies have shown gorilla blood is not reactive to anti-A and anti-B monoclonal antibodies
A monoclonal antibody (mAb, more rarely called moAb) is an antibody produced from a Lineage (evolution), cell lineage made by cloning a unique white blood cell. All subsequent antibodies derived this way trace back to a unique parent cell.
Mon ...
, which would, in humans, indicate type O blood
The ABO blood group system is used to denote the presence of one, both, or neither of the A and B antigens on erythrocytes (red blood cells). For human blood transfusions, it is the most important of the 47 different blood type (or group) cla ...
. Due to novel sequences, though, it is different enough to not conform with the human ABO blood group system
The ABO blood group system is used to denote the presence of one, both, or neither of the A and B antigens on erythrocytes (red blood cells). For human blood transfusions, it is the most important of the 47 different blood type (or group) c ...
, into which the other great apes fit.
A gorilla's lifespan is normally between 35 and 40 years, although zoo gorillas may live for 50 years or more in rare circumstances. At , Fatou is both the oldest living gorilla and oldest living female gorilla. The oldest male gorilla ever was Ozoum, who reached the final age of 61 years, 24 days. The oldest living male gorilla is Guhonda, aged .
Distribution and habitat
Gorillas have a patchy distribution. The range of the two species is separated by the Congo River
The Congo River, formerly also known as the Zaire River, is the second-longest river in Africa, shorter only than the Nile, as well as the third-largest river in the world list of rivers by discharge, by discharge volume, following the Amazon Ri ...
and its tributaries. The western gorilla lives in west central Africa, while the eastern gorilla lives in east central Africa. Between the species, and even within the species, gorillas live in a variety of habitats and elevations. Gorilla habitat ranges from montane forest
Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
to swampland. Eastern gorillas inhabit montane and submontane forests between above sea level.
Mountain gorillas live in montane forests at the higher end of the elevation range, while eastern lowland gorillas live in submontane forests at the lower end. In addition, eastern lowland gorillas live in montane bamboo forests, as well as lowland forests ranging from in elevation. Western gorillas live in both lowland swamp forests and montane forests, at elevations ranging from sea level to . Western lowland gorillas live in swamp and lowland forests ranging up to , and Cross River gorillas live in low-lying and submontane forests ranging from .
Ecology
Diet and foraging
A gorilla's day is divided between rest periods and travel or feeding periods. Diets differ between and within species. Mountain gorillas mostly eat foliage, such as leaves, stems, pith, and shoots, while fruit makes up a very small part of their diets. Mountain gorilla food is widely distributed and neither individuals nor groups have to compete with one another. Their home ranges vary from , and their movements range around or less on an average day. Despite eating a few species in each habitat, mountain gorillas have flexible diets and can live in a variety of habitats.
Eastern lowland gorillas have more diverse diets, which vary seasonally. Leaves and pith are commonly eaten, but fruits can make up as much as 25% of their diets. Since fruit is less available, lowland gorillas must travel farther each day, and their home ranges vary from , with day ranges . Eastern lowland gorillas will also eat insects, preferably ants. Western lowland gorillas depend on fruits more than the others and they are more dispersed across their range. They travel even farther than the other gorilla subspecies, at per day on average, and have larger home ranges of . Western lowland gorillas have less access to terrestrial herbs, although they can access aquatic herbs in some areas. Termites and ants are also eaten.
Gorillas rarely drink water "because they consume succulent vegetation that is almost half water as well as morning dew", although both mountain and lowland gorillas have been observed drinking.
Nesting
Gorillas construct nest
A nest is a structure built for certain animals to hold Egg (biology), eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of ...
s for daytime and night use. Nests tend to be simple aggregations of branches and leaves about in diameter and are constructed by individuals. Gorillas, unlike chimpanzees or orangutans, tend to sleep in nests on the ground. The young nest with their mothers, but construct nests after three years of age, initially close to those of their mothers. Gorilla nests are distributed arbitrarily and use of tree species for site and construction appears to be opportunistic. Nest-building by great apes is now considered to be not just animal architecture
Structures built by non-human animals, often called animal architecture, are common in many species. Examples of animal structures include termite mounds, ant hills, wasp and beehives, burrow complexes, beaver dams, elaborate nests of birds, an ...
, but as an important instance of tool use
Tool use by non-humans is a phenomenon in which a non-human animal uses any kind of tool in order to achieve a goal such as acquiring food and water, grooming, combat, defence, communication, recreation or construction. Originally thought to b ...
.
Gorillas make a new nest to sleep on each day; even if remaining in the same place, they do not use the previous one. Usually, they are made an hour before dusk, to be ready to sleep when night falls. Gorillas sleep longer than humans, an average of 12 hours per day.
Interspecies interactions
One possible predator of gorillas is the leopard
The leopard (''Panthera pardus'') is one of the five extant cat species in the genus ''Panthera''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a ...
