botany
Botany, also called plant science (or plant sciences), plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "bot ...
, a plant shoot consists of any
plant stem
A stem is one of two main structural axes of a vascular plant, the other being the root. It supports leaves, flowers and fruits, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrie ...
together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the spring,
perennial
A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
plant shoots are the new growth that grows from the ground in
herbaceous
Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials.
Definitions of "herb" and "herbaceous"
The fourth edition ...
plants or the new stem or flower growth that grows on woody plants.
In everyday speech, shoots are often synonymous with stems. Stems, which are an integral component of shoots, provide an axis for buds, fruits, and leaves.
Young shoots are often eaten by animals because the fibers in the new growth have not yet completed
secondary cell wall The secondary cell wall is a structure found in many plant cells, located between the primary cell wall and the plasma membrane. The cell starts producing the secondary cell wall after the primary cell wall is complete and the cell has stopped expan ...
development, making the young shoots softer and easier to chew and digest. As shoots grow and age, the cells develop secondary cell walls that have a hard and tough structure. Some plants (e.g.
bracken
Bracken (''Pteridium'') is a genus of large, coarse ferns in the family (biology), family Dennstaedtiaceae. Ferns (Pteridophyta) are vascular plants that have alternating generations, large plants that produce spores and small plants that produ ...
) produce
toxin
A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849 ...
s that make their shoots inedible or less palatable.
File:Cucumber leaf.jpg, The shoot of a
cucumber
Cucumber (''Cucumis sativus'') is a widely-cultivated creeping vine plant in the Cucurbitaceae family that bears usually cylindrical fruits, which are used as culinary vegetables.Sachaline
File:Sunflower seedlings.jpg, Sunflower seedlings
germinate
Germination is the process by which an organism grows from a seed or spore. The term is applied to the sprouting of a seedling from a seed of an angiosperm or gymnosperm, the growth of a sporeling from a spore, such as the spores of fungi, fer ...
File:Persea americana (Avocado) Sprout 08May2010.JPG, A young hass avocado shoot
Shoot types of woody plants
Many
woody plant
A woody plant is a plant that produces wood as its structural tissue and thus has a hard stem. In cold climates, woody plants further survive winter or dry season above ground, as opposite to herbaceous plants that die back to the ground until s ...
s have distinct short shoots and long shoots. In some
angiosperm
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. They include all forbs (flowering plants without a woody stem), grasses and grass-like plants, a vast majority of br ...
s, the short shoots, also called spur shoots or fruit spurs, produce the majority of flowers and fruit. A similar pattern occurs in some
conifer
Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All ex ...
s and in '' Ginkgo'', although the "short shoots" of some genera such as ''Picea'' are so small that they can be mistaken for part of the leaf that they have produced.
A related phenomenon is seasonal heterophylly, which involves visibly different leaves from spring growth and later lammas growth. Whereas spring growth mostly comes from buds formed the previous season, and often includes flowers, lammas growth often involves long shoots.
File: Suckers on stump.jpg, Long shoot growth
File: Pyrus pyrifolia (Hosui) blossom.jpg, A mature fruiting spur on a Nashi pear tree, '' Pyrus pyrifolia''
File: Cedrus deodara 02.JPG, On long shoots of '' Cedrus deodara'' individual leaves may have buds in the axils.
File:20130903Cedrus deodara2.jpg, ''Cedrus deodara'' forms short shoots (from buds) along the long shoots.
Seedling
A seedling is a young sporophyte developing out of a plant embryo from a seed. Seedling development starts with germination of the seed. A typical young seedling consists of three main parts: the radicle (embryonic root), the hypocotyl (emb ...
*
Sterigma
In biology, a sterigma (pl. sterigmata) is a small supporting structure.
It commonly refers to an extension of the basidium (the spore-bearing cells) consisting of a basal filamentous part and a slender projection which carries a spore at the t ...
, the "woody peg" below the leaf of some conifers
* Thorn (botany), true thorns, as distinct from spines or prickles, are short shoots