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Shrubland, scrubland, scrub, brush, or bush is a
plant community A plant community is a collection or Association (ecology), association of plant species within a designated geographical unit, which forms a relatively uniform patch, distinguishable from neighboring patches of different vegetation types. The comp ...
characterized by
vegetation Vegetation is an assemblage of plants and the ground cover they provide. It is a general term, without specific reference to particular Taxon, taxa, life forms, structure, Spatial ecology, spatial extent, or any other specific Botany, botanic ...
dominated by
shrubs A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
, often also including
grasses Poaceae ( ), also called Gramineae ( ), is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos, the grasses of natural grassland and species cultivated in ...
,
herbs Herbs are a widely distributed and widespread group of plants, excluding vegetables, with savory or aromatic properties that are used for flavoring and garnish (food), garnishing food, for medicinal purposes, or for fragrances. Culinary use typi ...
, and geophytes. Shrubland may either occur naturally or be the result of human activity. It may be the mature
vegetation type Vegetation classification is the process of classifying and mapping the vegetation over an area of the Earth's surface. Vegetation classification is often performed by state based agencies as part of land use, resource and environmental management ...
in a particular region and remain stable over time, or it may be a transitional community that occurs temporarily as the result of a disturbance, such as fire. A stable state may be maintained by regular natural disturbance such as fire or
browsing Browsing is a kind of orienting strategy. It is supposed to identify something of relevance for the browsing organism. In context of humans, it is a metaphor taken from the animal kingdom. It is used, for example, about people browsing open sh ...
. Shrubland may be unsuitable for human habitation because of the danger of fire. The term was coined in 1903. Shrubland species generally show a wide range of
adaptations In biology, adaptation has three related meanings. Firstly, it is the dynamic evolutionary process of natural selection that fits organisms to their environment, enhancing their evolutionary fitness. Secondly, it is a state reached by the p ...
to fire, such as heavy seed production, lignotubers, and fire-induced germination.


Botanical structural form

In
botany Botany, also called plant science, is the branch of natural science and biology studying plants, especially Plant anatomy, their anatomy, Plant taxonomy, taxonomy, and Plant ecology, ecology. A botanist or plant scientist is a scientist who s ...
and
ecology Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
a shrub is defined as a much-branched
woody plant A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposed to Herbaceous plant, herbaceous plants that die back to t ...
less than 8 m high, usually with many stems. Tall shrubs are mostly 2–8 m high, small shrubs 1–2 m high and
subshrubs A subshrub (Latin ''suffrutex'') or undershrub is either a small shrub (e.g. prostrate shrubs) or a perennial that is largely herbaceous but slightly woody plant, woody at the base (e.g. garden pink and florist's chrysanthemum). The term is ofte ...
less than 1 m high. There is a descriptive system widely adopted in
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
to describe different types of vegetation is based on structural characteristics based on
plant life-form Plant life-form schemes constitute a way of classifying plants alternatively to the ordinary species-genus-family scientific classification. In colloquial speech, plants may be classified as trees, shrubs, herbs (forbs and graminoids), etc. The sci ...
, as well as the height and foliage cover of the tallest stratum or dominant
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
. For shrubs that are high, the following structural forms are categorized: * dense foliage cover (70–100%) — closed-shrubs * mid-dense foliage cover (30–70%) — open-shrubs * sparse foliage cover (10–30%) — tall shrubland * very sparse foliage cover (<10%) — tall open shrubland For shrubs less than high, the following structural forms are categorized: * dense foliage cover (70–100%) — closed-
heath A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and is characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...
or closed low shrubland—(''North America'') * mid-dense foliage cover (30–70%) — open-heath or mid-dense low shrubland—(''North America'') * sparse foliage cover (10–30%) — low shrubland * very sparse foliage cover (<10%) — low open shrubland


