''Amelanchier'' ( ), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry (or just sarvis), juneberry, saskatoon, sugarplum, wild-plum or chuckley pear,A Digital Flora of Newfoundland and Labrador Vascular Plants /ref> is a
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of about 20 species of
deciduous
In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, a ...
tree
In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
s in the rose family (
Rosaceae
Rosaceae (), the rose family, is a medium-sized family of flowering plants that includes 4,828 known species in 91 genera.
The name is derived from the type genus ''Rosa''. Among the most species-rich genera are '' Alchemilla'' (270), '' Sorbu ...
United States
The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 U.S. state, states, a Washington, D.C., federal district, five ma ...
and adjacent southeastern
Canada
Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by tota ...
, and at least one species is native to every U.S. state except
Hawaii
Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only ...
and to every Canadian province and territory. Two species also occur in
Asia
Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an ...
, and one in
Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a subcontinent of Eurasia and it is located enti ...
. The taxonomic classification of shadbushes has long perplexed botanists, horticulturalists, and others, as suggested by the range in number of species recognized in the genus, from 6 to 33, in two recent publications. A major source of complexity comes from the occurrence of hybridization,
polyploidy
Polyploidy is a condition in which the cells of an organism have more than one pair of (homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two sets of chromosomes, where each set contains ...
, and
apomixis
In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cutt ...
(asexual seed production), making
species
In biology, a species is the basic unit of Taxonomy (biology), classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of ...
difficult to characterize and identify.University of Maine ''Amelanchier'' Systematics and Evolution /ref>
The various species of ''Amelanchier'' grow to 0.2–20 m tall; some are small trees, some are multistemmed, clump-forming shrubs, and yet others form extensive low shrubby patches ( clones). The
bark
Bark may refer to:
* Bark (botany), an outer layer of a woody plant such as a tree or stick
* Bark (sound), a vocalization of some animals (which is commonly the dog)
Places
* Bark, Germany
* Bark, Warmian-Masurian Voivodeship, Poland
Arts, ...
is gray or less often brown, and in tree species smooth or fissuring when older. The leaves are deciduous, cauline, alternate, simple, lanceolate to elliptic to orbiculate, 0.5–10 x 0.5–5.5 cm, thin to coriaceous, with surfaces above glabrous or densely tomentose at flowering, and glabrous or more or less hairy beneath at maturity. The
inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
s are terminal, with 1–20 flowers, erect or drooping, either in clusters of one to four flowers, or in
raceme
A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the sh ...
s with 4–20 flowers. The
flowers
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechani ...
have five white (rarely somewhat pink, yellow, or streaked with red), linear to orbiculate petals, 2.6–25 mm long, with the petals in one species (''A. nantucketensis'') often andropetalous (bearing apical microsporangia adaxially). The flowers appear in early spring, "when the
shad
The Alosinae, or the shads,Alosinae fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering.
Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
is a berry-like
pome
In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Well-known pomes include the apple, pear, and quince.
Etymology
The word ''pome'' entered English in the late 14th century, an ...
, red to purple to nearly black at maturity, 5–15 mm diameter, insipid to delectably sweet, maturing in summer.
''Amelanchier'' plants are valued horticulturally, and their fruits are important to wildlife.
Amelanchier alnifolia
''Amelanchier alnifolia'', the Saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub with an edible berry-like fruit, native to North America.
Description
It is a deciduous shrub or s ...
Amelanchier arborea
''Amelanchier arborea'' (downy serviceberry or common serviceberry), is native to eastern North America from the Gulf Coast north to Thunder Bay in Ontario and Lake St. John in Quebec, and west to Texas and Minnesota.
Other common names are "sh ...
'' – downy serviceberry
*'' Amelanchier asiatica'' – Korean juneberry or Asian serviceberry
*''
Amelanchier australis
''Amelanchier australis'' is a plant species native to Mexico and New Mexico. Some sources consider this to be the same species as ''A. utahensis'' but The Plant List
The Plant List was a list of botanical names of species of plants created ...