. Gorilla remains have been found in leopard scat, but this may be the result of scavenging. When the group is attacked by humans, leopards, or other gorillas, an individual silverback will protect the group, even at the cost of his own life. Gorillas do not appear to directly compete with chimpanzee
The chimpanzee (; ''Pan troglodytes''), also simply known as the chimp, is a species of Hominidae, great ape native to the forests and savannahs of tropical Africa. It has four confirmed subspecies and a fifth proposed one. When its close rel ...
s in areas where they overlap. When fruit is abundant, gorilla and chimpanzee diets converge, but when fruit is scarce gorillas resort to vegetation. The two apes may also feed on different species, whether fruit or insects. Gorillas and chimpanzees may ignore or avoid each other when feeding on the same tree, but they have also been documented to form social bonds. Conversely, coalitions of chimpanzees have been observed attacking families of gorillas including silverbacks and killing infants.
Behaviour
Social structure
Gorillas live in groups called troops. Troops tend to be made of one adult male or silverback, with a harem of multiple adult females and their offspring. However, multiple-male troops also exist. A silverback is typically more than 12 years of age, and is named for the distinctive patch of silver hair on his back, which comes with maturity. Silverbacks have large canine teeth that also come with maturity. Both males and females tend to emigrate from their natal groups. For mountain gorillas, females disperse from their natal troops more than males. Mountain gorillas and western lowland gorillas also commonly transfer to second new groups.
Mature males also tend to leave their groups and establish their own troops by attracting emigrating females. However, male mountain gorillas sometimes stay in their natal troops and become subordinate to the silverback. If the silverback dies, these males may be able to become dominant or mate with the females. This behaviour has not been observed in eastern lowland gorillas. In a single male group, when the silverback dies, the females and their offspring disperse and find a new troop. Without a silverback to protect them, the infants will likely fall victim to infanticide
Infanticide (or infant homicide) is the intentional killing of infants or offspring. Infanticide was a widespread practice throughout human history that was mainly used to dispose of unwanted children, its main purpose being the prevention of re ...
. Joining a new group is likely to be a tactic against this. However, while gorilla troops usually disband after the silverback dies, female eastern lowlands gorillas and their offspring have been recorded staying together until a new silverback transfers into the group. This likely serves as protection from leopards.
The silverback is the centre of the troop's attention, making all the decisions, mediating conflicts, determining the movements of the group, leading the others to feeding sites, and taking responsibility for the safety and well-being of the troop. Younger males subordinate to the silverback, known as blackbacks, may serve as backup protection. Blackbacks are aged between 8 and 12 years and lack the silver back hair. The bond that a silverback has with his females forms the core of gorilla social life. Bonds between them are maintained by grooming and staying close together. Females form strong relationships with males to gain mating opportunities and protection from predators and infanticidal outside males. However, aggressive behaviours between males and females do occur, but rarely lead to serious injury. Relationships between females may vary. Maternally related females in a troop tend to be friendly towards each other and associate closely. Otherwise, females have few friendly encounters and commonly act aggressively towards each other.
Females may fight for social access to males and a male may intervene. Male gorillas have weak social bonds, particularly in multiple-male groups with apparent dominance hierarchies and strong competition for mates. Males in all-male groups, though, tend to have friendly interactions and socialise through play, grooming, and staying together, and occasionally they even engage in homosexual interactions. Severe aggression is rare in stable groups, but when two mountain gorilla groups meet the two silverbacks can sometimes engage in a fight to the death, using their canines to cause deep, gaping injuries.
Reproduction and parenting
Females mature at 10–12 years (earlier in captivity), and males at 11–13 years. A female's first ovulatory cycle occurs when she is six years of age, and is followed by a two-year period of adolescent infertility. The estrous cycle lasts 30–33 days, with outward ovulation signs subtle compared to those of chimpanzees. The gestation period lasts 8.5 months. Female mountain gorillas first give birth at 10 years of age and have four-year interbirth intervals. Males can be fertile before reaching adulthood. Gorillas mate year round.
Females will purse their lips and slowly approach a male while making eye contact. This serves to urge the male to mount her. If the male does not respond, then she will try to attract his attention by reaching towards him or slapping the ground. In multiple-male groups, solicitation indicates female preference, but females can be forced to mate with multiple males. Males incite copulation by approaching a female and displaying at her or touching her and giving a "train grunt". Recently, gorillas have been observed engaging in face-to-face sex, a trait once considered unique to humans and bonobo
The bonobo (; ''Pan paniscus''), also historically called the pygmy chimpanzee (less often the dwarf chimpanzee or gracile chimpanzee), is an endangered great ape and one of the two species making up the genus ''Pan (genus), Pan'' (the other bei ...
s.
