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''Amelanchier'' ( ), also known as shadbush, shadwood or shadblow, serviceberry or sarvisberry (or just sarvis), juneberry, saskatoon, sugarplum, wild-plumCampbell, C. S., Dibble, A. C., Frye, C. T., & Burgess, M. B. (2015). ''Amelanchier''. In FNA Editorial Committee, ''Flora of North America'' 9. Magnoliophyta: Rosidae (in part): Rosales (in part). Oxford University Press, New York.
/ref> or chuckley pear,A Digital Flora of Newfoundland and Labrador Vascular Plants

/ref> is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of about 20 species of
deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...
-leaved
shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...
s and small
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...
s in the rose family (
Rosaceae Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...
). ''Amelanchier'' is native to temperate regions of the
Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...
, growing primarily in early successional habitats. It is most diverse taxonomically in
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
, especially in the northeastern
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
and adjacent southeastern
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
, and at least one species is native to every U.S. state except
Hawaii Hawaii ( ; ) is an island U.S. state, state of the United States, in the Pacific Ocean about southwest of the U.S. mainland. One of the two Non-contiguous United States, non-contiguous U.S. states (along with Alaska), it is the only sta ...
and to every Canadian province and territory. Two species also occur in
Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...
, and one in
Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...
. The taxonomic classification of shadbushes has long perplexed botanists, horticulturalists, and others, as suggested by the range in number of species recognized in the genus, from 6 to 33, in two recent publications. A major source of complexity comes from the occurrence of hybridization,
polyploidy Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
, and
apomixis In botany, apomixis is asexual development of seed or embryo without fertilization. However, other definitions include replacement of the seed by a plantlet or replacement of the flower by bulbils. Apomictically produced offspring are geneti ...
(asexual seed production), making
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
difficult to characterize and identify.University of Maine
''Amelanchier'' Systematics and Evolution
/ref> A pome fruit, the berries are commonly consumed by wildlife and picked by humans for uses in baked goods. The Canadian city of
Saskatoon Saskatoon () is the largest city in the Provinces and territories of Canada, Canadian province of Saskatchewan. It straddles a bend in the South Saskatchewan River in the central region of the province. It is located along the Trans-Canada Hig ...
,
Saskatchewan Saskatchewan is a Provinces and territories of Canada, province in Western Canada. It is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and to the south by the ...
is named after the berry via a
Cree The Cree, or nehinaw (, ), are a Indigenous peoples of the Americas, North American Indigenous people, numbering more than 350,000 in Canada, where they form one of the country's largest First Nations in Canada, First Nations. They live prim ...
term describing the sweet fruit which was wild-harvested for food by Aboriginal people and early settlers of western Canada.


Description

The various species of ''Amelanchier'' grow to tall; some are small trees, some are multistemmed, clump-forming shrubs, and yet others form extensive low shrubby patches ( clones). The
bark Bark may refer to: Common meanings * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Arts and entertainment * ''Bark'' (Jefferson Airplane album), ...
is gray or less often brown, and in tree species smooth or fissuring when older. The
leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...
are deciduous, cauline, alternate, simple, lanceolate to elliptic to orbiculate, 0.5–10 x 0.5–5.5 cm, thin to coriaceous, with surfaces above glabrous or densely tomentose at flowering, and glabrous or more or less hairy beneath at maturity. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
s are terminal, with 1–20 flowers, erect or drooping, either in clusters of one to four flowers, or in
raceme A raceme () or racemoid is an unbranched, indeterminate growth, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are ...
s with 4–20 flowers. The
flowers Flowers, also known as blooms and blossoms, are the reproductive structures of flowering plants ( angiosperms). Typically, they are structured in four circular levels, called whorls, around the end of a stalk. These whorls include: calyx, m ...
have five white (rarely somewhat pink, yellow, or streaked with red), linear to orbiculate petals, 2.6–25 mm long, with the petals in one species (''A. nantucketensis'') often andropetalous (bearing apical microsporangia adaxially). The flowers appear in early spring, "when the
shad The Alosidae, or the shads, are a family (biology), family of clupeiform fishes. The family currently comprises four genera worldwide, and about 32 species. The shads are Pelagic fish, pelagic (open water) schooling fish, of which many are anadr ...
run" according to North-American tradition (leading to names such as "shadbush"). The
fruit In botany, a fruit is the seed-bearing structure in flowering plants (angiosperms) that is formed from the ovary after flowering. Fruits are the means by which angiosperms disseminate their seeds. Edible fruits in particular have long propaga ...
is a berry-like
pome In botany, a pome is a type of fruit produced by flowering plants in the subtribe Malinae of the family Rosaceae. Pome fruits consist of a central "core" containing multiple small seeds, which is enveloped by a tough membrane and surrounded by a ...
, red to purple to nearly black at maturity, 5–15 mm diameter, insipid to delectably sweet, maturing in summer. ''Amelanchier'' plants are valued horticulturally, and their fruits are important to wildlife. Some orchards are cultivated in the
Canadian prairie provinces The Canadian Prairies (usually referred to as simply the Prairies in Canada) is a region in Western Canada. It includes the Canadian portion of the Great Plains and the Prairie provinces, namely Alberta, Saskatchewan, and Manitoba. These provin ...
which are subjected to severe winter cold of , indicating the hardiness of the plant having a lifespan up to 50 years.