Amelanchier bartramiana
''Amelanchier bartramiana'' is a species of serviceberry. Common names include mountain serviceberry, mountain shadbush, Bartram's serviceberry, mountain juneberry, Bartram juneberry, and the oblongfruit serviceberry.
The leaves of ''A. bartram ...
'' – mountain shadbush
*''
Amelanchier canadensis
''Amelanchier canadensis'' (bilberry,Canadian Wildlife Federation/ref> Canadian serviceberry, chuckle-berry, currant-tree, juneberry, shad-blow serviceberry, shad-blow, shadbush, shadbush serviceberry, sugarplum, thicket serviceberry) is a specie ...
Amelanchier interior
''Amelanchier interior'' or Wiegand's shadbush is type of serviceberry shrub. It produces a sweet tasting edible fruit called a pome, which can be eaten raw or cooked. The fruit has a sweet flavor. This species is a deciduous tree. It grows on hi ...
'' – Wiegand's shadbush
*''
Amelanchier intermedia
''Amelanchier intermedia'', also known as intermediate serviceberry, shadbush or juneberry, is a wetland
A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for we ...
''
*''
Amelanchier laevis
''Amelanchier laevis'', the smooth shadbush, smooth serviceberry or Allegheny serviceberry, is a North American species of tree in the rose family Rosaceae, growing up to tall. It is native to eastern Canada and the eastern United States, from ...
Amelanchier nantucketensis
''Amelanchier nantucketensis'', also known as the Nantucket serviceberry or the Nantucket shadbush, produces edible fruit called pomes. Nantucket serviceberry is of conservation concern in the wild. Its distribution extends from Nantucket and Mar ...
Amelanchier ovalis
''Amelanchier ovalis'', commonly known as snowy mespilus (a name which is also attached to the related '' A. lamarckii'') or serviceberry, is a deciduous shrub in the family Rosaceae. Its pome fruits are edible and can be eaten raw or cooked. The ...
Amelanchier pallida
''Amelanchier pallida'', the pale serviceberry or western serviceberry, is a species of ''Amelanchier'' native to the US states of California and Arizona. They are shrubs or small trees reaching , with attractive blue-green foliage. They typicall ...
'' – pale serviceberry or western serviceberry
*''
Amelanchier parviflora
''Amelanchier'' ( ), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry (or just sarvis), juneberry, saskatoon, sugarplum, wild-plum or chuckley pear,A Digital Flora of Newfoundland and Labrador Vascular Plants/ref> is a g ...
''
*''
Amelanchier sanguinea
''Amelanchier sanguinea'', known as red-twigged shadbush or roundleaf serviceberry, is a shrub native to eastern and central North America. Its native range stretches from New Brunswick to Saskatchewan south as far as northern Georgia (U.S. state ...
Amelanchier spicata
''Amelanchier spicata'', also referred to as the low juneberry, thicket shadbush, dwarf serviceberry, or low serviceberry (historically also called "pigeon berry"), is a species of serviceberry that has edible fruit, which are really pomes. The ...
Amelanchier stolonifera
''Amelanchier'' ( ), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry (or just sarvis), juneberry, saskatoon, sugarplum, wild-plum or chuckley pear,A Digital Flora of Newfoundland and Labrador Vascular Plants/ref> is a g ...
Since classifications have varied greatly over the past century, species names are often used interchangeably in the nursery trade. Several natural or horticultural
hybrid
Hybrid may refer to:
Science
* Hybrid (biology), an offspring resulting from cross-breeding
** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species
** Hybridity, the property of a hybrid plant which is a union of two diff ...
s also exist, and many ''A. arborea'' and ''A. canadensis'' plants that are offered for sale are actually hybrids, or entirely different species. ''A''. × ''grandiflora'' is another hybrid of garden origin, between ''A. arborea'' and ''A. laevis''. The cultivar 'La Paloma' has gained the
Royal Horticultural Society
The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity.