Gorilla infants are vulnerable and dependent, thus mothers, their primary caregivers, are important to their survival. Male gorillas are not active in caring for the young, but they do play a role in socialising them to other youngsters. The silverback has a largely supportive relationship with the infants in his troop and shields them from aggression within the group. Infants remain in contact with their mothers for the first five months and mothers stay near the silverback for protection. Infants suck at least once per hour and sleep with their mothers in the same nest.
Infants begin to break contact with their mothers after five months, but only for a brief period each time. By 12 months old, infants move up to from their mothers. At around 18–21 months, the distance between mother and offspring increases and they regularly spend time away from each other. In addition, nursing decreases to once every two hours. Infants spend only half of their time with their mothers by 30 months. They enter their juvenile period at their third year, and this lasts until their sixth year. At this time, gorillas are weaned and they sleep in a separate nest from their mothers. After their offspring are weaned, females begin to ovulate and soon become pregnant again. The presence of play partners, including the silverback, minimizes conflicts in weaning between mother and offspring.
Communication
Twenty-five distinct vocalisations are recognised, many of which are used primarily for group communication within dense vegetation. Sounds classified as grunts and barks are heard most frequently while traveling, and indicate the whereabouts of individual group members. They may also be used during social interactions when discipline is required. Screams and roar
A roar is a type of animal vocalization that is loud, deep and resonating. Many mammals have evolved to produce roars and other roar-like vocals for purposes such as long-distance communication and intimidation. These include various species of ...
s signal alarm or warning, and are produced most often by silverbacks. Deep, rumbling belches suggest contentment and are heard frequently during feeding and resting periods. They are the most common form of intragroup communication.
For this reason, conflicts are most often resolved by displays and other threat behaviours that are intended to intimidate without becoming physical. As a result, fights do not occur very frequently. The ritual
A ritual is a repeated, structured sequence of actions or behaviors that alters the internal or external state of an individual, group, or environment, regardless of conscious understanding, emotional context, or symbolic meaning. Traditionally ...
ized charge display is unique to gorillas. The entire sequence has nine steps: (1) progressively quickening hooting, (2) symbolic feeding, (3) rising bipedally, (4) throwing vegetation, (5) chest-beating with cupped hands, (6) one leg kick, (7) sideways running, two-legged to four-legged, (8) slapping and tearing vegetation, and (9) thumping the ground with palms to end display.
A gorilla's chest-beat may vary in frequency depending on its size. Smaller ones tend to have higher frequencies, while larger ones tend to be lower. They also do it the most when females are ready to mate.
Intelligence
Gorillas are considered highly intelligent. A few individuals in captivity, such as Koko, have been taught a subset of sign language
Sign languages (also known as signed languages) are languages that use the visual-manual modality to convey meaning, instead of spoken words. Sign languages are expressed through manual articulation in combination with #Non-manual elements, no ...
. Like the other great ape
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s, gorillas can laugh, grieve, have "rich emotional lives", develop strong family bonds, make and use tools, and think about the past and future. Some researchers believe gorillas have spiritual feelings or religious sentiments. They have been shown to have cultures in different areas revolving around different methods of food preparation, and will show individual colour preferences.
Tool use
The following observations were made by a team led by Thomas Breuer of the Wildlife Conservation Society
The Wildlife Conservation Society (WCS) is a global 501(c)(3) organization, 501(c)(3) non-governmental organization, headquartered at the Bronx Zoo in New York City, with a mission to save "wildlife and wild places across the globe". Founded in ...
in September 2005. Gorillas are now known to use tools in the wild. A female gorilla in the Nouabalé-Ndoki National Park in the Republic of Congo
The Republic of the Congo, also known as Congo-Brazzaville, the Congo Republic or simply the Congo (the last ambiguously also referring to the neighbouring Democratic Republic of the Congo), is a country located on the western coast of Central ...
was recorded using a stick as if to gauge the depth of water whilst crossing a swamp. A second female was seen using a tree stump as a bridge and also as a support whilst fishing in the swamp. This means all of the great ape
The Hominidae (), whose members are known as the great apes or hominids (), are a taxonomic family of primates that includes eight extant species in four genera: '' Pongo'' (the Bornean, Sumatran and Tapanuli orangutan); '' Gorilla'' (the ...
s are now known to use tools.
In September 2005, a two-and-a-half-year-old gorilla in the Republic of Congo
The Republic of the Congo, also known as Congo-Brazzaville, the Congo Republic or simply the Congo (the last ambiguously also referring to the neighbouring Democratic Republic of the Congo), is a country located on the western coast of Central ...
was discovered using rocks to smash open palm nuts inside a game sanctuary. While this was the first such observation for a gorilla, over 40 years previously, chimpanzees had been seen using tools in the wild 'fishing' for termites. Nonhuman great apes are endowed with semiprecision grips, and have been able to use both simple tools and even weapons, such as improvising a club from a convenient fallen branch.
