''Siamodon'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

iguanodontia Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...

ornithopod Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (). They represent one of the most successful groups of herbivorous dinosaurs during the Cretaceous. The most primitive members of the group were bipedal and relatively sm ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic Geological period, period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the #Evolutio ...

from

Early Cretaceous The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name) is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 143.1 ...

deposits of northeastern

Thailand Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...

Discovery

''Siamodon'' is known from the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

PRC-4, a well-preserved left

maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...

and from the referred materials PRC-5, an isolated maxillary

tooth A tooth (: teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tea ...

and PRC-6, a

braincase In human anatomy, the neurocranium, also known as the braincase, brainpan, brain-pan, or brainbox, is the upper and back part of the skull, which forms a protective case around the brain. In the human skull, the neurocranium includes the calv ...

. It was collected in the Ban Saphan Hin site, Nakhon Ratchasima Province, from the

Khok Kruat Formation The Khok Kruat Formation () is a geological formation, rock formation found in northeastern Thailand. It is the uppermost formation of the Khorat Group. It is dated to the Aptian Geological stage, stage of the Early Cretaceous Geological period, ...

, dating to the

Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...

stage of the late

, about 125-113

million years ago Million years ago, abbreviated as Mya, Myr (megayear) or Ma (megaannum), is a unit of time equal to (i.e. years), or approximately 31.6 teraseconds. Usage Myr is in common use in fields such as Earth science and cosmology. Myr is also used w ...

Description

''Siamodon'' shows a combination of plesiomorphic and apomorphic features, including a maxilla shaped like an

isosceles triangle In geometry, an isosceles triangle () is a triangle that has two Edge (geometry), sides of equal length and two angles of equal measure. Sometimes it is specified as having ''exactly'' two sides of equal length, and sometimes as having ''at le ...

, with the dorsal process located at about mid-length of the bone; a strong longitudinal bulge on the medial surface of the maxilla; at least 25 maxillary teeth, which bear a prominent median primary ridge, and one short weak subsidiary ridge or no subsidiary ridge at all, and mamillated denticles on the crown margins. The maxilla is 230 millimeters long, and has a height of 100 millimeters. The height of the isolated tooth is about 25-28 millimeters, and the width is about 14-17 millimeters. ''Siamodon'' differs from more basal iguanodontians, such as ''

Iguanodon ''Iguanodon'' ( ; meaning 'iguana-tooth'), named in 1825, is a genus of iguanodontian dinosaur. While many species found worldwide have been classified in the genus ''Iguanodon'', dating from the Late Jurassic to Early Cretaceous, Taxonomy (bi ...

'' and closely related forms, in the morphology of its maxillary teeth, which are narrower and bear a strong median primary ridge, sometimes accompanied by a weak subsidiary ridge, instead of a distally displaced primary ridge and several subsidiary ridges, and the apex of the maxilla is in a more posterior position. Its maxilla differs from that of

hadrosaurids Hadrosaurids (), also hadrosaurs or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod fami ...

in the articular area for the jugal, with forms a tab-like jugal process. Whereas in hadrosaurids, the expanded anterior end of the jugal contacts and overlaps a large sutural area on the maxilla. The combination of characters seen in the maxilla of ''Siamodon nimngami'' indicate that it belongs to a group of iguanodontians more derived than ''Iguanodon'' but basal to Hadrosauridae. It may be closely related to ''

Probactrosaurus ''Probactrosaurus'' (meaning "before ''Bactrosaurus''") is an early herbivorous hadrosauroid iguanodont dinosaur. It lived in China during the Early Cretaceous Period (geology), period. Discovery and species In 1959 and 1960 a Soviet-Chinese ex ...

'' from China but they differ in the number of tooth positions.

Etymology

''Siamodon'' was first named by

Éric Buffetaut Éric Buffetaut (born 19 November 1950) is a French paleontologist, author and researcher at the Centre national de la recherche scientifique since 1976 where he is a Doctor of Science and Director of Research. Buffetaut is a specialist of fossil ...

and Varavudh Suteethorn in

2011 The year marked the start of a Arab Spring, series of protests and revolutions throughout the Arab world advocating for democracy, reform, and economic recovery, later leading to the depositions of world leaders in Tunisia, Egypt, and Yemen ...

. The

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Siamodon nimngami''. The generic name is derived from

Siam Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...

, the ancient name of Thailand; and ',

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

for "tooth". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

is in honour of Witaya Nimngam, who donated the specimens to the authors.

References

{{Taxonbar, from=Q1705962 Hadrosauroidea Dinosaur genera Aptian dinosaurs Taxa named by Éric Buffetaut Taxa named by Varavudh Suteethorn Fossil taxa described in 2011 Dinosaurs of Thailand