The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (No ...
’s
Award of Garden Merit
The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions.
History
The Award of Garden Merit ...
.
A taxon called '' Amelanchier lamarckii'' (or ''A.'' x ''lamarckii'') is very widely cultivated and
naturalized
Naturalization (or naturalisation) is the legal act or process by which a non-citizen of a country may acquire citizenship or nationality of that country. It may be done automatically by a statute, i.e., without any effort on the part of the i ...
in Europe, where it was introduced in the 17th century. It is
apomictic
In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cutt ...
, breeding true from seed, and probably of hybrid origin, perhaps descending from a cross between ''A. laevis'' and either ''A. arborea'' or ''A. canadensis''. While ''A. lamarckii'' is known to be of North American origin, probably from eastern Canada, it is not known to occur naturally in the wild in North America.Bean, W. J. (1976). ''Trees and Shrubs Hardy in the British Isles'' 8th ed., vol. 1. John Murray .Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .
Etymology
The origin of the generic name ''Amelanchier'' is probably derived from ''amalenquièr'', ''amelanchièr'', the
Provençal
Provençal may refer to:
*Of Provence, a region of France
* Provençal dialect, a dialect of the Occitan language, spoken in the southeast of France
*''Provençal'', meaning the whole Occitan language
*Franco-Provençal language, a distinct Roman ...
names of the European ''
Amelanchier ovalis
''Amelanchier ovalis'', commonly known as snowy mespilus (a name which is also attached to the related '' A. lamarckii'') or serviceberry, is a deciduous shrub in the family Rosaceae. Its pome fruits are edible and can be eaten raw or cooked. The ...
''.Jepson Flora ''Amelanchier alnifolia'' /ref>
The name ''serviceberry'' comes from the similarity of the fruit to the related European ''
Sorbus
''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' (''s.l.'') are commonly known as whitebeam, rowan (mountain-ash) and service tree. The exact number of species is disputed dep ...
''.
A fanciful etymology explains the name 'serviceberry' by noting that the flowers bloom about the time roads in the Appalachian mountains became passable, allowing circuit-riding preachers to resume church services. A similar etymology says that blooming serviceberry indicated the ground had thawed enough to dig graves, so burial services could be held for those who died in the winter when the only way to deal with the bodies was to allow them to freeze and wait for spring. Both of these fanciful etymologies are unlikely to be correct since the term is attested for both English and New World species as early as the 16th century, well before settlement of English North America, and serviceberry is far from unique in blossoming early in the year.
Juneberry refers to the fruits of certain species becoming ripe in June. The name saskatoon originated from a Cree noun ''misâskwatômina'' (''misāskwatōmina'', ''misaaskwatoomina'') for ''
Amelanchier alnifolia
''Amelanchier alnifolia'', the Saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub with an edible berry-like fruit, native to North America.
Description
It is a deciduous shrub or s ...
''. The city of
Saskatoon
Saskatoon () is the largest city in the Canadian province of Saskatchewan. It straddles a bend in the South Saskatchewan River in the central region of the province. It is located along the Trans-Canada Yellowhead Highway, and has served as ...
,
Saskatchewan
Saskatchewan ( ; ) is a province in western Canada, bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and on the south by the U.S. states of Montana and North ...
, is named after this plant.
Shadberry refers to the
shad
The Alosinae, or the shads,Alosinae Caterpillar
Caterpillars ( ) are the larva, larval stage of members of the order Lepidoptera (the insect order comprising butterfly, butterflies and moths).
As with most common names, the application of the word is arbitrary, since the larvae of sawfly ...
brimstone moth
The brimstone moth (''Opisthograptis luteolata'') is a moth of the family Geometridae. The species was first described by Carl Linnaeus in his 1758 10th edition of ''Systema Naturae''. It should not be confused with the brimstone butterfly Gone ...