Scientific study
American physician and missionary Thomas Staughton Savage obtained the first specimens (the skull and other bones) during his time in Liberia
Liberia, officially the Republic of Liberia, is a country on the West African coast. It is bordered by Sierra Leone to Liberia–Sierra Leone border, its northwest, Guinea to Guinea–Liberia border, its north, Ivory Coast to Ivory Coast–Lib ...
. The first scientific description of gorillas dates back to an article by Savage and the naturalist Jeffries Wyman in 1847 in ''Proceedings of the Boston Society of Natural History'', where ''Troglodytes gorilla'' is described, now known as the western gorilla
The western gorilla (''Gorilla gorilla'') is a great ape found in Africa, one of two species of the hominid genus ''Gorilla''. Large and robust with males weighing around , the species is found in a region of midwest Africa, geographically iso ...
. Other species of gorilla were described in the next few years.[
]
The explorer Paul Du Chaillu
Paul Belloni Du Chaillu (July 31, 1831 (disputed)April 29, 1903) was a French-American traveler, zoologist, and anthropologist. He became famous in the 1860s as the first modern European outsider to confirm the existence of gorillas, and later t ...
was the first westerner to see a live gorilla during his travel through western equatorial Africa from 1856 to 1859. He brought dead specimens to the UK in 1861.
The first systematic study was not conducted until the 1920s, when Carl Akeley of the American Museum of Natural History
The American Museum of Natural History (AMNH) is a natural history museum on the Upper West Side of Manhattan in New York City. Located in Theodore Roosevelt Park, across the street from Central Park, the museum complex comprises 21 interconn ...
traveled to Africa to hunt for an animal to be shot and stuffed. On his first trip, he was accompanied by his friends Mary Bradley, a mystery writer, her husband, and their young daughter Alice, who would later write science fiction under the pseudonym James Tiptree Jr. After their trip, Mary Bradley wrote ''On the Gorilla Trail''. She later became an advocate for the conservation of gorillas, and wrote several more books (mainly for children). In the late 1920s and early 1930s, Robert Yerkes
Robert Mearns Yerkes (; May 26, 1876 – February 3, 1956) was an American psychologist, ethologist, eugenicist and primatologist best known for his work in intelligence testing and in the field of comparative psychology.
Yerkes was a pionee ...
and his wife Ava helped further the study of gorillas when they sent Harold Bigham to Africa. Yerkes also wrote a book in 1929 about the great apes.
After World War II
World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...
, George Schaller
George Beals Schaller (born 26 May 1933) is an American mammalogist, biologist, conservationist and author. Schaller is recognized by many as the world's preeminent field biologist, studying wildlife throughout Africa, Asia and South America. ...
was one of the first researchers to go into the field and study primates. In 1959, he conducted a systematic study of the mountain gorilla in the wild and published his work. Years later, at the behest of Louis Leakey
Louis Seymour Bazett Leakey (7 August 1903 – 1 October 1972) was a Kenyan-British palaeoanthropologist and archaeologist whose work was important in demonstrating that humans evolved in Africa, particularly through discoveries made at Olduvai ...
and the ''National Geographic
''National Geographic'' (formerly ''The National Geographic Magazine'', sometimes branded as ''Nat Geo'') is an American monthly magazine published by National Geographic Partners. The magazine was founded in 1888 as a scholarly journal, nine ...
'', Dian Fossey
Dian Fossey ( ; January 16, 1932 – ) was an American primatologist and conservationist known for undertaking an extensive study of mountain gorilla groups from 1966 until her murder in 1985. She studied them daily in the mountain forests of ...
conducted a much longer and more comprehensive study of the mountain gorilla. When she published her work, many misconceptions and myths about gorillas were finally disproved, including the myth that gorillas are violent.
Western lowland gorilla
The western lowland gorilla (''Gorilla gorilla gorilla'') is one of two Critically Endangered subspecies of the western gorilla (''Gorilla gorilla'') that lives in Montane ecosystems#Montane forests, montane, Old-growth forest, primary and sec ...
s (''G. g. gorilla'') are believed to be one of the zoonotic
A zoonosis (; plural zoonoses) or zoonotic disease is an infectious disease of humans caused by a pathogen (an infectious agent, such as a virus, bacterium, parasite, fungi, or prion) that can jump from a non-human vertebrate to a human. When h ...
origins of HIV/AIDS
The HIV, human immunodeficiency virus (HIV) is a retrovirus that attacks the immune system. Without treatment, it can lead to a spectrum of conditions including acquired immunodeficiency syndrome (AIDS). It is a Preventive healthcare, pr ...
. The SIVgor Simian immunodeficiency virus
Simian immunodeficiency virus (SIV) is a species of retrovirus that cause persistent infections in at least 45 species of non-human primates. Based on analysis of strains found in four species of monkeys from Bioko Island, which was isolated fr ...
that infects them is similar to a certain strain of HIV-1.