Biome plant group

Similarly, shrubland is a category that is used to describe a type of
biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...
plant group. In this context, shrublands are dense thickets of evergreen
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short Internode (botany), internodes (the distance between leaves along the stem) and leaf orientation which is parallel or ...
shrubs and small trees, called: *
Chaparral Chaparral ( ) is a shrubland plant plant community, community found primarily in California, southern Oregon, and northern Baja California. It is shaped by a Mediterranean climate (mild wet winters and hot dry summers) and infrequent, high-intens ...
in
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
*
Matorral 300px, Springtime in Santiago.html" ;"title="Chilean matorral a few kilometers north of Santiago">Chilean matorral a few kilometers north of Santiago along the Pan-American Highway Matorral is a Spanish language, Spanish word, along with ''tomil ...
in
Chile Chile, officially the Republic of Chile, is a country in western South America. It is the southernmost country in the world and the closest to Antarctica, stretching along a narrow strip of land between the Andes, Andes Mountains and the Paci ...
,
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
, and
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
* Maquis in
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
and elsewhere around the
Mediterranean The Mediterranean Sea ( ) is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean basin and almost completely enclosed by land: on the east by the Levant in West Asia, on the north by Anatolia in West Asia and Southern ...
* Macchia in
Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...
*
Fynbos Fynbos (; , ) is a small belt of natural shrubland or heathland vegetation located in the Western Cape and Eastern Cape provinces of South Africa. The area is predominantly coastal and mountainous, with a Mediterranean climate. The fynbos ...
in
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
* Eastern Suburbs Banksia Scrub in
Sydney Sydney is the capital city of the States and territories of Australia, state of New South Wales and the List of cities in Australia by population, most populous city in Australia. Located on Australia's east coast, the metropolis surrounds Syd ...
* Kwongan in Southwest
Australia Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
* Cedar scrub in
Texas Hill Country The Texas Hill Country is a geographic region of Central and South Texas, forming the southeast part of the Edwards Plateau. Given its location, climate, terrain, and vegetation, the Hill Country can be considered the border between the Ame ...
*
Caatinga Caatinga () is a type of semi-arid tropical vegetation, and an ecoregion characterized by this vegetation in interior northeastern Brazil. The name "Caatinga" comes from the Tupi word '' ka'atinga'', meaning "white forest" or "white vegetat ...
in northeastern Brazil In some places, shrubland is the mature vegetation type. In other places, it is the result of degradation of former
forest A forest is an ecosystem characterized by a dense ecological community, community of trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, ...
or
woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...
by
logging Logging is the process of cutting, processing, and moving trees to a location for transport. It may include skidder, skidding, on-site processing, and loading of trees or trunk (botany), logs onto logging truck, trucksovergrazing Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature ...
, or disturbance by major fires. A number of
World Wildlife Fund The World Wide Fund for Nature (WWF) is a Swiss-based international non-governmental organization founded in 1961 that works in the field of wilderness preservation and the reduction of human impact on the environment. It was formerly named the ...
biomes are characterized as shrublands, including the following: ;Desert scrublands ''Xeric'' or ''desert scrublands'' occur in the world's
deserts and xeric shrublands Deserts and xeric shrublands are a biome defined by the World Wide Fund for Nature. Deserts and xeric (Ancient Greek 'dry') shrublands form the largest terrestrial biome, covering 19% of Earth's land surface area. Ecoregions in this habitat ...
ecoregions or in fast-draining sandy soils in more humid regions. These scrublands are characterized by plants with adaptations to the dry climate, which include small leaves to limit water loss, thorns to protect them from grazing animals, succulent leaves or stems, storage organs to store water, and long taproots to reach groundwater. ;Mediterranean scrublands ''Mediterranean scrublands'' occur naturally in the Mediterranean scrub
biome A biome () is a distinct geographical region with specific climate, vegetation, and animal life. It consists of a biological community that has formed in response to its physical environment and regional climate. In 1935, Tansley added the ...
, located in the five
Mediterranean climate A Mediterranean climate ( ), also called a dry summer climate, described by Köppen and Trewartha as ''Cs'', is a temperate climate type that occurs in the lower mid-latitudes (normally 30 to 44 north and south latitude). Such climates typic ...
regions of the world. Scrublands are most common near the seacoast and have often adapted to the wind and salt air of the ocean. Low, soft-leaved scrublands around the Mediterranean Basin are known as ''
garrigue Garrigue or garigue ( ), also known as phrygana ( , n. pl.), is a type of low scrubland ecoregion and plant community in the Mediterranean forests, woodlands, and scrub biome. It is found on limestone soils in southern France and around the ...
'' in
France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...
, ''phrygana'' in
Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...
, ''tomillares'' in
Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...
, and ''batha'' in
Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...
.
Northern coastal scrub Northern coastal scrub is a diverse scrubland plant community found along the Pacific Coast from Northern California to Southern Oregon, as well as some offshore islands. It frequently forms a landscape mosaic with California coastal prairie, coast ...
and
coastal sage scrub Coastal sage scrub, also known as coastal scrub, CSS, or soft chaparral, is a low scrubland plant community of the California coastal sage and chaparral subecoregion, found in coastal California and northwestern coastal Baja California. It is ...
occur along the
California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...
coast, strandveld in the
Western Cape The Western Cape ( ; , ) is a provinces of South Africa, province of South Africa, situated on the south-western coast of the country. It is the List of South African provinces by area, fourth largest of the nine provinces with an area of , an ...
of
South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...
, coastal matorral in central Chile, and sand-heath and kwongan in
Southwest Australia Southwest Australia is a biogeographic region in Western Australia. It includes the Mediterranean-climate area of southwestern Australia, which is home to a diverse and distinctive flora and fauna. The region is also known as the Southwest Au ...
. ;Interior scrublands ''Interior scrublands'' occur naturally in semi-arid areas with nutrient-poor soils, such as on the matas of
Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...
, which are underlain by
Cambrian The Cambrian ( ) is the first geological period of the Paleozoic Era, and the Phanerozoic Eon. The Cambrian lasted 51.95 million years from the end of the preceding Ediacaran period 538.8 Ma (million years ago) to the beginning of the Ordov ...
and
Silurian The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of t ...
schists Schist ( ) is a medium-grained metamorphic rock generally derived from fine-grained sedimentary rock, like shale. It shows pronounced ''schistosity'' (named for the rock). This means that the rock is composed of mineral grains easily seen with a ...
.
Florida scrub Florida scrub is a forest ecoregion found throughout Florida in the United States. It is found on coastal and inland sand ridges and is characterized by an evergreen Xerophyte, xeromorphic plant community dominated by shrubs and dwarf oaks. Becau ...
is another example of interior scrublands. ;Dwarf shrubs Some vegetation types are formed of '' dwarf-shrubs'', low-growing or creeping shrubs. They include the maquis and the garrigues of Mediterranean climates and the acid-loving dwarf shrubs of
heathland A heath () is a shrubland habitat found mainly on free-draining infertile, acidic soils and is characterised by open, low-growing woody vegetation. Moorland is generally related to high-ground heaths with—especially in Great Britain—a coole ...
and
moorland Moorland or moor is a type of Habitat (ecology), habitat found in upland (geology), upland areas in temperate grasslands, savannas, and shrublands and the biomes of montane grasslands and shrublands, characterised by low-growing vegetation on So ...
.


See also

* Prostrate shrub * Semi-desert * Shrub-steppe *
Shrub swamp Shrub swamps — also called scrub swamps or buttonbush swamps — are a type of freshwater wetland ecosystem occurring in areas too wet to become swamps ("true" or freshwater swamp forest), but too dry or too shallow to become marshes. They a ...


Notes and references


External links

* {{Vegetation