,
brown-tail
The brown-tail moth (''Euproctis chrysorrhoea'') is a moth of the family Erebidae. It is native to Europe, neighboring countries in Asia, and the north coast of Africa. Descriptions of outbreaks, i.e., large population increases of several year ...
rough prominent
The rough prominent (''Nadata gibbosa'') is a moth of the family Notodontidae, subfamily Phalerinae. It is also known as the white-dotted prominent and the tawny prominent. The species was first described by James Edward Smith in 1797. This com ...
winter moth
:''In North America, "winter moth" usually denotes the invasive species ''Operophtera brumata'', but may also mean refer to a native species, '' Erannis tiliaria'' (linden looper) or ''Operophtera bruceata'' (bruce spanworm).''
The winter moth (' ...
, and the red-spotted purple and the white admiral (both ''
Limenitis arthemis
''Limenitis arthemis,'' the red-spotted purple or white admiral, is a North American butterfly species in the cosmopolitan genus ''Limenitis''. It has been studied for its evolution of mimicry, and for the several stable hybrid wing patterns wi ...
''), as well as various other herbivorous insects feed on ''Amelanchier.'' Many insects and diseases that attack orchard trees also affect this genus, in particular trunk borers and ''
Gymnosporangium
''Gymnosporangium'' is a genus of heteroecious plant-pathogenic fungi which alternately infect members of the family Cupressaceae, primarily species in the genus ''Juniperus'' (junipers), and members of the family Rosaceae in the subfamily Amyg ...
'' rust. In years when late flowers of ''Amelanchier'' overlap those of wild roses and
brambles
A bramble is any rough, tangled, prickly shrub, usually in the genus '' Rubus'', which grows blackberries, raspberries, or dewberries. "Bramble" is also used to describe other prickly shrubs, such as roses (''Rosa'' species). The fruits in ...
The fruit of several species are excellent to eat raw, sweetish, and strongly accented by the almond-like flavour of the seeds. Selections from ''
Amelanchier alnifolia
''Amelanchier alnifolia'', the Saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub with an edible berry-like fruit, native to North America.
Description
It is a deciduous shrub or s ...
'' have been chosen for fruit production, with several named
cultivar
A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s. Other cultivars appear to be derived from hybridization between ''A. alnifolia'' and '' A. stolonifera''. Propagation is by seed, divisions, and grafting. Serviceberries graft so readily that grafts onto other genera, such as '' Crataegus'' and ''
Sorbus
''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' (''s.l.'') are commonly known as whitebeam, rowan (mountain-ash) and service tree. The exact number of species is disputed dep ...
'', are often successful.
The fruit can be harvested for pies, muffins, jams, and wine. The saskatoon berry is harvested commercially. One version of the Native American food
pemmican
Pemmican (also pemican in older sources) is a mixture of tallow, dried meat, and sometimes dried berries. A calorie-rich food, it can be used as a key component in prepared meals or eaten raw. Historically, it was an important part of indigenous ...
was flavored by serviceberry fruits in combination with minced dried meat and fat.
The
wood
Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of ligni ...
is brown, hard, close-grained, and heavy. The heartwood is reddish-brown, and the sapwood is lighter in color. It can be used for tool handles and fishing rods. Native Americans used it for arrow shafts. Members of the
Pit River Tribe
The Pit River Tribe is a federally recognized tribe of eleven bands of indigenous peoples of California. They primarily live along the Pit River in the northeast corner of California.ornamental shrubs, grown for their flowers, bark, and fall color. All need similar conditions to grow well, requiring good drainage, air circulation (to discourage leaf diseases), watering during drought, and
soil
Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
appropriate for the species.
George Washington planted specimens of ''Amelanchier'' on the grounds of his estate,
Mount Vernon
Mount Vernon is an American landmark and former plantation of Founding Father, commander of the Continental Army in the Revolutionary War, and the first president of the United States George Washington and his wife, Martha. The estate is ...
, in
Virginia
Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the East Coast of the United States, Atlantic Coast and the Appalachian Mountains. The geography an ...