Genome sequencing
The gorilla became the next-to-last great ape genus to have its genome sequenced. The first gorilla genome was generated with short read and Sanger sequencing using DNA from a female western lowland gorilla named Kamilah. This gave scientists further insight into the evolution and origin of humans. Despite the chimpanzees being the closest extant relatives of humans, 15% of the human genome was found to be more like that of the gorilla. In addition, 30% of the gorilla genome "is closer to human or chimpanzee than the latter are to each other; this is rarer around coding genes, indicating pervasive selection throughout great ape evolution, and has functional consequences in gene expression." Analysis of the gorilla genome has cast doubt on the idea that the rapid evolution of hearing genes gave rise to language in humans, as it also occurred in gorillas.
Captivity
Gorillas became highly prized by western zoo
A zoo (short for zoological garden; also called an animal park or menagerie) is a facility where animals are kept within enclosures for public exhibition and often bred for conservation purposes.
The term ''zoological garden'' refers to zoology, ...
s since the 19th century, though the earliest attempts to keep them in captive facilities ended in their early death. In the late 1920s, the care of captive gorillas significantly improved. Colo (December 22, 1956 – January 17, 2017) of the Columbus Zoo and Aquarium
The Columbus Zoo and Aquarium is a non-profit zoo located near Powell, Ohio, Powell in Liberty Township, Delaware County, Ohio, United States, north of the city of Columbus, Ohio, Columbus. The land lies along the eastern banks of the O'Shaughnes ...
was the first gorilla to be born in captivity.
Captive gorillas exhibit stereotypic behaviors, including eating such as regurgitation, reingestion and self-injurious or conspecific aggression, pacing, rocking, sucking of fingers or lip smacking, and overgrooming. Negative vigilance of visitor behaviors have been identified as starting, posturing and charging at visitors. Groups of bachelor gorillas containing young silverbacks have significantly higher levels of aggression and wounding rates than mixed age and sex groups.
The use of both internal and external privacy screens on exhibit windows has been shown to alleviate stresses from visual effects of high crowd densities, leading to decreased stereotypic behaviors in the gorillas. Playing naturalistic auditory stimuli as opposed to classical music, rock music, or no auditory enrichment (which allows for crowd noise, machinery, etc. to be heard) has been noted to reduce stress behavior as well. Enrichment modifications to feed and foraging, where clover-hay is added to an exhibit floor, decrease stereotypic activities while simultaneously increasing positive food-related behaviors.
Recent research on captive gorilla welfare emphasizes a need to shift to individual assessments instead of a one-size-fits-all group approach to understanding how welfare increases or decreases based on a variety of factors. Individual characteristics such as age, sex, personality and individual histories are essential in understanding that stressors will affect each individual gorilla and their welfare
Welfare may refer to:
Philosophy
*Well-being (happiness, prosperity, or flourishing) of a person or group
* Utility in utilitarianism
* Value in value theory
Economics
* Utility, a general term for individual well-being in economics and decision ...
differently.
Conservation status
All species (and subspecies) of gorilla are listed as endangered
An endangered species is a species that is very likely to become extinct in the near future, either worldwide or in a particular political jurisdiction. Endangered species may be at risk due to factors such as habitat loss, poaching, inv ...
or critically endangered
An IUCN Red List critically endangered (CR or sometimes CE) species is one that has been categorized by the International Union for Conservation of Nature as facing an extremely high risk of extinction in the wild. As of December 2023, of t ...
on the IUCN Red List
The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
. All gorillas are listed in Appendix I of the Convention on International Trade in Endangered Species
Convention may refer to:
* Convention (norm), a custom or tradition, a standard of presentation or conduct
** Treaty, an agreement in international law
** Convention (political norm), uncodified legal or political tradition
* Convention (meeting ...
(CITES), meaning that international export/import of the species, including in parts and derivatives, is regulated. Around 316,000 western lowland gorillas are thought to exist in the wild, 4,000 in zoos, thanks to conservation; eastern lowland gorillas have a population of under 5,000 in the wild and 24 in zoos. Mountain gorillas are the most severely endangered, with an estimated population of about 880 left in the wild and none in zoos. Threats to gorilla survival include habitat destruction
Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
and poaching
Poaching is the illegal hunting or capturing of wild animals, usually associated with land use rights.
Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set against the huntin ...
for the bushmeat
Bushmeat is meat from wildlife species that are Hunting, hunted for human consumption. Bushmeat represents a primary source of animal protein and a cash-earning commodity in poor and rural communities of humid tropical forest regions of the worl ...
trade. Gorillas are closely related to humans, and are susceptible to diseases that humans also get infected by. In 2004, a population of several hundred gorillas in the Odzala National Park, Republic of Congo
The Republic of the Congo, also known as Congo-Brazzaville, the Congo Republic or simply the Congo (the last ambiguously also referring to the neighbouring Democratic Republic of the Congo), is a country located on the western coast of Central ...
was essentially wiped out by the Ebola virus
''Orthoebolavirus zairense'' or Zaire ebolavirus, more commonly known as Ebola virus (; EBOV), is one of six known species within the genus ''Ebolavirus''. Four of the six known ebolaviruses, including EBOV, cause a severe and often fatal vira ...