Species

Species accepted by the
Plants of the World Online Plants of the World Online (POWO) is an online taxonomic database published by the Royal Botanic Gardens, Kew. History Following the Convention on Biological Diversity, the Royal Botanic Gardens in Kew launched Plants of the World Online i ...
as of :


Natural hybrids

* '' Amelanchier × lamarckii'' ( A. arborea × A. laevis.)– Juneberry * '' Amelanchier × neglecta'' (A. bartramiana × A. laevis.) * '' Amelanchier × quinti-martii'' (A. arborea × A. bartramiana. ) * '' Amelanchier × spicata'' (A. alnifolia × A. humilis.) - low juneberry


Garden hybrids

Since classifications have varied greatly over the past century, species names are often used interchangeably in the nursery trade. Several natural or horticultural hybrids also exist, and many ''A. arborea'' and ''A. canadensis'' plants that are offered for sale are actually hybrids, or entirely different species. ''A''. × ''grandiflora'' is another hybrid of garden origin, between ''A. arborea'' and ''A. laevis''. The cultivar 'La Paloma' has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. It includes the full range of cultivated p ...
. A taxon called '' Amelanchier lamarckii'' (or ''A.'' x ''lamarckii'') is very widely cultivated and
naturalized Naturalization (or naturalisation) is the legal act or process by which a non-national of a country acquires the nationality of that country after birth. The definition of naturalization by the International Organization for Migration of the ...
in Europe, where it was introduced in the 17th century. It is
apomictic In botany, apomixis is asexual reproduction, asexual development of seed or embryo without Fertilisation, fertilization. However, other definitions include replacement of the seed by a plantlet or replacement of the flower by bulbils. Apomictic ...
, breeding true from seed, and probably of hybrid origin, perhaps descending from a cross between ''A. laevis'' and either ''A. arborea'' or ''A. canadensis''. While ''A. lamarckii'' is known to be of North American origin, probably from eastern Canada, it is not known to occur naturally in the wild in North America.Bean, W. J. (1976). ''Trees and Shrubs Hardy in the British Isles'' 8th ed., vol. 1. John Murray .Rushforth, K. (1999). ''Trees of Britain and Europe''. Collins .


Etymology

The origin of the generic name ''Amelanchier'' is probably derived from ''amalenquièr'', ''amelanchièr'', the Provençal names of the European '' Amelanchier ovalis''.Jepson Flora
''Amelanchier alnifolia''
/ref> The name ''serviceberry'' comes from the similarity of the fruit to the related European ''
Sorbus ''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' ('' s.str.'') are commonly known as rowan or mountain-ash. The genus used to include species commonly known as whitebeam, cheque ...
''. Juneberry refers to the fruits of certain species becoming ripe in June. The name saskatoon originated from a Cree noun ''misâskwatômina'' (''misāskwatōmina'', ''misaaskwatoomina'') for ''
Amelanchier alnifolia ''Amelanchier alnifolia'', the saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub native to North America. It is a member of the rose family, and bears an edible berry-like fruit. D ...
''. The city of
Saskatoon Saskatoon () is the largest city in the Provinces and territories of Canada, Canadian province of Saskatchewan. It straddles a bend in the South Saskatchewan River in the central region of the province. It is located along the Trans-Canada Hig ...
,
Saskatchewan Saskatchewan is a Provinces and territories of Canada, province in Western Canada. It is bordered on the west by Alberta, on the north by the Northwest Territories, on the east by Manitoba, to the northeast by Nunavut, and to the south by the ...
, is named after this plant. Shadberry refers to the
shad The Alosidae, or the shads, are a family (biology), family of clupeiform fishes. The family currently comprises four genera worldwide, and about 32 species. The shads are Pelagic fish, pelagic (open water) schooling fish, of which many are anadr ...
runs in certain New England streams, which generally took place about when the trees bloomed.