. A 2006 study published in ''Science
Science is a systematic discipline that builds and organises knowledge in the form of testable hypotheses and predictions about the universe. Modern science is typically divided into twoor threemajor branches: the natural sciences, which stu ...
'' concluded more than 5,000 gorillas may have died in recent outbreaks of the Ebola virus in central Africa. The researchers indicated in conjunction with commercial hunting of these apes, the virus creates "a recipe for rapid ecological extinction". In captivity, it has also been observed that gorillas can also be infected with COVID-19.
Conservation efforts include the Great Apes Survival Project, a partnership between the United Nations Environment Programme
The United Nations Environment Programme (UNEP) is responsible for coordinating responses to environmental issues within the United Nations system. It was established by Maurice Strong, its first director, after the Declaration of the United Nati ...
and the UNESCO
The United Nations Educational, Scientific and Cultural Organization (UNESCO ) is a List of specialized agencies of the United Nations, specialized agency of the United Nations (UN) with the aim of promoting world peace and International secur ...
, and also an international treaty, the Agreement on the Conservation of Gorillas and Their Habitats
The Agreement on the Conservation of Gorillas and Their Habitats, also known as the Gorilla Agreement (French language, French: ''Accord Gorille''), is a multilateral environmental agreement that binds the parties to conserve gorillas in their te ...
, concluded under UNEP-administered Convention on Migratory Species
The Convention on the Conservation of Migratory Species of Wild Animals, also known as the Convention on Migratory Species (CMS) or the Bonn Convention, is an international agreement that aims to conserve migratory species throughout their r ...
. The Gorilla Agreement is the first legally binding instrument exclusively targeting gorilla conservation; it came into effect on 1 June 2008. Governments of countries where gorillas live placed a ban on their killing and trading, but weak law enforcement still poses a threat to them, since the governments rarely apprehend poachers, traders and consumers that rely on gorillas for profit.
Cultural significance
In Cameroon's Lebialem
Lebialem is a department of Southwest Region in Cameroon. The Division covers an area of 617 km and as of 2005 had a total population of 113,736. The capital of the department lies at Menji
Menji is a town and capital of the Fontem distri ...
highlands, folk stories connect people and gorillas via totem
A totem (from or ''doodem'') is a spirit being, sacred object, or symbol that serves as an emblem of a group of people, such as a family, clan, lineage (anthropology), lineage, or tribe, such as in the Anishinaabe clan system.
While the word ...
s; a gorilla's death means the connected person will die also. This creates a local conservation ethic. Many different indigenous peoples interact with wild gorillas. Some have detailed knowledge; the Baka have words to distinguish at least ten types of gorilla individuals, by sex, age, and relationships. In 1861, alongside tales of hunting enormous gorillas, the traveller and anthropologist Paul Du Chaillu
Paul Belloni Du Chaillu (July 31, 1831 (disputed)April 29, 1903) was a French-American traveler, zoologist, and anthropologist. He became famous in the 1860s as the first modern European outsider to confirm the existence of gorillas, and later t ...
reported the Cameroonian story that a pregnant woman who sees a gorilla will give birth to one.
In 1911, the anthropologist Albert Jenks noted the Bulu people
The Beti-Pahuin are a Bantu ethnic group located in Center region of Cameroon. Though they separate themselves into several individual clans, they all share a common origin, history and culture. Estimated to be well over 8 million individuals i ...
's knowledge of gorilla behaviour and ecology, and their gorilla stories. In one such story, "The Gorilla and the Child", a gorilla speaks to people, seeking help and trust, and stealing a baby; a man accidentally kills the baby while attacking the gorilla. Even far from where gorillas live, savannah tribes pursue "cult-like worship" of the apes. Some beliefs are widespread among indigenous peoples. The Fang
A fang is a long, pointed tooth. In mammals, a fang is a modified maxillary tooth, used for biting and tearing flesh. In snakes, it is a specialized tooth that is associated with a venom gland (see snake venom). Spiders also have external fangs, ...
name for gorilla is ''ngi'' while the Bulu name is ''njamong''; the root ''ngi'' means fire, denoting a positive energy. From the Central African Republic to Cameroon and Gabon, stories of reincarnations as gorillas, totems, and transformations similar to those recorded by Du Chaillu are still told in the 21st century.
Since gaining international attention, gorillas have been a recurring element of many aspects of popular culture and media. They were usually portrayed as murderous and aggressive. Inspired by Emmanuel Frémiet's ''Gorilla Carrying off a Woman'', gorillas have been depicted kidnapping human women. This theme was used in films such as ''Ingagi
''Ingagi'' is a 1930 pre-Code Hollywood, pre-Code pseudo-documentary exploitation film directed by William S. Campbell (director), William S. Campbell. It purports to be a documentary about "Sir Hubert Winstead" of London on an expedition to the ...
'' (1930) and most notably ''King Kong
King Kong, also referred to simply as Kong, is a fictional giant monster resembling a gorilla, who has appeared in various media since 1933. The character has since become an international pop culture icon,Erb, Cynthia, 1998, ''Tracking Kin ...
'' (1933). The comedic play '' The Gorilla'', which debuted in 1925, featured an escaped gorilla taking a woman from her house. Several films would use the "escaped gorilla" trope including '' The Strange Case of Doctor Rx'' (1942), ''The Gorilla Man
''The Gorilla Man'' is a 1943 American drama film directed by D. Ross Lederman. Despite the title and marketing, it is not a horror film, but a World War II espionage thriller.
Plot
Military officer Captain Craig Killian is wounded in the lin ...
'' (1943), ''Gorilla at Large
''Gorilla at Large'' is a 1954 American horror mystery film made in 3-D. The film stars Cameron Mitchell, Anne Bancroft, Lee J. Cobb and Raymond Burr, with Lee Marvin and Warren Stevens in supporting roles. Directed by Harmon Jones, it was ...
'' (1954) and the Disney
The Walt Disney Company, commonly referred to as simply Disney, is an American multinational mass media and entertainment industry, entertainment conglomerate (company), conglomerate headquartered at the Walt Disney Studios (Burbank), Walt Di ...
cartoons '' The Gorilla Mystery'' (1930) and '' Donald Duck and the Gorilla'' (1944).
Gorillas have been used as opponents to jungle-themed heroes such as Tarzan
Tarzan (John Clayton, Viscount Greystoke) is a fictional character, a feral child raised in the African jungle by the Mangani great apes; he later experiences civilization, only to reject it and return to the wild as a heroic adventurer.
Creat ...
and Sheena, Queen of the Jungle
Sheena, Queen of the Jungle, is a fictional American comic book jungle girl heroine during the Golden Age of Comic Books. She originally debuted in the British magazine ''Wags'' #46 (January 1938). and later made her first American appearance i ...
, as well as superheroes. The DC Comics
DC Comics (originally DC Comics, Inc., and also known simply as DC) is an American comic book publisher owned by DC Entertainment, a subsidiary of Warner Bros. Discovery. DC is an initialism for "Detective Comics", an American comic book seri ...
supervillain Gorilla Grodd
Gorilla Grodd is a supervillain appearing in American comic books published by DC Comics. Created by John Broome and Carmine Infantino, the character first appeared in ''The Flash'' #106 (May 1959).
He is an evil, super-intelligent gorilla who ...
is an enemy of the Flash
The Flash is the name of several superheroes appearing in American comic books published by DC Comics. Created by writer Gardner Fox and artist Harry Lampert, the original Flash first appeared in ''Flash Comics'' #1 (cover date, cover-dated Jan ...
. Gorillas also serve as antagonists in the 1968 film ''Planet of the Apes
''Planet of the Apes'' is a science fiction media franchise consisting of films, books, television series, comics, and other media about a Apocalyptic and post-apocalyptic fiction, post-apocalyptic world in which humans and intelligent apes c ...
''. More positive and sympathetic portrayals of gorillas include the films '' Son of Kong'' (1933), '' Mighty Joe Young'' (1949), ''Gorillas in the Mist
''Gorillas in the Mist'' is a 1988 American biographical drama film directed by Michael Apted from a screenplay by Anna Hamilton Phelan and a story by Phelan and Tab Murphy. The film is based on a book of the same name by Dian Fossey and fr ...
'' (1988) and ''Instinct
Instinct is the inherent inclination of a living organism towards a particular complex behaviour, containing innate (inborn) elements. The simplest example of an instinctive behaviour is a fixed action pattern (FAP), in which a very short to me ...
'' (1999) and the 1992 novel ''Ishmael
In the Bible, biblical Book of Genesis, Ishmael (; ; ; ) is the first son of Abraham. His mother was Hagar, the handmaiden of Abraham's wife Sarah. He died at the age of 137. Traditionally, he is seen as the ancestor of the Arabs.
Within Isla ...
''. Gorillas have been featured in video games as well, notably ''Donkey Kong
is a video game series and media franchise created by the Japanese game designer Shigeru Miyamoto for Nintendo. It follows the adventures of Donkey Kong (character), Donkey Kong, a large, powerful gorilla, and other members of the List of Don ...
''.
In April 2025, a viral online debate arose regarding whether 100 men could beat a gorilla, prompting responses from celebrities and experts.