Ecology

''Amelanchier'' plants are preferred browse for deer and rabbits, and heavy browsing pressure can suppress natural regeneration.
Caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...
s of such
Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...
as the brimstone moth,
brown-tail The brown-tail moth (''Euproctis chrysorrhoea'') is a moth of the family Erebidae. It is native to Europe, neighboring countries in Asia, and the north coast of Africa. Descriptions of outbreaks, i.e., large population increases of several years ...
,
grey dagger The grey dagger (''Acronicta psi'') is a moth of the family Noctuidae. Distribution This species can be found from Europe and North Africa to northern Iran, central Asia, southern and central Siberia and Mongolia. In the Levant it is found in L ...
,
mottled umber The mottled umber (''Erannis defoliaria'') is a moth of the family Geometridae. It is common throughout much of the Palearctic region. The species was Species description, first described by Carl Alexander Clerck in 1759. Distribution The specie ...
, rough prominent,
satellite A satellite or an artificial satellite is an object, typically a spacecraft, placed into orbit around a celestial body. They have a variety of uses, including communication relay, weather forecasting, navigation ( GPS), broadcasting, scient ...
,
winter moth The winter moth (''Operophtera brumata'') is a moth of the family Geometridae. It is an abundant species in Europe and the Near East and a famous study organism for evaluating insect population dynamics. It is one of very few lepidopterans of te ...
, red-spotted purple and white admiral (both ''
Limenitis arthemis ''Limenitis arthemis,'' the red-spotted purple or white admiral, is a North American butterfly species in the cosmopolitan genus ''Limenitis''. It has been studied for its evolution of mimicry, and for the several stable hybrid wing patterns wit ...
''), as well as various other herbivorous insects feed on ''Amelanchier.'' Many insects and diseases that attack orchard trees also affect this genus, in particular trunk borers and '' Gymnosporangium'' rust. In years when late flowers of ''Amelanchier'' overlap those of wild roses and
brambles ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...
, bees may spread bacterial
fireblight Fire blight, also written fireblight, is a contagious disease affecting apples, pears, and some other members of the family Rosaceae. It is a serious concern to apple and pear producers. Under optimal conditions, it can destroy an entire orchard ...
.


Uses and cultivation

The fruit of several species is safe to eat raw, possessing a mild sweetness strongly accented by the almond-like flavour of the seeds. Selections from ''
Amelanchier alnifolia ''Amelanchier alnifolia'', the saskatoon berry, Pacific serviceberry, western serviceberry, western shadbush, or western juneberry, is a shrub native to North America. It is a member of the rose family, and bears an edible berry-like fruit. D ...
'' have been chosen for fruit production, with several named
cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...
s. Other cultivars appear to be derived from hybridization between ''A. alnifolia'' and '' A. stolonifera''. Propagation is by seed, divisions, and grafting. Serviceberries graft so readily that grafts onto other genera, such as ''
Crataegus ''Crataegus'' (), commonly called hawthorn, quickthorn, thornapple, Voss, E. G. 1985. ''Michigan Flora: A guide to the identification and occurrence of the native and naturalized seed-plants of the state. Part II: Dicots (Saururaceae–Cornacea ...
'' and ''
Sorbus ''Sorbus'' is a genus of over 100 species of trees and shrubs in the rose family, Rosaceae. Species of ''Sorbus'' ('' s.str.'') are commonly known as rowan or mountain-ash. The genus used to include species commonly known as whitebeam, cheque ...
'', are often successful. The fruit can be harvested for pies, muffins, jams, and wine. The saskatoon berry is harvested commercially. One version of the Native American food
pemmican Pemmican () (also pemican in older sources) is a mixture of tallow, dried meat, and sometimes dried berries. A calorie-rich food, it can be used as a key component in prepared meals or eaten raw. Historically, it was an important part of indigeno ...
was flavored by serviceberry fruits in combination with minced dried meat and fat. The
wood Wood is a structural tissue/material found as xylem in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulosic fibers that are strong in tension and embedded in a matrix of lignin t ...
is brown, hard, close-grained, and heavy. The heartwood is reddish-brown, and the sapwood is lighter in color. It can be used for tool handles and fishing rods. Native Americans used it for arrow shafts. Members of the Pit River Tribe would use the wood to create a sort of body armor, crafting it into a heavy robe or overcoat and corset armor worn during fighting.Merriam, C. Hart 1966 Ethnographic Notes on California Indian Tribes. University of California Archaeological Research Facility, Berkeley (p. 222)


Garden history

Several species are very popular ornamental shrubs, grown for their flowers, bark, and fall color. All need similar conditions to grow well, requiring good drainage, air circulation (to discourage leaf diseases), watering during drought, and
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
appropriate for the species.
George Washington George Washington (, 1799) was a Founding Fathers of the United States, Founding Father and the first president of the United States, serving from 1789 to 1797. As commander of the Continental Army, Washington led Patriot (American Revoluti ...
planted specimens of ''Amelanchier'' on the grounds of his estate,
Mount Vernon Mount Vernon is the former residence and plantation of George Washington, a Founding Father, commander of the Continental Army in the Revolutionary War, and the first president of the United States, and his wife, Martha. An American landmar ...
, in
Virginia Virginia, officially the Commonwealth of Virginia, is a U.S. state, state in the Southeastern United States, Southeastern and Mid-Atlantic (United States), Mid-Atlantic regions of the United States between the East Coast of the United States ...
.


References


External links


''Juneberry''
in What Am I Eating? A Food Dictionary {{Authority control Rosaceae genera Plants used in Native American cuisine Fruit trees