See also
* Bili ape
The Bili apes, or Bondo mystery apes, were names given in 2003 in sensational reports in the popular media to a purportedly new species of highly aggressive, giant ape supposedly inhabiting the Bili Forest, wetlands and savannah around of the vill ...
* Gorilla (advertisement)
''Gorilla'' is a British advertising campaign launched by the advertising agency Fallon Worldwide, Fallon London on behalf of Cadbury Schweppes in 2007, to promote Dairy Milk, Cadbury Dairy Milk brand chocolate. The centrepiece of the campaign ...
* Gorilla suit
Gorilla suits are a type of creature suit resembling a gorilla. The gorilla suit is a popular Halloween and costume party costume, and is also used as a source of humour, while more realistic suits have been used both to represent real gorillas ...
* Great Ape Project
The Great Ape Project (GAP), founded in 1993, is an international organization of primatologists, anthropologists, ethicists, and others who advocate a United Nations Declaration of the Rights of Great Apes that would confer basic legal rights ...
* Harambe
Harambe ( ; May 27, 1999 – May 28, 2016) was a western lowland gorilla who lived at the Cincinnati Zoo and Botanical Garden, Cincinnati Zoo. On May 28, 2016, a three-year-old boy visiting the zoo climbed under a fence into an outdoor gorilla ...
– a captive gorilla, shot while interacting with a child who fell into his zoo enclosure, who became a popular internet meme
* International Primate Day
International Primate Day, September 1, is an annual educational observance event organized since 2005 largely by British-based Animal Defenders International (ADI)
and supported annually by various primate-oriented advocacy organizations, speak ...
* King Kong
King Kong, also referred to simply as Kong, is a fictional giant monster resembling a gorilla, who has appeared in various media since 1933. The character has since become an international pop culture icon,Erb, Cynthia, 1998, ''Tracking Kin ...
* List of individual apes
This is a list of non-human apes of encyclopedic interest. It includes individual chimpanzees, gorillas, orangutans, bonobos, and gibbons that are in some way famous or notable.
Actors
* J. Fred Muggs (a chimpanzee born 1952) was a "co-host" ...
* List of fictional apes
This list of fictional primates is a subsidiary to the list of fictional animals. The list is restricted to notable non-human primate characters from the world of fiction including chimpanzees, gorillas, orangutans, monkeys, lemurs, and other prim ...
* Monkey Day
References
Literature cited
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External links
Animal Diversity Web
– includes photos, artwork, and skull specimens of ''Gorilla gorilla''
Pulitzer Center on Crisis Reporting
International Gorilla Conservation Programme
The International Gorilla Conservation Programme (IGCP; French language, French: ''Programme international de conservation des gorilles'') was formed in 1991 to ensure that the critically endangered mountain gorillas are conserved in their habitat ...
(Video) (archived 5 September 2007)
Primate Info Net ''Gorilla'' Factsheet
– taxonomy, ecology, behavior and conservation
– features a video and photos
World Wildlife Fund: Gorillas
– conservation, facts and photos (archived 16 October 2004)
Gorilla protection
– Gorilla conservation (archived 29 November 2010)
Welcome to the Year of the Gorilla 2009
* – The Official Website for Virunga National Park, the Last Refuge for Congo's Mountain Gorillas (archived 22 September 2010)
Human Timeline (Interactive)
– Smithsonian, National Museum of Natural History
The National Museum of Natural History (NMNH) is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. With 4.4 ...
(August 2016).
Genome of ''Gorilla gorilla''
via Ensembl
Ensembl genome database project is a scientific project at the European Bioinformatics Institute, which provides a centralized resource for geneticists, molecular biologists and other researchers studying the genomes of our own species and other v ...
Genome of ''Gorilla gorilla'' (version Kamilah_GGO_v0/gorGor6)
via UCSC Genome Browser
The UCSC Genome Browser is an online and downloadable genome browser hosted by the University of California, Santa Cruz (UCSC). It is an interactive website offering access to genome sequence data from a variety of vertebrate and invertebrate spec ...
Data of the genome of ''Gorilla gorilla''
via NCBI
The National Center for Biotechnology Information (NCBI) is part of the National Library of Medicine (NLM), a branch of the National Institutes of Health (NIH). It is approved and funded by the government of the United States. The NCBI is loca ...
Data of the genome assembly of ''Gorilla gorilla'' (version Kamilah_GGO_v0/gorGor6)
via NCBI
The National Center for Biotechnology Information (NCBI) is part of the National Library of Medicine (NLM), a branch of the National Institutes of Health (NIH). It is approved and funded by the government of the United States. The NCBI is loca ...
{{Authority control
Gorillini
Herbivorous mammals
Fauna of Sub-Saharan Africa
Primate genera
Taxa named by Isidore Geoffroy Saint-Hilaire
Taxa described in 1852